NATIONAL LIBRARY OF MEDICINE Washington Founded 1836 U. S. Department of Health, Education, and Welfare Public Health Service SOLD BY JAMES WEBSTER, No. 10, South Eighth street, i PHILADELPHIA. e~~^^— passes obliquely outward from the apex behind. As the figure is irregular, the side next the nose does not partake of this general obliquity, but extends in a straight direction from behind forwards. The orbit is somewhat contracted at its orifice, and en- larged immediately within. The form of the orifice is ra- ther oval; as the transverse diameter is longer than the verti' al. Seven bones are concerned in the formation of this cavity: The os frontis and a portion of the lesser wing ot the sphenoid bone above; the os planum or ethmoid, the os unguis, and the nasal process of the upper maxillary bone, on the inside; the orbitar plate of the upper maxil- lary bone, and the os palati, below; the os malae, and or- bitar plate of the sphenoid bone, on the outside. On the upper surface is the depression for the lacrymal gland; and at the orifice is the notch or foramen for the sujjraorbitary vessels, &c. which have already been men- tioned. Orbit of the Eye. 55 On the inner surface are two longifodinal sutures, which connect the os planum and the os unguis, to the os irontis above, and the os maxillare below. In the upper suture are the two internal orbitary foramina mentioned in the des- cription of the os frontis, the anterior of which transmits a fibre of the ophthalmic nerve, with an artery and vein; the posterior transmits only an artery and vein. There are also two smaller vertical sutures on each side of the os unguis. On the anterior part of this inner surface is the ridge of the os unguis, and the groove for accommodat- ing the lacrymal sac, which passes into the canal of the same name immediately below. On the lower surface is the aforesaid canal, formed by the nasal and orbitar processes of .the upper maxillary bone, and that part of the os unguis which is anterior to the ridge. On the posterior part of this surface is a groove which pro- ceeds forwards, and penetrating into the bone becomes a canal that terminates in the infra-orbitar foramen; this groove in the bone is made a canal by the periosteum. The thin plate which forms this surface is the partition between the antrum maxillare and the orbit of the eye, and is more or less absorbed in those cases where polypi of the antrum maxillare occasion a protrusion of the eye. The external surface, formed by the malar bone and the orbitar plate of the sphenoid, is almost flat. In the poste- rior part of the orbit it is bounded by two large fissures, which are now to be described. In the posterior part of the orbit are three apertures: The optic foramen, the sphenoidal fissure, and the sphenoc maxillary fissure. The optic foramen ope us almost at the bottom of the wbit on the inside; its direction is forwards and outwards. 56 Cavities of the Nose. The sphenoidal fissure, formed principally by the les- ser and greater wings of the sphenoidal bone, begins at the bottom of the orbit, and extends forward, upward and outward. It is broad a the commencement, and gradually diminishes to a fissure. This fissure opens directly into the cavity of the cranium, and admits the third, fourth, sixth, and one branch of the fifth pair of nerves, an artery and a vein. The spheno-maxillary fissure commences also at the bottom of the orbit, and extends forward, outward and downward, between the maxillary bone and the orbitar plate of the sphenoid, from the body of the sphenoid to the malar bone. This fissure opens from the orbit direct- ly into the zygomatic fossa. In the recent subject it is closed, and only transmits the infraorbitary nerve and vessels, and a small branch of the superior maxillary nerve. • The Cavities of the Nose. These cavities, which are separated from each other by the septum narium, are contained between the cribri- form plate of the ethmoid and the palatine processes of the upper maxillary and palate bones, and between the an- terior and posterior nares. They are therefore of consi- derable extent in these directions; but the distance from the septum to the opposite side of the nose is so small, that each cavity is very narrow. The upper surface of each cavity consists of that por- tion of the cribriform plate of the ethmoid which is be- tween the septum and the cellular portions. Anterior to this, each cavity is bounded by the internal surface of the os nasi of its respective side; and posterior to it, by the interior surface of the body of the sphenoid bone. These anterior and posterior surfaces form obtuse angles with Cavities cfthe Nose. 57 the upper surface of the nose, and are immediately above the openings culled antes ior and posterior nares. The an- terior surface partakes of the figure of the os nasi; the up- per surface h:;s the perforations of the cribriform plate; the posterior surface has an opening, equal in diameter to a small quill, that leads into the sphenoidal cell, a.id it is also broader than -he anterior or superior surface. The internal surface, formed by the septum of the nose, which is composed of the vomer, the nasal plate of the eth- moid, and a cartilaginous plate, is flat, but rather inclined to one side or the other, so as to make a difference in the nasal cavities. The external surface is very irregular: it is formed by the cellular portions of the ethmoid; bv a small portion of the os unguis; by the upper maxillary bone; the os turbina- ting, inferius; the os palati; and the internal pterygoid pro- cess of the os sphenoides. The upper part of this surface is formed by the internal surface of the cellular portions of the ethmoid, which have been described at page 26. It ex- tends from the sphenoid bone, very ne.-'.r to the ossa nasi; and is uniformly flat and rough. About the middle of it begins a deep groove, which penetrates into the cellular structure of the ethmoides, and passes obliquely downwards and backwards. At the upper end of this groove is the foramen by which the pos- terior ethmoidal cells communicate with the nasal cavity. This is the upper channel or meatus of the nose. At the posterior end of it is a large foramen formed by the nasal plate of the os palati and the pterygoid process of the os sphenoides, and therefore called pterygo-palatine foramen. It opens externally and transmits a nerve and an arterv to the nose. Below the meatus is the upper spongy bone, which pre- Vol. I. II 58 Cavities of the Nose. sents a convex surface; its lower edge is rolled up and not connected with the parts about it. This spongy bone co- vers a foramen in the ethmoid bone, bv which its anterior cells and the frontal sinuses communicate with the nose. Below this spongy bone, is the middle channel or meatus of the nose. This channel extends from the anterior to the posterior part of the cavity. It is verydeep, as it penetrates to the maxillary bone. The cells of the ethmoid are above it; the inferior turbinated bone below it; and the upper spongy bone projects over it. In this channel is the open- ing of the great cavity of the upper maxillary bone. At the anterior extremity of it is a small portion of the os unguis, which intervenes between the nasal process of the upper maxillary bone and the cells of the ethmoid, and continues down to tht lower spongy bone. The lower spongy bone is nearly horizontal, and very conspicuous. It extends* almost from one opening of the nose to the other. Under this bone is the third and largest channel or mea- tus of the nose. It is made large by an excavation of the upper maxillary bone, particularly at the anterior part. It affords a direct and very easy passage to the posterior opening of the nose and the throat. Near the anterior extremity of this meatus is the lower orifice of the lacrymal duct, which is so situated that a probe properly curved can be readily passed into it through the nostril. There are then four foramina on each side, which form communications between the cavities of the nose and the adjacent cells, viz. One in the upper meatus which leads to the posterior ethmoid cells. Cavity of the Cranium. 59 A second in the middle meatus which leads to the an- terior ethmoid cells and the frontal sinuses. A third in the same meatus which opens into the max- illary sinus. A fourth in the anterior surface of the body of the sphe- noidal bone, which opens into the sphenoidal sinus. To these must be added the opening of the lacrymal canal. It will be useful to the student of anatomy, after placing three or four of the uppermost cervical vertebrae in their natural situation, to take a view of The Cavity between the Spine and the posterior Nares, which is bounded, above, by the cuneiform process, pass- ing obliquely upward and forward; laterally, by soft parts not yet described; behind, by the bodies of the cervical vertebrae; and before, by the posterior nares, each of which is oblong in form, rounded above, flat below, and separat- ed from the other by a thin partition, the vomer. The Cavity of the Cranium. The upper concave surface of < is cavity corresponds with the figure of the cranium. The ri ige in it for sup- porting the falciform process of the dura mater, the groove made by the longitudinal sinus, the impressions of the ar- teries, and the pits made by the convolutions of the brain, are particularly to be noticed. 60 Basis oj'the Cranium. The Basis of the Cranium is much more important, it is divided into three fossae on each side: the anterior of those are most superficial, and the posterior the deepest. The bottoms of the anterior fossa are formed by the orbitar processes of the os frontis, and consequently are convex; between them is the cribri- form plate of the ethmoid, which is commonly sunk below the adjoining surface. The crista galli is very conspicuous; and the foramen c;ecum can almost always be seen. The crista galli is evidently the beginning of the prominent ridge, which continues on the os frontis, and supports the falx of the dura mater. The posterior margins of these fossae are formed by the less r wings of the sphenoid bone. The middle foss.e are formed by the great wings of the sphenoidal bone*, and by the squamous and petrous por- tions of the temporal bone. They are lower than the ante- rior, and higher than the posterior fossae. The projection of the margin of the anterior fossae into these cavities, corresponds with the separation between the anterior and middle lobes of the brain. The suture between the sphenoidal and temporal bones is evident in these fossae. The upper surface of ihe body of the sphenoid bone, or the sella turcica, is between them; and all the peculiarities of its surface are very conspicuous. The first five foramina of the sphenoidal bone can be easily ascertained, and also, the anterior foramen lacerum and termination of the fora- men caroticum, with the impressions made by the carotid arteries on the sides of the sella turcica. The petrous por- tions of the temporal bones are the posterior boundaries dTthe middle fossa;. Their oblique direction, inwards and forwards, is particularly remarkable; being formed like Basis of the Cranium. 6.1 triangular pyramids. Two of their sides are in the cavity of the cranium; one, which is anterior, forms a portion of the middle fossa; and the other forms a part of the posterior fossa. The edge between them is very prominent, and has the tentorium or horizontal process of the dura mater at- tached to it. On the anterior surface, in the middle fossa, mav be traced the groove and the foramen for the vidian nerve. The posterior fossae are larger as well as deeper than the other two. Their boundaries are well defined by the edges of the petrous bones above mentioned, and b,y the grooves for the horizontal parts of the lateral sinuses. These fossae are nearly separated from the general cavity by the ten- torium, which is attached to the edge of the petrous bone and also to the edge of the horizontal part of the groove for the lateral sinuses. On the tentorium lie the posterior lobes of the cerebrum; and under it, in the^se fossae, is the cerebellum. These fossae may be considered as one great cavity, which is circular behind, and somewhat angular before. The angular surfaces are formed by the posterior sides of the petrous portions. Between them, is the oblique surface of the cuneiform process of the occipital bone, which de- scends to the great foramen. On the surface of each pe- trous bone is the meatus auditorius internus, and the ori- fice of the aqueduct of the vestibule. Behind the petrous portion, the groove for the lateral sinus is very conspicu- ous : it terminates in the posterior foramen lacerum, which is evidently formed by the temporal and the occipital bones. At the anterior part of this foramen is most com- monly a small bony process, which separates the eighth pair of nerves from the internal jugular vein, as they pass out here. 52 Basis of the Cranium. The anterior condyloid foramen for the passage of the ninth pair of nerves, appears in the surface of the great occipital hole, immediately below the foramen lacerum. From the back part of this hole the spine, which forms the lower limb of the cross, passes up; and on each side of it are the great depressions which accommodate the two lobes of the cerebellum. External Basis of the Cranium. When the head is inverted we see the external protu- berances of the os occipitis, formerly described. The mas- toid processes of the ossa temporum are on the same trans- verse line with the great foramen of the os occipitis; but the foramen being larger extends farther forward. On the inside of the mastoid process, the fissure for the digastric muscle is very conspicuous, and also the suture between the mastoid process and the occipital bone. The oblique direction of the occipital condyles and the slanting position of their articulating surfaces are particu- larly striking. The posterior and anterior condyloid fora- mina for the cervical veins, and the ninth pair of nerves, are also in view. The position of the cuneiform process of tiie os occipitis is by no means horizontal, but extends forwards and upwards. The petrous or pyramidal portion of the temporal bone commences between the mastoid process, and the glenoid cavity for the condyle of the lower jaw, and extends obliquely forwards and inwards, having the occipital bone behind it, and the glenoid cavity and the os sphenoides before it. At the commencement, the sur- face of the petrous portion is not horizontal but oblique, sloping into the glenoid cavity with a sharp edge down- wards. This edge in some cases is curved so as to sur- round the basis of the stiloid process, which arises in con- Basis of the Cranium. 63 tact with it, and projects downwards, on each side of the vertebrae. Between the mastoid and stiloid process, is the foranu-n stilo-mastoideum. On the inside of the stiloid process, and rather anterior to it, is the foramen lacerum posterior, for the internal jugular vein, the eighth pair of nerves, Sec. This foramen passes obliquely backwards and upwards, and is bounded behind by the jugular pro- cess of the os occipitis, which bone seems to contribute most to its formation. Very near to this holt, on the inside, is the anterior condyloid foramen; and rather anterior to it is the opening of the carotid canal, which forms a curve in the bone as it passes upwards, inwards, and forwards. From the foramen lacerum posterior, the suture, between the cuneiform process of the occipital and the petrous por- tion of the temporal, extends to the foramen lacerum ante- rior; which is closed by cartilage in the recent subject, but is of an irregular and rather triangular form in the mace- rated head: this hole is formed by the occipital, sphenoid- al and petrous bones. The suture or connexion between the petrous bone and the os sphenoides, is continued on the anterior side of the petrous bone, from the fissure of the glenoid cavity to the anterior foramen lacerum. The stiloid process of the os sphenoides, which is seldom more than four lines in length, appears at the edge of this suture. On the inside of the glenoid cavity, and on the inside of this process, in the suture formed between the petrous and sphenoid bones, is the bony orifice of the eustachian tube. The foramen spinale for the middle artery of the dura mater is at a very small distance from the eustachian tube, immediately anterior to it; and at a small distance on the inside and in front of this foramen, is the foramen ovale, for tlu inferior maxillary nerve, or the third branch of the fifth pair. 64 Side of the Head. The Side of the Head. Those portions of the side of the head which are form- ed bv the frontal, parietal and occipital bones, and by the squamous part of the temporal, require no explanation here; but the region which is behind the malar and upper maxillary bone, and within the zygomatic processes of the temporal and malar bones, which comprises part of the temporal and zygomatic fossae of some anatomists, is both important and obscure. To obtain a view of this, the lower jaw should be re- moved, and the zygoma sawed away, in one preparation; and in another, the upper maxillary and palate bones, of one side, should be applied in their natural position, to the os sphenoides, without an\ of the other bones. The upper part of this region, formed by the sphenoi- dal, frontal and malar bones, is made concave by the form of the external angular part of the os frontis and of the os malae; which projects backward so as to cover a large por- tion of it. ' The lower part is formed principally by the external surface of the pterygoid process of the sphenoid bone, and by the posterior surface of the upper maxillary. Between the lower end of the pterygoid process and the upper max- illary bone, a small portion of the os palati intervenes; but in many adult subjects it is not to be distinguished from the other bones. At this place, the pterygoid process and these bones appear to be in close contact; but as they pass upwards they recede from each other so as to form a considerable aperture, which continues the whole length of th..- pterygoid process. This Jiswre, which may be call- ed pterygopalatine orpterygo-maxillary, would open into the posterior part of the cavity of the nose, if the nasal Form of the Cranium. 65 plate of the os palati did not intervene: this plate forms a partition, which separates the nose from this fissure; and the spheno-palatine foramen, formed principally by it, transmits a nerve and bloodvessels to the nose. The fissure is vertical: at the back of the orbit, it unites with the spheno-maxillary fissure of the orbit, which is almost horizontal; and at the place of their junction, the sphenoidal, or upper fissure of the orbit, opens also. The foramen rotundum, which transmits the second branch of the fifth pair, or the upper maxillary nerve, is likewise situated near this place; and when the upper max- illary, the sphenoidal, and the palate bones are in their na- tural situation, the distribution of the branches of this im- portant nerve can be easily understood : for the same view presents the course of its various branches; viz. to the nose, by the spheno-palatine foramen; to the cavity of the cra- nium, by the pterygoid foramen; to the orbit, and the in- ferior orbitary canal, by the spheno-maxillary fissure; and to the roof of the mouth, by the palato-maxillary canal. The Form of the Cranium. The form of the cranium is that of an irregular oval. The greatest length of its cavity is between a part of the os frontis above the crista galli, and of the os occipitis above the center of the crucial ridge. The greatest breadth is at about two thirds of the dis- tance from the first to the last of these positions. This transverse diameter touches the sides of the cranium near the posterior part of the basis of the petrous portion of the temporal bone. The difference between these longitudinal and transverse diameters varies greatly in different per- sons, as their craniums approach to the oval or round figures. Vol. I. I 66 Form of the Cranium. The greatest depth of the cavity is between the posterior part of the cuneiform process of the occipital bone, and a part of the cranium which is nearly over it, about the mid- dle of the sagittal suture. The figure of the cranium is somewhat varied in differ- ent races of men; and it has been much changed by the particular management of several savage nations. In North America, the Choctaw tribe of Indians were formerly accustomed to make their foreheads perfectly flat, and sloping obliquely backwards. They have latterly dis- used this practice; but one of their nation, whose head had this form, was in Philadelphia about the year 1796. At this time, a tribe who inhabit a district of country near the sources of the Missouri river, are in the practice of flattening both the frontal and occipital regions of the head; so that a small part only, of the middle of it, remains of the natural form, between these flattened sloping sur- faces. In the case of the Choctaw man above mentioned, it did not appear that his health, or his intellectual operations, were any way affected by this form of his head. During infancy, the cranium sometimes increases to a preternatural size, as disproportionate to the face as if it were affected by hydrocephalus. In many of these in- stances that disease ultimately shows itself; but in other cases, the preternatural increase of the cranium finally stops without the occurrence of disease; and the dispro- portion is lessened by the increase of the face in the ordi- nary progress of growth. In many cases where men have deviated from the ordi- nary stature, the head has preserved the common size. It is therefore said to be small in giants and large in dwarfs. Head of the Foetus. 67 The Head of the Foetus. In the foetus, those bones, which form the vault of the cranium, originally consist of one plate only; which is composed of radiated fibres. At birth, the os frontis consists of two pieces, which join each other in the middle of the forehead. The parietal bones are each in a single piece; but they are incomplete at their edges and their angles. The temporal bones have no appearance of mastoid or stiloid processes. Instead of a meatus auditorius exter- nus, there is a bony ring in which the membrana tympani is fixed. The squamous and petrous portions, and this ring, are originally formed separate; but at the period of birth they often adhere to each other. The os occipitis is composed of four pieces: the first and largest, extends from the beginning or angle of the lamb- doidal suture to the upper edge of the great occipital fora- men. Each side of the foramen, and the condyle on it, is formed by a distinct piece. The front part is formed by the cuneiform process, which is separate from the other parts and forms the fourth piece. The sphenoidal bone may be separated by maceration into hree pieces. The body and the little wings form one piece. Each of the great wings, with the pterygoid pro- cesses united to it, forms also a piece. The body of the bone is entirely solid. A large part of the ethmoid is in a cartilaginous state. It is divided into two portions by a partition of cartilage, which occupies the place of the nasal plate and the crista galli. In consequence of the imperfect formation of the bones which compose the vault of the cranium, there are several 68 Head of the Foetus. deficiencies in it. Thus the superior anterior angles of the parietal bones being incomplete, and also the upper angles of the pieces which compose the os frontis, a vacuity with four sides is occasioned, which is termed the Anterior fontanel. This opening may be distinguished by its form, as well as its greater size, from another vacu- ity which is produced in a.similar way at the other end of the sagittal suture, and called the , Posterior fontanel: but as there are only three bones concerned in its formation, viz. the two parietal and the Occipital, this vacuity is triangular. Besides these, there are two other vacuities or fontanels on each side, at the two lower corners of each parietal bone: these however are much less than those first des- cribed. The smaller fontanels do not continue open long; but the anterior fontanel is seldom completely closed before the end of the third year. It is very obvious, upon an examination of the cranium, that the center of the base is better calculated to resist pressure than any other part; as the cuneiform process of the occipital bone, the petrous portions of the temporal, and the body of the sphenoidal bone, which compose a large part of it, are very firm and substantial. The face of the foetus differs very essentially from that of the adult. Although the orbits of the eyes are very large when compared with the size of the head, that por- tion of the face which is below them is very small and has little depth. The upper maxillary bones have no sinuses in them; and their orbitar plates are not m .ch ehvated above the cavi- ties for containing the posterior teeth: in consequence, the Head of the Foetus. 69 depth of the face is very small; and its whole aspect is af- fected. The nose of the foetus differs greatly from that of the adult in respect to its sinuses; for not only are the maxil- lary cavities wanting, but those of the frontal and sphenoi- dal bones also. The lower jaw is formed in two pieces, which unite at the middle; and hence the term symphysis is used in des- cribing the chin. This bone is not only less broad in pro- portion than that of the adult, but the angles are more ob- tuse, and the processes which arise from them are more sloping. The head of the foetus is much larger in proportion to the body than that of the adult. 70 SECTION II. Of the Trunk. THE Trunk consists of the Spine, Thorax, and Pel- vis. The Spine* The spine is the long pile of bones extended from the con: Ils of the occiput to the end of the os coccygis. It som-what resembles two unequal pyramids joined in a common base. It is not, however, straight; for its upper pan 'o. mg drawn backwards by strong muscles, it gradu- ally advances forwards, to support the oesophagus, vessels of the head, &c. Then it turns backwards, to make room for the heart and lungs. It is next bent forwards, to sup- port the viscera of the abdomen. It afterwards turns backwards, for the enlargement of the pelvis. And, lastly, it is reflected forwards, for sustaining the lowest great intestines. The spine is commonly divided into true and false ver- tebrae: the former constituting the long upper pyramid, which has its base below; while the false vertebrae make the shorter lower pyramid, whose base is above. True Vertebrae, The true vertebrae are the twenty-four upper bones of the spine, on which the several motions of the trunk of our bodies are performed. Their name is derived from the Larin verb vertere. Each of these vertebrae is composed of its body and processes. The body is the thick spongy forepart, which is convex The Vertebra:. 7-1 before, concave backwards, horizontal and flat in most of them above and below. Numerous small holes, especi- ally on the fore and back part of their surface, give passage to their vessels, and allow the ligaments to enter their sub- stance. The edges of the body of each vertebra are cover- ed, especially at the forepart, with a ring of bone firmer and more solid than the substance of the body any where else. These rings seem to be joined to the vertebrae in the form of epiphyses. They are of great use in preventing the spongy bodies from being broken in the motions of the trunk. Between the bodies of each two adjoining vertebrae, a substance between the nature of ligament and cartilage is interposed; which seems to consist of concentrical curved fibres, when it is cut horizontally; but when it is divided perpendicularly, the fibres appear oblique and decussating. The outer part of these intervertebral ligaments is the most solid and hard; and they gradually become softer till they are almost in the form of a glairy liquor in the center. The external fibrous part of each is capable of being greatly extended, and of being compressed into a smaller space, while the middle fluid part is incompressible, or nearly so. The middle point is therefore a fulcrum or pivot, on which the motion of a ball and socket may be made, with such a gradual yielding of the substance of the ligament, in whatever direction our spines are moved, as saves the body from violent shocks, and their dangerous consequences. This ligamento-cartilaginous substance is firmly fixed to the horizontal surfaces of the bodies of the vertebrae, to connect them: in which it is assisted by a strong membranous ligament, which lines all their con- cave surface, and by a still stronger ligament that covers all their anterior convex surface. 72 The Vertebra:. The elastic substance seems to be in a state of com- pression by the exterior ligaments and the bones; for, if a section be made through a portion of the vertebrae and the intervertebral substance, this substance will expand, so that its surface will be much higher than that of the ver- tebrae. It is so elastic, and so much confined, in some sub- jects, that a sharp knife, if plunged into it, will be gradu- ally ejected when the hand is withdrawn. The bodies of the vertebrae are, with some exceptions, smaller and more solid above, but more spongy as they descend. The cartilages between them are thick, and the surrounding ligaments are strong in proportion to the size of the vertebrae. By this disposition, the greatest weight is supported on the broadest best secured base, and the middle of the body is allowed a large and secure motion. From each side of the body of each vertebra, a bony bridge is produced backwards, and to one side; from the posterior end of which one slanting process rises and ano- ther descends. The smooth, and generally the flattest, side of each of these four processes is covered with a smooth cartilage; and the two lower processes of each upper ver- tebra are fitted to, and articulated with, the two upper pro- cesses of the vertebra below, having their articular liga- ments fixed into the rough line round their edges. These processes are termed the oblique or articulating. From between the oblique processes of each side, an- other process extends laterally, which is called the trans- verse. From the back part of the roots of the two oblique pro- cesses, and of the transverse process of each side, a broad oblique bony plate is extended backwards: where these meet, the seventh process of the vertebrae takes its rise, and stands out backwards. This being generally sharp- The Vertebra. 73 pointed and narrow-edged, it has therefore been called spinal process; from which this whole chain of bones has got its name. Besides the common ligament which lines all the inter- nal surface of the spinal processes, as well as of the bodies, particular ligaments connect the bony bridges and pro- cesses of the contiguous vertebrae together. The substance of the processes is considerably stronger and firmer, and has a thicker external plate, than the bodies of the vertebrae themselves. The seven processes form a concavity at their forepart, which, joined to the one at the back part of the bodies, makes a great hole; and when the vertebrae are placed upon each other in their natural order, these holes form a long tube for containing the spinal marrow. In the upper and lower edge of each lateral bridge, there is a notch. These are so adapted to each other in the con- tiguous vertebrae, as to form a round hole in each side, be- tween each two vertebrae, through which the nerves that proceed from the spinal marrow and its bloodvessels pass. The articulations of these true vertebras are conse- quently double: for their bodies are joined by the interven- ing cartilage above described; and their oblique processes, being tipped with cartilages, are so connected by their li- gaments as to allow a small degree of motion on every side. Hence it is evident, that their center of motion is altered in different positions of the trunk: for, when we bow for- wards, the weight bears entirely on the bodies of the ver- tebrae; if we bend back, the oblique processes support it; if we recline to one side, we rest upon the oblique pro- cesses of that side and part of the bodies; if we stand erect, all the bodies and oblique processes have their share in our support. Vol. I. K 74 The Vertebra:. The true vertebra: are divided into three classes which agree with each other in their general structure, but are distinguished by several peculiarities. These classes are named Cervical, Dorsal, and Lumbar. The cervical are the seven uppermost vertebrae; which are distinguished from the rest by these marks: Their bo- dies are smaller and more solid than any others; and are flatted on the front surface. They are also flat behind, where small processes rise, to which the internal ligaments are fixed. The upper surface of the body of each vertebra is made hollow, by a slanting thin process which is raised on each side. The lower surface is also hollowed, but in a different manner; for here the posterior edge is raised a little, and the anterior one is considerably extended. Hence the cartilages between these vertebrae are firmly connect- ed, and their articulations are secure. These cartilages are thick, especially at their forepart; which is one reason why the vertebrae project forward as they descend, and have larger motion. Their oblique processes more justly deserve that name than those of any other vertebrae. They are situated slant- ing; the upper ones having their smooth and almost flat surfaces facing obliquely backwards and upwards; while the inferior oblique processes have these surfaces facing obliquely forwards and downwards. The transverse processes of these vertebrae are framed in a different manner from those of any other bones of the spine: for, besides the common transverse process rising from between the oblique processes of each side, there is a second one that comes out from the side of the body of each vertebra; and these two processes, after leaving a cir- cular hole for the passage of the cervical artery and vein, The Vertebra. 75 unite and form a groove on their upper surface to protect the nerves that pass in it. They terminate obtusely on each side, for the insertion of muscles. The spinal processes project backwards almost horizon- tally. They are shorter than those of any other vertebrae, and are forked or double at their ends; they therefore allow a more convenient insertion to muscles. The thick cartilages between the bodies of these cervi- cal vertebrae, the obliquity of their oblique processes, and the shortness and horizontal situation of their spinal pro- cesses, all conspire to allow them large motion. The holes between the bony cross bridges, for the pas- sage of the nerves from the spinal marrow, have their largest share formed in the lowest of the two vertebrae, to which they are common. So far most of the cervical vertebrae agree; but they have some particular differences, which require a separate consideration. The first, from its use in supporting the head, has the name of atlas. Contrary to all the other vertebrae of the spine, it has no body; but instead of it, there is a bony arch. In the convex forepart of this arch a small rising appears; and on each side of this protuberance, a small cavity may be observed. The upper and lower parts of the arch are rough and unequal, where the ligaments that con- nect this vertebra to the os occipitis, and to the second vertebra, are fixed. The back part of the arch is concave, smooth, and covered with a cartilage, in a recent subject, to receive the tooth-like process of the second vertebra. On each side of it a small rough sinuosity may be remark- ed, where the ligaments going to the sides of the tooth- like process of the following vertebra are fastened; and on f 6 The Vertebra:. each side a small rough protuberance and depression is observable, where the transverse ligament, which secures the tooth-like process in the sinuosity, is fixed, and hinders that process from injuring the medulla spinalis in the flex- ions of the head. The atlas has as little spinal process as body; but, in- stead of it, there is a large bony arch, that the muscles which pass over this vertebra at that place might not be hurt in extending the head. On the posterior and upper part of this arch there are two depressions, where the recti postici minores take their rise ; and at the lower part are two other sinuosities, into which the ligaments that connect this bone to the following one are fixed. The superior oblique processes of the atlas are large, and more horizontal than those of any other vertebra. They form an oblong concav% surface which has an inter- nal aspect, and corresponds exactly with the articulating surface on the external side of each condyle of the os oc- cipitis. Under the external edge of the posterior part of each of these cavities is the fossa, or deep open channel, in which the vertebral arteries make the circular turn, as they are about to enter the great foramen of the occipital bone, and where the tenth pair of nerves go out. In some subjects this fossa is covered with bone. The inferior oblique pro- cesess, extending from within outwards and downwards, are large, circular, and slightly concave. So that this ver- tebra, contrary to the other six, receives the bones with which it is articulated, both above and below. The transverse processes of this vertebra are not much hollowed or forked; but are longer and larger than those of any other vertebra of the neck, for the origin and in- sertion of several muscles j and therefore those muscles The Vertebra. 77 which move this vertebra on the second have a consider- able le\rer to act with, because of the distance of their in- sertion from the axis of revolution. The hole for the medulla spinalis is larger in the atlas than in any other vertebra, not only on account of the me- dulla being largest here, but also to prevent its being hurt by the motions of this vertebra on the second. This large hole, and the long transverse processes, make this the broadest vertebra of the neck. The condyles of the os occipitis move forwards and backwards in the superior oblique processes of this verte- bra; but from the figure of the bones forming these articu- lations, it is evident, that very little motion can here be allowed to either side; and there must be still less circu- lar motion. The second vertebra of the neck is called dentata. It is somewhat of a pyramidal figure, being large, and ex- tended downwards, especially in front, to enter into a hollow of the vertebra below; while the upper part has a long process, with its extremity formed into an obtuse point. This process from its supposed resemblance to a tooth, has given name to the vertebra. The side of it, on which the concave surface of the anterior arch of the first vertebra plays, is convex, smooth, and covered with a car- tilage; and it is of the same form behind, to accommodate the ligament, which is extended transversely from one rough protuberance of the first vertebra to the other, and is cartilaginous in the middle. A ligament likewise goes out in an oblique transverse direction, from each side of the processus dentatus, to be fixed at its other end to the first vertebra, and to the occipital bone; and another liga- ment rises up from near the point of the process to the os occipitis. 78 The Vertebra. The superior oblique processes of the vertebra dentata are large, circular, very nearly in an horizontal position, and slightly convex, to be adapted to the inferior oblique processes of the first vertebra. The inferior oblique pro- cesses of this vertebra answer exactly to the description given of those common to all the cervical vertebrae. The transverse processes of the vertebra dentata are short, very little hollowed at their upper part, and not forked at their ends; and the canals through which the cervical arteries pass are reflected outwards about the middle of each process; so that the course of these ves- sels may be directed towards the transverse processes of the first vertebra. Had this curvature of the arte- ries been made in a part so movable as the neck is, while they were not defended by a bone, and placed in a cavity of that bone, scarce a motion could have been performed without the utmost hazard of compression. This is the third instance of similar mechanism in cases of sudden curvature of arteries. The first is the passage of the caro- tids through the temporal bones; and the second is that lately described, where the vertebral arteries turn round the oblique processes of the first vertebra, to come at the great hole of the occipital bone. The spinal process of this vertebra is thick, strong, and short, to give sufficient origin to the musculi recti majores and obliqui inferiores, and to prevent the con- tusion of these and other muscles in pulling the head back. The four cervical vertebrae which are next in order, have nothing particular in their structure, but agree with the general description. The seventh vertebra approaches the form of those of the back, having the upper and lower The Vertebra. 79 surfaces less excavated than the others. The oblique pro- cesses are more perpendicular; and the spinal as well as transverse processes are without bifurcation. After an examination of the condyles of the os occipitis, and of the whole structure of the atlas and vertebra den- tata, it will be evident, that the flexion and extension of the head, or its motion backwards and forwards, is effected by the movements of the condyles of the occipital bone on the atlas; and that in the rotation of the head, the atlas re- volves to a certain degree round the processus dentatus of the second vertebra: the head necessarily moving with it. The twelve dorsal may be distinguished from the other vertebra of the spine by the following marks. Their bodies are of a middle size, between those of the neck and loins. They are more convex before than either of the other two sorts; and are flatted laterally by the pressure of the ribs, which are inserted into small cavities formed in their sides. This flatness of their sides, which makes the figure of these vertebrae almost an half oval, is of great use; as it affords a firm articulation to the ribs, allows the trachea arteria to divide at a small angle, and the other large vessels to run secure from the action of the vital organs. Their bodies are more concave be- hind than any of the other two classes. Their upper and lower surfaces are horizontal. The cartilages interposed between the bodies of these vertebrae are thinner than in any other of the true verte- brae; and contribute to the concavity of the spine in the thorax, by being thinnest at their forepart. The oblique processes are placed almost perpendicu- larly: the upper ones slanting but a little forwards, and the lower ones slanting as much backwards. Their convexity 80 The Vertebra. / or conc?.vity is not so remarkable as to require particular notice. Between the oblique processes of opposite sides, several sharp processes stand out from the upper and lower parts of the plates which join to form the spinal processes: into these sharp processes strong ligaments are fixed for connecting the vertebrae. The transverse processes of the dorsal vertebrae are long, thicker at their ends than in the middle, and turned obliquely backwards; which may be owing to the pressure of the ribs; the tubercles of which are inserted into a de- pression near the end of these processes. The spinal processes are long, small-pointed, and sloping downwards and backwards. From their upper and back part a ridge rises, which is received by a small channel in the forepart of the spinal process immediately above, which is here connected to it by a ligament. The canal for the spinal marrow is here more circular, but, corresponding to the size of that cord, is smaller than in any of the other vertebrae; and a larger share of the holes, in the bony bridges for the transmission of the nerves, is formed in the vertebra above than in the one below. The connexion of the dorsal vertebrae to the ribs, the thinness of their cartilages, the erect situation of the oblique processes, the length, sloping and connexion of the spinal processes, all contribute to restrain these verte- brae from much motion, which might disturb the actions of the heart and lungs; and in consequence of the little mo- tion allowed here, the intervertebral cartilages sooner shrivel, by becoming more solid} and therefore the first remarkable curvature of the spine observed, as people ad- vance to old age, is in the least stretched vertebrae of the back; or old people first become round-shouldered. The Vertebra. 81 The bodies of the four uppermost dorsal vertebrae devi- ate from the rule, that the vertebrae become larger as they descend; for the first of the four is the largest, and the other three below gradually become smaller, to allow the trachea and large vessels to divide at smaller angles. The two uppermost vertebrae of the back, instead of be- ing very prominent forwards, are flattened by the action of the musculi longi colli and recti majores. The proportional size of the two little depressions in the body of each vertebra for receiving the heads of the ribs, seems to vary in the following manner; the depres- sion on the upper edge of each vertebra decreases as far down as the fourth, and after that increases. The transverse processes are longer in each lower ver- tebra to the seventh or eighth, with their smooth surfaces, for the tubercles of the ribs, facing gradually more down- wards; but afterwards, as they descend, they become shorter, and the smooth surfaces are directed more up- wards. The spinous processes of the vertebrae of the back be- come gradually longer and more slanting from the first, as far down as the eighth or ninth vertebra; from which they manifestly turn shorter and more erect. The first vertebra, besides an oblong hollow in its lower edge, that assists in forming the cavity wherein the second rib is received, has the whole cavity for the head of the first rib formed in it. The eleventh often has the whole cavity for the eleventh rib in its body, and wants the smooth surface on each transverse process. The twelfth always receives the whole head of the last rib, and has no smooth surface on its transverse processes, which are very short. The smooth surfaces of its inferior Vol. I. E 82 The Vertebra. oblique processes face outwards as the lumbar do. In general, the upper vertebrae of the back lose gradually their resemblance to those of the neck, and the lower ones approach gradually to the figure of the lumbar. The lumbar vertebra are five bones, that may be dis- tinguished from any others by these marks: 1. Their bodies, though of a circular form at their forepart, are somewhat oblong from one side to the other. The epiphyses on their edges are larger; and therefore the upper and lower sur- faces of their bodies are more concave than in the verte- brae of the back. 2. The cartilages between these vertebra are very thick, and render the spine convex within the ab- domen, by their great thickness anteriorly. 3. The oblique processes are strong and deep: the superior, which are concave, facing inwards, and the convex inferior ones facing outwards: and therefore each of these vertebra receives the one above it, and is received by the one be- low; which is not so evident in the other two classes already described. 4. Their transverse processes are small, long, and almost horizontal, for allowing large motion to each bone, and sufficient insertion to muscles, and for sup- porting and defending the internal parts. 5. Between the roots of the superior oblique and transverse processes a small protuberance may be observed, where some of the muscles that raise the trunk of the body are inserted. 6. Their spinal processes are strong, straight, and hori- zontal, with broad flat sides, and a narrow edge above and below; this last being depressed on each side by muscles. And at the root of these edges, we see rough surfaces for fixing the ligaments. 7. The medullary canal is larger in these bones than in the dorsal vertebrae. 8. The holes for the passage of the nerves are more equally formed out of both the contiguous vertebrae than in the other classes; The Vertebra. 83 the upper one furnishes, however, the larger share of each hole. The thick cartilages between these lumbar vertebrae, their deep oblique processes, and their erect spinal pro- cesses, are all fit for allowing large motion: though it is not so great as what is performed in the neck; which ap- pears from comparing the arches which the head describes when moving on the neck or the loins only. The lumbar vertebrae, as they descend, have their ob- lique processes at a greater distance from each other, and facing more backwards and forwards. The transverse and spinal processes of the first and last lumbar vertebrae are shorter than those in the middle. The epiphyses round the edges of the bodies of the lumbar vertebrae are most raised in the two lowest; which consequently make them appear hollower in the middle than the others are. The body of the fifth vertebra is rather thinner than that of the fourth. The spinal process of this fifth is smaller, and the oblique processes face more backwards and for- wards, than those of any other lumbar vertebra. In consequence of this particular construction, the spine is capable of flexion, principally in an anterior and lateral direction, and also of extension. It ought to be remarked that during flexion it forms a curve, and not an angle; for, in this last case, the spinal marrow would be more or less compressed. The cervical vertebrae have most motion, and the dorsal the least. This circumstance is fully explained by the form of the different parts of these vertebrae, and by the differ- ence in the thickness of their intervertebral substance. The necessity of fixing the dorsal vertebrae is very evident: as their motion would greatly interfere with the motions of the ribs in respiration. 84 Os Sacrum. The lumbar vertebrae have more motion than is com- monly supposed; for, in addition to a certain degree of flexion, they perform a species of rotation or twisting, which is very observable in persons who are diseased in one of their hip joints; such persons move the whole pel- vis, by a rotation of the lumbar vertebrae, to avoid moving the diseased joint. False Vertebra. The lower pyramid, or under part of the spine, consists of one large triangular bone called the os sacrum, and of some small bones denominated the os coccygis. These bones are called the false vertebrae, because the sacrum in young subjects is composed of five distinct bones, each of which has some resemblance to a vertebra; but they are completely united in the adult, and form but one bone, which is supposed to have been denominated sacrum, because it was offered in sacrifice by the ancients. The os sacrum is of a triangular form, with its base up- wards. It is concave anteriorly, and convex posteriorly. The middle of the bone, when viewed anteriorly, appears to be composed of the bodies of five vertebrae, united to each other, and their union is marked by four transverse lines. At the two extremities of each of these lines, are large round holes, which communicate with the vertebral cavity of the bone. On the exterior sides of these holes the surface is free from any marks of the original separation. The middle of the upper surface, or base of the bone, is formed for articulating with the last lumbar vertebra, and has two oblique processes, with a groove on each side, which forms part of the foramen for transmitting the twenty-fourth pair of nerves. Os Sacrum. 85 The back part of the os sacrum is rough and convex: in the middle there are commonly three processes similar to the spinous processes of the lumbar vertebrae, and a fourth which is much smaller. Below this, there is a deficiency of the bony spine, and the vertebral cavity is consequently open behind; but the sides of the canal continue lower down. On each side of the spinous processes are four smaller holes, which are opposite to the larger holes on the ante- rior surface. Between the spinous processes and the ante- rior part, which resembles the bodies of vertebrae, is the continuation of the vertebral cavity which contains the spi- nal marrow. From the cauda equina, contained in this cavity, the great nerves of the lower extremities pass off, through the large holes on the anterior surface, and some small nerves through the posterior holes. In some bones the spinous processes are entirely defi- cient, and the cavity above mentioned is completely open behind; but the contained parts are defended by strong membranes. The anterior part of each lateral surface is covered by a plate of cartilage, and articulated to the os ilium. The posterior part is rough and perforated by the fibres of the strong ligaments, which are inserted into it. On the posterior surface of the sacrum, the sides of the open part of the vertebral canal terminate, so as to form a notch, through which passes the twenty-ninth pair of nerves. The os sacrum is very spongy, and is lighter in propor- tion to its bulk than any bone in the body: it is defended by the muscles that cover it and the ligaments which ad- here to it. It is articulated, above, to the last lumbar vertebra; be- low, to the os coccygis; and on the sides, to the ossa ilia. 86 Os Coccygis. That triangular chain of bones depending from the os sacrum, in which each bone becomes smaller as it de- scends, till the last ends in a small tubercle, is called os coccygis. It is convex behind, and concave before; from which crooked pyramidal figure, which was thought to resemble a cuckow's beak, the name is derived. There are four pieces in people of middle age. In chil- dren, they are almost wholly cartilaginous. In old subjects, all the bones are united, and become frequently one con- tinued bone with the os sacrum. The highest of the four bones is the largest, with shoul- ders extended farther to each side than the end of the os sacrum; which enlargement may serve as a distinguishing mark to fix the limits of either bone. The upper surface of this bone is a little hollow. From the back of that bulbous part called its shoulders, a process often rises up on each side, to join with the os sacrum. Sometimes these shoul- ders are joined to the sides of the open end of the verte- bral canal', to form the hole in each side common to these two bones, for the passage of the twenty-ninth pair of spi- nal nerves. Immediately below the shoulders of the os coccygis, a notch may be remarked on each side, where the thirtieth pair of the spinal nerves passes. The lower end of this bone is formed into a small head, which very often is hollow in the middle. The three lower bones gradually become smaller, and are spongy, but are strengthened by a strong ligament which covers and connects them. Their ends, by which they are articulated, are formed in the same manner as those of the first bone. Between each of these four bones of young subjects a cartilage is interposed; therefore their articulation is ana- logous to that of the bodies of the vertebras of the neck: Os Coccygis. 87 for the lower end of the os sacrum, and of each of the three superior bones of the os coccygis, has a small de- pression in the middle; and the upper part of all the bones of the os coccygis is a little concave, and consequently the interposed cartilages are thickest in the middle, to fill up both cavities; by which they connect the bones more firmly. When the cartilages ossify, the upper end of each bone is formed into a cavity, exactly adapted to the protuberant lower end of the bone immediately above. From this sort of articulation, it is evident, that, unless when these bones grow together, all of them are capable of motion; of which the first and second enjoy the largest share. The lower end of the fourth bone terminates in a rough point, to which a cartilage is appended. To the sides of these bones of the os coccygis, the coc- cygaei muscles, and part of the levatores ani, and of the glutaei maximi, are fixed. The connexions of these bones hinder them from being moved to either side; and their motion backwards and forwards is much confined: yet as their ligaments can be stretched by a considerable force, it is of great advantage in the excretion of the faeces alvinae, and much more in childbearing, that these bones should remain movable; and the right management of them, in delivering women, is very important. The mobility of the os coccygis dimin- ishing as people advance in age, especially when its liga- ments and cartilages have not been kept flexible by being stretched, is probably one reason why women, who are advanced in years before they marry, have generally diffi- cult parturition. These bones serve to sustain the intestinum rectum; and, therefore, are curved forwards; by which they are preserved, as well as the muscles and teguments, from any injury when sitting with the body reclined back. 88 The Ribs. The Vertebral Cavity for containing the Spinal Marrow. The canal formed by the foramina of the different ver- tebrae, when these bones are placed in their natural order, extends from the great occipital foramen to the end of the sacrum. Its direction varies with the different curvatures of the spine, and its figure and diameter are also very dif- ferent in different places. In the cervical vertebrae, it is largest, and nearly trian- gular in form; in the dorsal, it is much smaller and almost cylindrical; in the lumbar, it is somewhat enlarged, and ap- proaches again to the triangular figure; in the sacrum, it is broad, but flat, and diminishes gradually, so as to assume the form of a long triangle. It has a ligamentous lining, which will be described when an account is given of the fresh bones and their liga- ments. The Thorax. The thorax resembles a flattened cone, cut away oblique- ly at its basis; and regularly truncated at its apex. It is formed by the dorsal vertebrae behind, the ribs on the sides, and the sternum before. The Ribs, Are long crooked bones, placed in an oblique direc- tion downwards as respects the backbone. Their number is generally twelve on each side; though sometimes eleven or thirteen have been found. They are convex externally and concave internally. They are made smooth by the action of the contained parts, which, on this account, are in no danger of being hurt bv them. The Ribs, 89 The ribs approach towards a round form at their extre- mities near the vertebrae. Further forwards they are flat and broad, and have an upper and lower edge; each of which is made rough by the action of the intercostal mus- cles inserted into them. These muscles, being all of nearly equal force, and equally stretched in the interstices of the ribs, prevent the broken ends of these bones, in a fracture, from being removed far out of their natural place, to inter- rupt the motion of the vital organs. The upper edge of the ribs is more obtuse and rounder than the lower, which is depressed on its internal side by a long fossa, for lodg- ing the intercostal vessels and nerves: on each side of which there is a ridge, to which the intercostal muscles are fixed. The fossa is not observable at the ends of the ribs; for, at the posterior, or root, the vessels have not yet reached the bones; and, at the fore end, they are split away into branches, to serve the parts between the ribs. From this situation of the bloodvessels, has originated the rule adopted by surgeons, that the incision in cases of empyema, &c. should be made midway between the spine and sternum, and that the lower edge of the upper rib should be avoided. At the posterior end of each rib, a little head is formed, which is divided by a middle ridge into two flat or hol- low surfaces; the lowest of which is generally the broad- est and deepest. The two surfaces are joined to the bodies of two different vertebrae, and the ridge forces itself into the intervening cartilage. A little way from this head, we find, on the external surface, a small cavity, where mucila- ginous glands are lodged; and round the head, the bone appears spongy, where the capsular ligament of the articu- lation is fixed. Immediately beyond this, a flatted tubercle rises, with a small cavity at its root, which is surrounded Vol. I. M 90 The Ribs. by a roughness, for the articulation of the rib with the transverse process of the lowest of the two vertebrae, with which the head of the rib is joined. Advancing further on this external surface, another smaller tubercle may be ob- served in most cases, into which, ligaments connecting the ribs to each other, and to the transverse processes of the vertebrae and portions of the longissimus dorsi, are insert" ed. Beyond this, these bones are made flat by the sacro- lumbalis muscle, which is inserted into the part of this flat surface furthest from the spine, where each rib makes a considerable curve, called by some its angle. Then the rib begins to turn broad, and continues so to its anterior end, which is hollow and spongy, for the reception of, and firm coalition with, the cartilage that runs thence to be inserted into the sternum, or to be joined with some other carti- lage. In adults, the cavity at this end of the ribs is gene- rally smooth. The substance of the ribs is spongy, cellular, and only covered with a very thin external lamellated surface, which increases in thickness and strength as it approaches the vertebrae. To the fore end of each rib a long broad and strong cartilage is fixed, which reaches to the sternum, or is joined to the cartilage of the next fib. This course, how- ever, is not in a straight line with the rib: for the cartilages generally make a considerable flexure, the concave part of which is upwards; therefore, at their insertion into the sternum, they make an obtuse angle above, and an acute one below. These cartilages are of such a length as never to allow the ribs to come to a right angle with the spine; but they keep them situated so obliquely as to make the angle very considerably obtuse above, till a force exceed- ing the elasticity of the cartilage is applied. These cartila- The Ribs. 91 ges, as all others, are firmer and harder internally than they are on their external surface; and sometimes, in old peo- ple, all their middle substance becomes bony, while a thin cartilaginous lamella appears externally. The ossification, however, begins frequently at the external surface. The greatest alternate motions of the cartilages being made at their great curvature, that part remains frequently cartila- ginous after all the rest is ossified. The ribs then are articulated at each end, and that be- hind is doubly joined to the vertebrae; for the head is received into the cavities of two bodies of the vertebrae, and the larger tubercle is received into the depression in the transverse process of the lower vertebra. When we examine the double articulation, we must immediately see, that no other motion can here be allowed than upwards and downwards. Since the transverse process hinders the rib to be thrust back, the resistance of the sternum on the other side prevents the ribs coming forward; and each of the two joints, with the other parts attached, oppose its turning round. But then it is likewise as evident, that even the motion upwards and downwards can be but small in any one rib at the articulation itself. But as the ribs ad- vance forwards, the distance from their center of motion increasing, the motion must be larger; and it would be very conspicuous at their anterior ends, were they not re- sisted there by the cartilages, which yield so little, that the principal motion is performed by the middle part of the ribs, which turns outwards and upwards, and occasions the twist remarkable in the long ribs at the place near their fore end where they are most resisted. The ribs differ from each other in the following respects: The upper rib is the most crooked; and as they descend they become straight. Their obliquity, with respect to the 92 The Ribs. spine, increases as they descend; so that though their dis- tances from each other are nearly equal at their back part, yet at their fore ends the distances between the lower ribs must increase. In consequence of this increased obliquity of the lower ribs, each of their cartilages makes a greater curve in its progress from the rib towards the sternum; and the tubercles, that are articulated to the transverse processes of the vertebrae, have their smooth surfaces gra- dually facing more upwards. The ribs becoming thus more oblique, while the sternum advances forwards in its de- scent, makes the distance between the sternum and the anterior end of the lower ribs greater than between the sternum and the ribs above; consequently, the cartilages of those ribs that are joined to the breast bone are longer in the lower than in the higher ones. These cartilages are placed nearer to each other as the ribs descend, which oc- casions their curvature to be greater. The length of the ribs increases from the first and up- permost rib, as far down as the seventh; and from that to the twelfth, it gradually diminishes. The superior of the two surfaces, by which the ribs are articulated to the bo- dies of the vertebrae, gradually increases from the first to the fourth rib, and is diminished after that in each lower rib. The distance of their angles from the heads always increases as they descend to the ninth, because of the greater breadth of the sacro-lumbalis muscle. The ribs are commonly divided into true and false. The true ribs are the seven uppermost of each side. Their cartilages are all gradually longer as they descend, and are joined to the breast bone: so that, being pressed constantly between two bones, they are flatted at both ends; and are thicker, harder, and more liable to ossify, than the other cartilages that are not subject to so much pressure. The Ribs. 93 These bones include the heart and lungs; and therefore are called true ribs. The five inferior ribs of each side are the false, whose cartilages do not reach to the sternum; but on this account, having less pressure, their substance is softer. To these five ribs the circular edge of the diaphragm is connected. The first rib of each side is so situated, that the flat sides are above and below, while one edge is placed inwards, and the other outwards, or nearly so: therefore sufficient space is left above it for the subclavian vessels and muscles; and the broad concave surface is opposed to the lungs. But in consequence of this situation, the chan- nel for the intercostal vessels is not to be found. The head of this rib is not divided into two plane surfaces by a mid- dle ridge, because it is only articulated with the first ver- tebra of the thorax. Its cartilage is ossified in adults, and is united to the sternum at right angles. This first rib fre- quently has a ridge rising near the middle of its posterior edge, where one of the heads of the scalenus muscles rises. Farther forward it is flatted, or sometimes depressed by the clavicle. The position of the second rib is such that its two broad surfaces have oblique aspects, inward and downward, out- wards and upwards, so as to make the surface of the tho- rax uniform: and it may be observed of all the ribs, that the aspect of their surfaces is varied upon this principle, according to their situation in the thorax. The sixth, seventh, and eighth ribs, have their cartilages nearly contiguous. They are frequently joined to each other by cross cartilages; and frequently the cartilages of the eighth, ninth, and tenth, are connected to the former, and to each other by firm ligaments. The eleventh, and sometimes the tenth rib, has no uv 94 The Sternum. bercle for its articulation with the transverse process of the vertebra, to which it is only loosely fixed by ligaments. The fossa in its lower edge is not so deep as in the upper ribs; because the vessels run more towards the interstice between the ribs. Its front end is smaller than its body; and its short small cartilage is but loosely connected to the cartilage of the rib above. The twelfth rib is the shortest and straightestu Its head is only articulated with the last vertebra of the thorax; therefore is not divided into two surfaces. This rib is not joined to the transverse process of the vertebra; and there- fore has no tubercle, being often pulled necessarily inwards by the diaphragm, which an articulation with the trans- verse process would not have allowed. The fossa is not found at its under edge,, because the vessels run below it. The forepart of this rib is smaller than its middle, and has -only a very small pointed cartilage fixed to it. To its whole internal side the diaphragm is connected. The Sternum is the broad flat bone, in the front part of the thorax. In adults it is composed of three pieces, which easily separate after the cartilages connecting them are destroyed. The two lower pieces are frequently found intimately united; and very often, in old people, the sternum is a continued bony substance from one end to the other; though we still observe two, sometimes three, transverse lines on its sur- face; which are marks of the former divisions. The sternum, considered as one bone, is broadest and thickest above, and smaller as it descends. The internal surface of this bone is somewhat concave for enlarging the ro\vs to the two bones at both its internal and external The Clavicle. 109 end: its substance at the external end being soft, but very strong, and resembling the intervertebral cartilages. From its internal end, the clavicle, for about two fifths of its length, is bended obliquely forwards. On the upper and front part of this curvature a small ridge is seen, with a plain rough surface before it; whence the sterno- hyoideus and sterno-mastoideus muscles have in part their origin. Near the lower angle, a small plain surface is often to be remarked, where the first rib and this bone are con- tiguous, and are connected by a firm ligament. From this a rough plain surface is extended outwards, where the pectoral muscle has part of its origin. Behind, the bone is made flat and rough by the insertion of the larger share of the subclavian muscle. The clavicle is then curved back- wards, and at first is round: but it soon after becomes broad and thin; which shape it retains to its external end. Along the external concavity, a rough sinuosity runs; from which some part of the deltoid muscle takes its rise: op- posite to this, on the convex edge, a scabrous ridge gives insertion to a share of the cucullaris muscle. The upper surface of the clavicle is here flat; but the lower is hollow, for lodging the beginning of the musculus subclavius; and towards its back part, a tubercle rises; to which, and to a roughness near it, the strong short thick ligament, con- necting this bone to the coracoid process of the scapula, is fixed. The external end of this bone is oblong horizontally, smooth, sloping at the posterior side, and tipped in a re- cent subject with a cartilage, for its articulation with the acromion scapulae. Round this the bone is spongy, for the firmer connexion of the ligaments. The surfaces of contact of this bone, and the scapula, are remarkably small, and flat also. The medullary arteries, having their direction obliquely 1 lo The Scapula. outwards, enter the clavicles by one or more small passa- ges in the middle of their back part. The substance of this bone is the same as of the other round long bones. The ligaments, which surround the articulation of this bone with the sternum, are so short and strong, that little motion can be allowed any way; and the strong ligament that is stretched across the upper furcula of the sternum, from the posterior prominent angle of the one clavicle to the same place of the other clavicle, serves to keep each of these bones more firmly in its place. By the assistance, however, of the movable intervening cartilage, the clavicle can move at this articulation so that the external extremity may be elevated or depressed, and moved backwards or forwards. The whole bone may be moved so as to describe a cone; of which the end at the sternum is the apex The movements of the scapula and arm are the objects of these motions of the clavicle; and the general use of the bone is to regulate the motions of these parts. From the situation, figure, and use of the clavicles, it is evident that they are much exposed to fractures; that their broken parts must generally pass each other; and, that they will be kept in their places with difficulty. The Scapula, Or shoulderblade, is the triangular bone situated on the upper and back part of the thorax. The back part of the scapula has nothing but the thin ends of the serratus ami- cus major, and subscapularis muscles between it and the ribs: but as this bone advances forwards, its distance from the ribs increases. The longest side of this bone is nearest the spine, and has an oblique position as respects it. The upper or shortest side, called the superior costa of the sca- pula, is nearly horizontal, and parallel with the second rib. The Scapula. Ill The lower side, which is named the inferior costa, is ex- tended obliquely from the third to the eighth rib. The situation of this bone, here described, is when people are sitting or standing in a state of inactivity, and allowing the members to remain in the most natural easy posture. The inferior angle of the scapula is very acute; the upper one is near to a right angle; and what is called the anterior does not deserve the name, for the two sides do not meet to form an angle. The body of this bone is concave towards the ribs, and convex behind, where it has the name of dorsum. Three processes are generally reckoned to proceed from the scapula. The first is the large spine that rises from its convex surface behind, and divides it une- qually. The second process stands out from the fore part of the upper side; and, from its imaginary resemblance to a crow's beak, is named coracoides. The third process is the whole thick bulbous fore part of the bone. Into the oblique space the musculus patientiae is insert- ed. At the root of the spine, on the back part of the base, a triangular flat surface is formed by the pressure of the lower fibres of the trapezius. Below this, the edge of the scapula is scabrous and rough, for the insertion of the ser- ratus major anticus and rhomboid muscles. The back part of the inferior angle is made smooth by the latissimus dorsi passing over it. This muscle also alters the direction of the inferior costa some way forwards from this angle: and so far it is flatted behind by the ori- gin of the teres major. As the inferior costa advances for- ward, it is of considerable thickness, is slightly hollowed, and made smooth behind, by the teres minor; while it has a fossa formed into it below, by part of the subscapulars; and between the two, a ridge with a small depression ap- pears, where the longus extensor cubiti has its origin. 112 The Scapula. The superior costa is very thin: and near its fore part there is a semilunar notch, from one end of which to the other, a ligament is stretched; and sometimes the bone is continued to form one, or sometimes two holes, for the passage of the scapular bloodvessels and nerves. Imme- diately behind this semilunar cavity, the coraco-hyoideus muscle has its rise. From the notch, to the termination of the fossa for the teres minor, the scapula is narrower than any where else, and supports the third process. This part has the name ot cervix. The whole dorsum of the scapula is always said to be convex; but, by reason of the raised edges that surround it, it is divided into two cavities by the spine, which is stretched from behind forwards, much nearer to the supe- rior than to the inferior costa. The cavity above the spine is really concave, where the supra-spinatus muscle is lodg- ed; while the surface of this bone below the spine, on which the infra-spinatus muscle is placed, is convex, ex- cept a fossa that runs at the side of the inferior costa. The internal or anterior.surface of this bone is hollow, except in the part above the spine, which is convex. The subscapulars muscle is extended over this surface, where it forms several ridges and intermediate depressions, com- monly mistaken for prints of the ribs: they point out the interstices of the bundles of fibres of which the subscapu- laris muscle is composed. The spine rises small at the base of the scapula, and be- comes higher and broader as it advances forwards. On the sides it is unequally hollowed and crooked, by the ac- tions of the adjacent muscles. Its ridge is divided into two rough flat surfaces: into the upper one, the trapezius muscle is inserted; and the lower one has part of the del- toid fixed to it. The end of the spine, called acromion, or The Scapula. 113 top of the shoulder, is broad and flat, and is sometimes only joined to the spine by a cartilage. The anterior edge of the acromion is flat, smooth, and covered with a car- tilage, for its articulation with the external end of the cla- vicle; and it is hollowed below, to allow a passage to the infra and supra spinati muscles, and free motion to the os . humeri. The coracoid process is crooked, with its point inclining forwards; so that a hollow is left at the lower side of its root, for the passage of the subscapularis muscle. The end of this process is marked with three plain surfaces. Into the internal, the serratus minor amicus is inserted; from the external, one head of the biceps flexor cubiti rises; and from the lower one, the coraco-brachialis has its origin. At the upper part of the root of this process, immediately before the semilunar cavity, a smooth tuber- cle appears, where a ligament from the clavicle is fixed. From the whole of the external side of this coracoid apophysis a broad ligament goes out, which becomes nar- rower where it is fixed to the acromion. From the cervix scapulae the third process is produced. The fore part of this is formed into a glenoid cavity, which is of the shape of the longitudinal section of an egg, being broad below and narrow above. Between the margin of this cavity and the fore part of the root of the spine, a large sinuosity is left for the transmission of the supra and infra spinati muscles; and on the upper part of this margin we may remark a smooth surface, where the second head of the biceps flexor cubiti has its origin. The root of the margin is rough all round, for the firmer adhesion of the capsular ligament of the articulation, and of the cartilage; which is thick on the margin, but becomes very thin as it Vol. I. P 114 The Scapula. is continued towards the middle of the cavity, which it lines all over. The medullary vessels enter the scapula near the base of the spine. The substance of the scapula, as in all other broad flat bones, is cellular, but of an unequal thickness: for the neck and third process are thick and strong; the inferior costa, spine, and coracoid process, are of a middle thickness; and the body is so pressed by the muscles, as to become thin and transparent. The scapula and clavicle are joined by plain surfaces, tipped with cartilage; by which neither bone is allowed any considerable motion, being tightly tied down by the common capsular ligament, and by a very strong one which proceeds from the coracoid process; but divides into two before it is fixed into the clavicle, with such a direction, as can either allow this bone to have a small rotation, in which its posterior edge turns more backwards, while the anterior one rises farther forwards; or it can yield to the fore part of the scapula moving downwards, while the back part of it is drawn upwards; in both which cases, the oblong smooth articulated surfaces of the clavi- cle and scapula are not in the same plain, but stand a little transversely, or across each other, and thereby preserve this joint from luxations, to which it would be subject if cither of the bones was to move on the other perpendicu- larly up and down, without any rotation. Sometimes a movable ligamentous cartilage is found in this joint; and sometimes such a cartilage is only interposed at the ante- rior half of it; and in some old subjects a sesamoid bone has been found here. The scapula is connected to the head, os hyoides verte- brae, ribs, and arm bone, by muscles, that have one end The Os Humeri. 115 fastened to these parts, and the other to the scapula, which can move it upwards, downwards, backwards, or forwards; fey the quick succession of these motions, its whole body is carried in a circle. But being also often moved as upon an axis perpendicular to its plain, its circumference turns in a circle whose center this axis is. Whichever of these motions it performs, it always carries the outer end of the clavicle and the arm along with it. The glenoid cavity of this bone receives the os humeri, which plays in it, as will be more fully explained hereafter. The use of the scapula is, to serve as a fulcrum to the arm; and, by altering its position on different occasions, to allow always to the head of the os humeri a socket to move in properly situated; and thereby, to assist and to enlarge greatly the motions of the superior extremity, and to afford the muscles which rise from it more advanta- geous actions, by altering their directions with respect to the bone which they are to move. This bone also serves to defend the back part of the thorax, and is often em- ployed to sustain weights, or to resist forces, too great for the arm to bear. The Arm. The arm has only one bone, best known by the Latin name of os humeri; which is long, round, and nearly straight. The upper end of this bone consists of a large round smooth head, which forms the segment of a sphere, whose axis is not in a straight line with the axis of the bone, but stands obliquely backwards from it. The extent of the head is distinguished by a circular fossa surrounding its base, where the head is united to the bone, and the capsu- lar ligament of the joint is fixed. Below the fore part of 116 The Os Humeri. its base, two tubercles stand out: the smallest one, which is situated most to the inside, has the tendon of the sub- scapulars muscle inserted into it. The larger more exter- nal protuberance is divided, at its upper part, into three smooth plain surfaces: into the anterior of which, the musculus supra-spinatus; into the middle or largest, the infra-spinatus; and into the one behind, the teres minor, is inserted. Between these two tubercles, exactly in the fore part of the bone, a deep long groove is formed, for lodging the tendinous head of the biceps flexor cubiti; which, after passing, in a manner peculiar to itself, through the cavity of the articulation, is tied down, by a tendinous sheath extended across the groove; in which, and in the neighbouring tubercles, are several remarkable holes, which are penetrated by the tendinous and ligamentous fibres, and by vessels. On each side of this groove, as it descends in the os humeri, a rough ridge, gently flatted in the middle, runs from the roots of the tubercles. The ten- don of the pectoral muscle is fixed into the anterior of these ridges, and the latissimus dorsi and teres major are inserted into the internal one. A little behind the lower end of this last, another rough ridge may be observed, where the coraco-brachialis is inserted. From the back part of the root of the largest tubercle, a ridge also is con- tinued; from which the brevis extensor cubiti rises. This bone is flatted on the inside, about its middle, by the belly of the biceps flexor cubiti. In the middle of this plain sur- face, the entry of the medullary artery is seen slanting ob- liquely downwards. At the fore side of this plain the bone rises in a sort of ridge, which is rough, and often has a great many small holes in it, where the strong deltoid muscle is inserted; on each side of which the bone is smooth and flat, where the brachius internus rises. The The Os Humeri. 117 exterior of these two flat surfaces is the largest: behind it a superficial spiral channel, formed by the muscular nerve and the vessels that accompany it, runs from behind for- wards and downwards. The body of the os humeri is flatted behind by the ex- tensors of the fore arm. Near the lower end of this bone, a large sharp ridge is extended on its outside; from which the musculus spinator radii longus, and the longest head of the extensor carpi radialis, rise. Opposite to this, there is another small ridge to which the aponeurotic tendon, that gives origin to the fibres of the internal and external brachial muscles, is fix- ed; and from a little depression on the fore side of it, the pronator radii teres rises. The body of the os humeri becomes gradually broader towards the lower end, where it has several processes; at the roots of which there is a cavity before, and another behind. The anterior is divided by a ridge into two; the external, which is the least, receives the end of the radius; and the internal receives the coronoid process of the ulna, in the flexions of the fore arm; while, the posterior deep triangular cavity lodges the olecranon in the extensions of that limb. The bone between these two cavities is pressed so thin by the processes of the ulna, as to appear transpa- rent in many subjects. The sides of the posterior cavity are stretched out into two processes, one on each side: These are called condyles; from each of which a strong ligament goes out to the bones of the fore arm. The exter- nal condyle, which has an oblique direction forwards with respect to the internal, when the arm is in the most natu- ral posture, is equally broad, and has an obtuse smooth head rising from it forwards. From the rough part of the condyle, several muscles arise; and #n the smooth head 118 The Os Humeri. the upper end of the radius plays. The internal condyle is more pointed and protuberant than the external, to give origin to the flexor muscles of the wrist and hand, &c. Between the two condyles, is the trochlea or pulley; which consists of two lateral protuberances, and a middle cavity, that are smooth and covered with cartilage. When the fore arm is extended, the tendon of the internal brachiaeus muscle is lodged in the fore part of the cavity of this pul- ley. The external protuberance, which is less than the other, has a sharp edge behind; but forwards, this ridge is obtuse, and only separated from the little head, already described, by a small fossa, in which the adjoining edges of the ulna and radius move. The internal protuberance of the pulley is largest and highest; and, therefore, in the motions of the ulna upon it, that bone would be inclined outwards, were it not supported by the radius on that side. Between this internal protuberance and condyle, a sinuo- sity may be remarked, where the ulnar nerve passes. The substance and the internal structure of the os hu- meri are the same, and disposed in the same way,as in other long bones. The round head, at the upper end of this bone, is articu- lated with the glenoid cavity of the scapula; which being superficial, and having long ligaments, allows the arm a free and extensive motion. These ligaments are, however, considerably strong. For besides the common capsular ligament, the tendons of the muscles perform the office, and have been described under the name of ligaments. Then the acromion and coracoid process, with the strong broad ligaments stretched between them, secure the articu- lation above, where the greatest and most frequent force is applied to thrust the head of the bone out of its place. It is true, that there is not near so strong a defence at the The Os Humeri. 119 lower part of the articulation; but, in the ordinary pos- tures of the"arm, that is, so long as it is at an acute angle with the trunk of the body, there cannot be any force ap- plied at this place to occasion a luxation, since the joint is protected so well above. The motions which the arm enjoys by this articulation, are to every side; and by the succession of these different motions, a circle may be described. Besides which, the bohe performs a small rotation round its own axis; but when the axis of the bone is the center of motion, the movements are very different from those which take place when the axis of its head is the center; for the axis of the head forms a very large angle with the axis of the body of the bone. Thus, when the arm swings backwards and for- wards, the axis of the head is the center of motion; but when the elbow is bent, and the fore arm forms a right angle with the os humeri, the motion which applies the fore arm to the thorax, or removes it, is a rotation of this bone on its axis. Though the motions of the arm seem to be very exten- sive, yet the larger share of them depends on the motions of the scapula; for the surface of the glenoid cavity is directed upwards or downwards, and to a certain degree backwards or forwards, to support the head of the os hu- meri, This is exemplified when we press the hand against a body which is before, or above, or to one side of us. The lower end of the os humeri is articulated to the bones of the fore arm, and carries them with it in all its motions j but serves as a base, on which they perform the motions peculiar to themselves; as will shortly be descri- bed. 120 The Fore Arm.—Ulna. The Fore Arm Consists of two bones, one of which is called ulna, from its being used as a measure; and the other radius, from a supposed resemblance to the spoke of a wheel. These bones are concerned in very different operations. The ulna forms the elbow joint with the os humeri: the radius is the movable basis of the hand. Ulna. The length of this bone is equal to the fore arm, of which it is a part. It is thickest above, and gradually di- minishes until near its lower end. The body of the bone is nearly triangular in form. At the upper extremity of the ulna, on its anterior surface, is a semicircular notch. The end of the bone which forms the posterior part of this notch is denominated olecranon. The anterior part of the notch is formed by a process called coronoid. This notch applies to the pulley-like surface on the internal side of the lower extremity of the os humeri, to form the ar- ticulation of the elbow. In the middle of the concave sur- face is a ridge, in consequence of which a small rocking motion is performed by the ulna. The external surface of the olecranon is rough, and strongly marked. The extensor muscle of the fore arm is inserted into the end of it, and below this is a flat surface on which we lean. On the out- side of the coronoid process is a semilunated smooth ca- vity, lined with cartilage; in which, and in a ligament ex- tended from the one to the other end of this cavity, the round head of the radius plays. Immediately below it, a rough hollow gives lodging to mucilaginous glands. Be- low the root of the coronoid process, this bone is scabrous and unequal, where the brachiaeus internus is inserted. On Ulna. 121 the outside of that, we observe a smooth concavity, where the beginning of the flexor digitorum profundus sprouts out. The external angle of the triangular part of the ulna is very sharp, where the ligament that connects the two bones is fixed: the sides which make this angle are flat and rough, by the action and adhesion of the many muscles which are situated here. At the distance of one third of the length of the ulna from the top, in its fore part, the passage of the medullary vessels may be seen slanting up- wards. The internal side of this bone is smooth, some- what convex, and the angles at each edge of it are blunted by the pressure of the muscles equally disposed about them. As this bone descends, it becomes gradually smaller; so that its lower end terminates in a little head, standing on a small neck; towards the inner and back part of which last, an oblique ridge runs, that gives rise to the pronator radii quadratus. The head is somewhat cylindrical, smooth, and covered with a cartilage on its external side, to be received into the semilunar cavity of the radius; while a styloid process rises from its inside, to which is fixed a strong ligament that is extended to the os cunei- forme and pisiforme of the wrist. At the root of the pro- cess, the end of the bone is smooth, and covered with a cartilage. Between it and the bones of the wrist, a doubly concave movable cartilage is interposed; which is a con- tinuation of the cartilage that covers the lower end of the radius, and is connected loosely to the root of the styloid process, and to the rough cavity there; in which mucila- ginous glands are lodged. The ulna is principally concerned in the articulation with the os humeri, and forms a hinge-like joint, which Vol. I. Q 122 Radius. allows extension nearly to a straight line, and flexion to an acute angle. By the sloping of the pulley-like surface, the lower part of the arm is turned outwards in the extension, and inwards in the flexion; which greatly facilitates the motion of the hand towards the head. Radius. Before the radius is described, it is necessary to ob- serve, that the lower end of this bone occasionally revolves half round the lower end of the ulna, and the hand with it. The relative situation of these parts is therefore different in different positions of the hand. In the following description, the palm of the hand is supposed to present forwards, and the thumb outwards; in which case, the two bones of the fore arm will be parallel to each other. The radius is situated on the outside of the fore arm, and is rather shorter than the ulna. Its extremities are the reverse of those of the ulna in their proportionate size; and the body is not so triangular, although it approaches towards that form. Its upper end is formed into a cylindrical head, which is hollowed on the top for an articulation with the tubercle at the side of the pulley of the os humeri; and the half cylindrical circumference next to the ulna is smooth, and covered with a cartilage, in order to be received into the semilunated cavity of that bone. Below the head, the radius is much smaller; and therefore this part is named its cervix. At the external root of this neck is a flat tuber- cle, into the outer part of which the biceps flexor cubiti is inserted. From this a ridge runs downwards and inwards, where the supinator radii brevis is inserted; and a little below, and behind this ridge, there is a rough scabrous surface, where the pronator radii teres is fixed. Radius. 123 The body of the radius is not straight, but curved ex- ternally, the greater part of its length. Its external surface is rounded; the anterior and posterior surfaces are flatten- ed; and between them is a sharp spine, to which the strong ligament extended between the two bones of the fore arm is fixed. On the anterior surface, at a distance from its head nearly equal to one third of the length of the bone, is the orifice of the canal for the medullary vessels, which has a direction obliquelv upwards. Towards the lower end, the radius becomes broader and flatter, especially on its forepart, where the pronator quadratus muscle is situated. Its back part, at this end, has a flat strong ridge in the middle, and fossae on each side. In a small groove, immediately on the inside of the ridge, the tendon of the extensor of the last joint of the thumb plays. In a large one, inside of this, the tendons of the indicator, and of the common extensor muscles of the fingers pass. On the outside of the ridge there is a broad depression, which seems again subdivided, where the two tendons of the extensor carpi radialis are lodged. The external side of this end of the radius is also hollowed by the extensors of the first and second joint of the thumb. The ridges at the sides of the grooves, in which the tendons play, have an annular ligament fixed to them, by which the several sheaths for the tendons are formed. The fore part of this end of the radius is also de- pressed, where the flexors of the fingers and flexor carpi radialis pass. The internal side is formed into a semilu- nated smooth cavity, lined with a cartilage, for receiving the lower end of the ulna. The lowest part of the radius is formed into an oblong cavity; in the middle of which is a small transverse rising, gently hollowed, for lodging mu- cilaginous glands; while the rising itself is insinuated into 124 Radius. the conjunction of the two bones of the wrist that are received into the cavity. The external side of this articu- lation is defended by a remarkable process of the radius, from which a ligament passes to the wrist; and this struc- ture resembles that of the styloid process of the ulna with its ligament. The ends of both the bones of the fore arm being thicker than the middle, and the radius being curved, there is a considerable distance between the bodies of these bones; in the larger part of which a strong tendinous, but thin ligament, is extended, to give a sufficient surface for the origin of the numerous fibres of the muscles situated here, that are so much sunk between the bones as to be protected from injuries, to which they would otherwise be exposed. But this ligament is wanting near the upper end of the fore arm, where the supinator radii brevis, and flexor digitorum profundus, are immediately connected. As the head of the radius receives the tubercle of the os humeri, it is not only bended and extended along with the ulna, but may be moved almost half round its axis: and, that this motion round its axis may be sufficiently large, the ligament of the articulation is extended," further down than ordinary, on the neck of this bone, before it is connected to it; and it is very thin at its upper and lower part, but makes a firm ring in the middle. This bone is also joined to the ulna by a double articulation: for above, a tubercle of the radius plays in a socket of the ulna; whilst below, the radius gives the socket, and the ulna the tuber- cle. But then the motion performed at the two ends is very different: for, at the upper end, the radius does little more than turn round its axis; while, at the lower end, it moves nearly half round the cylindrical end of the ulna; and, as the hand is articulated and firmly connected here with the The Haiid. 125 radius, they must move together. When the palm is turn- ed uppermost, the radius is said to perform supination: when the back of the hand is above, it is said to be prone. But then the quickness and large extent of these two mo- tions are assisted by the ulna, which, as was before ob- served, can move with a kind of small rotation on the slo- ping sides of the pulley. This rocking motion, though very inconsiderable in the elbow joint itself, is conspicuous at the lower end of such a long bone; and the strong liga- ment connecting this lower end to the carpus, makes the hand more readily obey these motions. The Hand. The hand comprehends the whole structure, from the end of the radius to the points of the fingers. Its back part is convex, for greater firmness and strength; and it is con- cave before, for containing more surely and conveniently such bodies as we take hold of. One half of the hand has an obscure motion in comparison of what the other has: it serves as a base to the movable half, which can be extended back very little farther than to a straight line with the fore arm, but can be considerably bent forwards. The hand consists of the carpus or wrist; metacarpus, or part adjoining the wrist; and the fingers, among which the thumb is reckoned. Carpus. No part of the skeleton is more complex than the car- pus. The following description will therefore be of little use to a young student, unless the bones are before him when he is reading it. Great advantage will be derived from examining two sets of carpal bones; each set belonging to the same side. 126 Carpus. In one of these sets the bones should be connected by their natural ligaments; but the two rows separated from each other. The bones of the other set should be accurately cleaned, so that their forms and sur- faces may be examined. The carpus is composed of eight small bones, arranged in two rows; one of which rows is attached to the bones of the fore arm, and the other to the body of the hand. These bones are named from their figure, and shall be mentioned in the order in which they occur, beginning with the row next to the fore arm; and with the external bone in each row. They are, Os Scaphoides, Lunare, Cuneiforme, Pisiforme. Os Trapezium, Trapezoides, Magnum and Unciforme. First Row. Os scaphoides is the largest of the eight, excepting one. It is convex above, concave and oblong below; from which small resemblance to a boat, it has got its name. Its smooth convex surface is divided by a rough middle fossa, which runs obliquely across it. The upper largest division is arti- culated with the radius. The common ligament of the joint of the wrist is fixed into the fossa; and the lower division is joined to the trapezium and trapezoides. The concavity receives more than half of the round head of the os magnum. The internal side of this hollow is formed into a semilunar plain, to be articulated with the following bone. The external, posterior, and anterior edges are rough, for fixing the ligaments that connect it to the sur- rounding bones. Os lunare has a smooth convex upper surface, by which it is articulated with the radius. The external side, which gives the name to the bone, is in the form of a crescent, Carpus, 127 and is joined with the scaphoid: the lower surface is hol- low, for receiving part of the head of the os magnum. On the inside of this cavity is another smooth, but narrow, oblong sinuosity, for receiving the upper end of the os unciforme: and on the inside of this a small convexity is found, for its connexion with the os cuneiforme. Between the great convexity above, and the first deep inferior ca- vity, there is a rough fossa, in which the circular ligament of the joint of the wrist is fixed. Os cuneiforme is broader above, and towards the back of the hand, than it is below and forwards; which gives it the resemblance of a wedge. The superior slightly convex surface is included in the joint of the wrist, being opposed to the lower end of the ulna. Below this the cuneiforme bone has a rough fossa, wherein the ligament of the articu- lation of the wrist is fixed. On the external side of this bone, where it is contiguous to the os lunare, it is smooth and slightly concave. Its lower surface, where it is conti- guous to the os unciforme, is oblong, somewhat spiral, and concave. Near the middle of its anterior surface a circular plain appears, where the os pisiforme is sustained. Os pisiforme is almost spherical, except one circular plain, or slightly hollow surface, which is covered with cartilage for its motion on the cuneiforme bone, from which its whole rough body is prominent forwards into the palm; having the tendon of the flexor carpi ulnaris, and a liga- ment from the styloid process of the ulna, fixed to its up- per part; the transverse ligament of the wrist is connected to its external side; ligaments extended to the unciforme bone, and to the os metacarpi of the little finger, are at- tached to its lower part; the Abductor minimi digiti has its origin from its fore part; and, at the external side of it, a small depression is formed, for the passage of the ulnar nerve. 128 Carpus. Second Row. Trapezium has four unequal sides and angles in its back part, from which it has got its name. Above, its surface is smooth, slightly hollowed, and semicircular, for its con- junction with the os scaphoides. Its internal side is an oblong concave square, for receiving the following bone. The inferior surface is formed into a pulley; which faces obliquely outwards and downwards when the palm pre- sents forward. On this pulley the first bone of the thumb is moved. At the internal side of the external protuberance, a small oblong smooth surface is formed by the os metacarpi indicis. The fore part of the trapezium is prominent in the palm, and near to the internal side has a sinuosity in it, where the tendon of the flexor carpi radialis is lodged; on the ligamentous sheath of which the tendon of the flexor tertii internodii pollicis plays: and near this the bone is scabrous, where the transverse ligament of the wrist is 'connected, the abductor and flexor primi internodii pol- licis have their origin, and ligaments go out to the first bone of the thumb. Os trapezoides, so called from the irregular quadrangu- lar figure of its back part, is the smallest bone of the wrist except the pisiforme. The figure of it is an irregular cube. It has a small hollow surface above, by which it joins the scaphoides; a long convex one externally, where it is con- tiguous to the trapezium; a small internal one, for its con- junction with the os magnum; and an inferior convex sur- face, the edges of which are, however, so raised before and behind, that a sort of pulley is formed, where it sustains the os metacarpi indicis. Os magnum, so called because it is the largest bone of Carpus. 129 the carpus, is oblong, having four quadrangular sides, with a round upper end, and a triangular plain one below. The round head is divided by a small rising, opposite to the connexion of the os scaphoides and lunare, which together form the cavity for receiving it. On the outside a short plain surface joins the os magnum to the trapezoides. On the inside is a long narrow concave surface, where it is contiguous to the os unciforme. The lower end, which sustains the metacarpal bone of the middle finger, is trian- gular, slightly hollowed, and farther advanced on the ex- ternal side than on the internal, having a considerable ob- long depression made on the advanced outside by the me- tacarpal bone of the fore finger; and generally there is a small mark of the os metacarpi digiti annularis on its internal side. Os unciforme has got its name from a thin broad process that stands out from it forwards into the palm, and is hol- low for affording passage to the tendons of the flexors of the fingers. To this process also the transverse ligament is fixed that binds down and defends these tendons; and the flexor and abductor muscles of the little finger have part of their origin from it. The upper plain surface is small, convex, and joined with the os lunare: the external side is long, and slightly convex, adapted to the contigu- ous os magnum. The external surface is oblique, and irregularly convex, to be articulated with the cuneiforme bone. The lower end is divided into two concave surfaces; the internal is joined with the metacarpal bone of the lit- tle finger; and the external one is fitted to the metacarpal bone of the ring finger. The nature of the carpus will be best understood by stu- dying the bones placed together, in their natural order, in the two rows. Vol. I. K 130 Carpus. When thus placed they compose a structure of an ob- long form, whose greatest length extends across the wrist and forms a concavity in front, while it is convex poste- riorly. Two bones of the first row, viz. the scaphoides and lunare, form an oblong convex surface, which has a trans- verse position with respect to the arm, and applies to the concave surface at the end of the radius. These surfaces are particularly calculated for flexion and extension, and also for a considerable motion to each side; and by a suc- cession of these flexures, in different directions, the hand performs a circular motion, although it cannot perform at this joint, a rotation, or revolution on the axis of the car- pus. The under surface of these bones has a deep concavity which is composed by the scaphoides, lunare and cunei- forme, and receives a prominence of the second row. It also presents a convex surface formed by the scaphoides, which is received by the second row. The upper surface of the second row, which is concern- ed in this articulation, is very irregular; it has a head formed by the magnum and unciforme, which penetrates deeply into the cavity of the first row. On the outside of this head the trapezium and trapezoides form a surface which receives the projecting part of the scaphoides; so that the first row receives, and is received by the second, and the two surfaces are well calculated for moving to a certain extent, in the way of flexion and extension, upon each other* The lower surface of the second row, which is connected to the metacarpal bones, appears like the, side of an arch, which is partly induced by the wedge-like form of the two bones in the center, viz. the trapezoides and the mag- Metacarpus. 131 num. When the hand hangs by the side, and the palm is forward, all of this surface presents downwards, except that portion of it which is formed by the trapezium. This bone is placed obliquely between the two rows, and its surface for supporting the thumb presents obliquely down- wards and outwards. The trapezoides supports the fore finger, the magnum the middle finger, and the unciforme both of the remain- ing fingers. The scaphoides and the trapezium are very prominent at the external side of the anterior concave surface of the carpus; and the unciforme process, and the os pisiforme on the internal. The Metacarpus Consists of four bones which sustain the fingers. Each bone is long and round, with its ends larger than its body. The upper end, which some call the base, is flat and ob- long, inclining somewhat to the wedge-like form, without any considerable head or cavity; but it is however some- what hollowed, for the articulation with the carpus: It is made flat and smooth on the sides where these bones are contiguous to each other. Their bodies are flatted on their back part, particularly below the middle, by the tendons of the extensors of the fingers. The anterior surface of these bodies is a little convex, especially in their middle; along which a sharp ridge stands out, separating the musculi interossei placed on each side of these bones, which are there made flat and plain by these muscles. Their lower ends are raised into large oblong smooth heads, whose greatest extent is forwards from the axis of the bone. At the fore part of each side of the root of each of these heads, one or two tubercles stand out, for fixing 132 Metacarpus. the ligaments that go from one metacarpal bone to another, to preserve them from being drawn asunder. Round the heads a rough ring may be remarked, for the capsular ligaments of the first joints of the fingers to be fixed to; and both sides of these heads are flat, by pressing on each other. The substance of the metacarpal bones is the same with that of all long bones. The metacarpal bones are joined above to the bones of the carpus, and to each other, by surfaces almost flat. These connexions do not admit of much motion. The articulation of the round heads, at their lower ends, with the cavities of the first bones of the fingers, will soon be described. The concavity on the fore part of these metacarpal bones, and the position of their bases on the arched carpus, cause them to form a hollow in the palm of the hand, which is often useful to us. The spaces between them lodge muscles, and their small motion makes them fit sup- porters for the fingers to play on. Though the ossa metacarpi so far agree, yet they ma\ be distinguished from each other by the following marks. The metacarpal bone of the forefinger is generally the longest. Its base, which is articulated with the os trape- zoides, is hollow in the middle. The small ridge on the external side of this oblong cavity is smaller than the one opposite to it, and is made flat on the side by the trape- zium. The internal ridge is also smooth, and flat on its outside, for its conjunction with the os magnum; imme- diately below which a semicircular smooth flat surface shows the articulation of this to the second metacarpal bone. The back part of this base is flatted where the long head of the extensor carpi radialis is inserted, and its fore Metacarpus. 133 part is prominent where the tendon of the flexor carpi radia'.is is fixed. The tubercle at the internal root of its head is larger than the external. Its base is so firmly fixed to the bone it is connected with, that it has no motion. The metacarpal bone of the middle finger is generally the second in length: but often it is as long as the former; sometimes it is longer; and it frequently appears only to equal the first by the os magnum being farther projected downwards than any other bone of the wrist. Its base is a broad superficial cavity, slanting inwards; the external posterior angle of which is so prominent, as to have the appearance of a process. The external side of this base is made plain in the same way as the external side of the former bone, while its internal side has two hollow circu- lar surfaces, for joining the third metacarpal bone; and between these surfaces there is a rough fossa, for the ad- hesion of a ligament, and lodging mucilaginous glands. The extensor carpi radialis brevior is inserted into the back part of this base. The two sides of this bone are almost equally flatted; but the ridge on the fore part of the body inclines inwards. The tubercles at the fore part of the root of the head are equal. The motion of this bone is very little more than that of the former; and therefore these two firmly resist bodies pressed against them by the thumb or fingers, or both. The metacarpal bone of the ring finger is shorter than the second metacarpal bone. Its base is semicircular and convex, for its conjunction with the os unciforme; On its external side are two smooth convexities, and a middle fossa, adapted to the second metacarpal bone. The inter- nal side has a triangular smooth concave surface to join it with the fourth one. The anterior ridge of its body is situ- ated more to the inside than to the outside. The tubercles 134 Thumb. near the head are equal. The motion of this third meta- carpal bone is greater than the motion of the second. The metacarpal bone of the little finger is the smallest and sharpest. Its base is irregularly convex, and rises slanting inwards. Its external side is exactly adapted to the third metacarpal bone. The internal has no smooth surface, because it is not contiguous to any other bone; but it is prominent where the extensor carpi ulnaris is in- serted. As this metacarpal bone is furnished with a proper moving muscle, has the plainest articulation, is most loose- ly connected and least confined, it not only enjoys a much larger motion than any of the rest, but draws the third bone with it, when the palm of the hand is to be made hollow by its advancement forwards, and by the promi- nence of the thumb opposite to it. Thumb and Fingers. The thumb and fourfingers are each composed of three bones. The thumb is situated obliquely in respect of the fin- gers; neither opposite directly to them, nor in the same plane with them All its bones are much thicker and stronger in proportion to their length, than the bones of the- fingers are: which is extremely necessary, as the thumb conteracts all the fingers. The first bone of the thumb has its base adapted to the peculiar articulating surface of the trapezium: for, in viewing it from one side to the other, it appears convex in the middle; but when viewed from behind forwards, it is concave there. The edge at the fore part of this base is extended farther than any other part; and round the back part of the base a rough fossa may be seen, for the Thumb. 135 connexion of the ligaments of this joint. The body and head of this bone are of the same shape as the ossa meta* carpi; only that the body is shorter, the head flatter, and the tubercles at the fore part of its root larger. The articulation of the upper end of this bone is re- markable: for, though it has protuberances and depres- sions adapted to the double pulley of the trapezium, yet it enjoys a circular motion, as the joints do where a round head of the one plays in the orbicular socket of another: it is however more confined, and less expeditious, but stronger and more secure than such joints generally are. The second bone of the thumb has a large base formed into an oblong cavity, whose greatest length is from one side to the other. Round it several tubercles may be re- marked, for the insertion of ligaments. Its body is convex, or half-round behind; but flat before, for lodging the tendon of the long flexor of the thumb, which is tied down by ligamentous sheaths, that are fixed on each side to the angle at the edge of this flat surface. The lower end of this second bone has two lateral round protuberances, and a middle cavity, whose greatest extent of smooth surface is forwards and backwards. The articulation of the upper end of this second bone would,seem calculated for motion in all directions; yet, on account of the strength of its lateral ligaments, the ob- long figure of the joint itself, and mobility of the first joint, it only allows flexion and extension; and these are generally much confined. The third bone of the thumb is the smallest, with a large base, whose greate'st extent is from one side to the other. This base is formed into two cavities and a middle protu- berance, to be adapted to the pulley of the former bone. This bone becomes gradually smaller, till near the lower 136 lingers. end, where it is a little enlarged, and has an oval scabrous edge. Its body is rounded behind, but is flatter than in the former bone, for sustaining the nail. It is flat and rough before, by the insertion of the flexor tertii internodii. The motion of this third bone is confined to flexion and extension. The regular arrangement of the bones of the fingers in three rows, has obtained for them the name of the three phalanges. All of them have half round convex surfaces, covered with an aponeurosis, formed by the tendons of the extensors, lumbricales, and interossei, and placed directly backwards, for their greater strength; and their flat con- cave part is forwards, for taking hold more surely, and for lodging the tendons of the flexor muscles. The ligaments for keeping down these tendons are fixed to the angles that are between the convex and concave sides. The bones of the first phalanx of the fingers answer to the description of the second bone of the thumb; only that the cavity in their base is not so oblong; nor is their mo- tion on the metacarpal bones so much confined: for they can move laterally or circularly, the fore finger in particu- lar, but have no rotation, or a very small degree of it, round their axis. The second bone of the fingers has its base formed into two lateral cavities, and a middle protuberance; while the lower end has two lateral protuberances and a middle ca- vity; therefore it is joined at both ends in the same man- ner; which none of the bones of the thumb are. The third bone differs nothing from the description of the third bone of the thumb, except in the general distin- guishing marks; and therefore the second and third pha-" lanxof the fingers enjoy only flexion and extension. The Fingers. 137 All the difference of the phalanges of the several fingers consists in their magnitude. The bones of the middle fin- ger being the longest and largest; those of the forefinger come next to these in thickness, but not in length, for those of the ring finger are a little longer. The little finger has the smallest bones. Which disposition is the best contri- vance for holding the largest bodies; because the longest fingers are applied to the middle largest periphery of such substances as are of a spherical figure. Vol. I. S 138 SECTION IV. The Inferior Extremities. THE inferior extremities consist of the Thigh, Leg ;md Foot. The Thigh Consists of one bone only; the os femoris, which is very strong, and larger than any other in the skeleton. It is nearly cylindrical in the middle, and slighdy curved. The upper extremity is a spherical head, connected to the body of the bone by a neck. The lower extremity is much larger than the body, and is formed into two condyles. The upper end of this bone is not continued in a straight line with the body of it, but the axis of it inclines obliquely inwards and upwards, whereby the distance between these two bones at their upper part is considerably increased. The head is the greater portion of a sphere. Towards its lower internal part a round rough spongy pit is observable, where the strong ligament, commonly, but inaccurately, called the round one, is fixed, to be extended from thence to the lower internal part of the receiving cavity, where it is considerably broader than near to the head of the thigh bone. The neck of the os femoris, has a great many large holes, into which the fibres of the strong ligament, conti- nued from the capsular, enter, and are thereby firmly uni- ted to it; and round the root of the neck, where it rises from the bone, a rough ridge is found, where the capsular ligament of the articulation itself is connected. Below this root, a large unequal protuberance, called trochanter major, Os Femoris. 139 stands out; the external convex part of which is distin- guished into three different surfaces: whereof the one on the upper and front part is scabrous and rough, for the in- sertion of the glutaeus minimus; the superior one is smooth, and has the glutaeus medius inserted into it; and the one behind is made flat and smooth, by the tendon of the glu- taeus maximus passing over it. The upper edge of this pro- cess is sharp and pointed at its back part, where the glu- taeus medius is fixed; but forwards it is more obtuse, and under it is a depression, into which some of the muscles which rotate the thigh outwards, are fixed. From the pos- terior prominent part of this great trochanter, a rough ridge runs backwards and downwards, into which the quadratus is inserted. In the deep hollow, at the internal upper side of this ridge, the obturator externus is implant- ed. More internally, a conical process, called trochanter minor, rises, for the insertion of the musculus psoas and iliacus internus; and the pectineus is implanted into a rough hollow below its internal root. The muscles insert- ed into these two processes being the principal instruments of the rotatory motion of the thigh, have occasioned the name of trochanters to be given to these processes. The body of the os femoris is convex on the fore part, and concave behind; which enables us to sit without lean- ing too much on the posterior muscles. On the posterior concave surface is a broad rough ridge called linea aspera, which commences near the great tro- chanter, and continues downwards, more than two-thirds of the length of the bone, when it divides into two ridges which descend towards each condyle. The internal of these ridges is the most smooth; and the space between them is nearly flat. Near the end of each of these ridges, a small 140 Os Femoris. smooth protuberance may often be remarked, where the two heads of the external gastrocnemius muscle take their rise; and from the fore part of the internal tubercle a strong ligament is extended to the inside of the tibia. The lower end of the os femoris is larger than any other part of it, and is formed into two great protuberances, one on each side, which are called its condyles: between them a considerable cavity is found, especially at the back part, in which the crural vessels and nerves lie. The internal condyle is longer than the external, which must happen from the oblique position of this bone, to give less obliquity to the leg. These processes are of an oblong form, and are placed obliquely with respect to each other; being in con- tact before, and separated to a considerable distance be- hind. They form in front a smooth pulley-like surface, the external side of which is highest, on which the patella moves. Below, they are flat; and posteriorly, they are regularly convex. Between these convex portions is a rough cavity, from which the crucial ligament arises, to be attached to the tibia. Round the lower end of the thigh bone, large holes are found, into which the ligaments for the security of the joint are fixed, and bloodvessels pass to the internal sub- stance of the bone. The thigh bone being articulated above with the aceta- bulum of the ossa innominata, which affords its round head a secure and extensive play, can be moved to every side; but it is restrained in its motion outwards, by the high brims of the cavity, and by the round ligament; for other- wise the head of the bone would have been frequently Os Femoris. 141 thrust out at the breach of the brims on the inside, which allows the thigh to move considerably inwards. The body of this bone enjoys little or no rotatory motion, though the head most commonly moves round its own axis; because the oblique direction of the neck and head, from the bone, is such, that the rotatory motion of the head can only bring the body of the bone forwards and backwards. Nor is the head, as in the arm, ever capable of being brought to a straight direction with its body; so far, however, as the head can move within the cavity backwards and forwards, the rest of the bone may have a partial rotation. From the oblique position of these bones it results, that there is a considerable distance between them above, while the knees are almost contiguous. Sufficient space is there- by left for the external parts of generation, for the two great outlets of urine and faeces, and for the large thick muscles that move the thigh inwards. At the same time this situation of the thigh bones renders our progression quicker, surer, straighter, and in less room: for, had the knees been at a greater distance from each other, we must have been obliged to describe some part of a circle with the trunk of our body in making a long step; and when one leg was raised from the ground, our center of gravity- would have been too far from the base of the other, and we should consequently have been in danger of falling; so that our steps would neither have been straight nor firm, nor would it have been possible to walk in a narrow path, had our thigh bones been otherwise placed. In conse- quence, however, of the weight of the body bearing so obliquely on the joint of the knee by this situation of the thighbones, weak rickety children become knock-kneed. 142 The Leg.—Tibia. The Leg Is composed of two bones, the tibia and fibula. The patella, being evidently appropriated to the knee joint, may be regarded as common, both to the thigh and leg. , 77ieles of tlie Superior Extremities. THESE may be divided into the muscles that are situ- ated on the scapula, on the os humeri, on the cubit or fore arm, and on the hand. Muscles situated on the Scapula. These are called muscles of the os humeri; and are three behind, one along its inferior costa, two before, and one beneath it. Behind are, 1. Supraspinatus. Arises, fleshy, from all that part of the base of the sca- pula that is above its spine; also from the spine and supe- rior costa; passes under the acromion, and adheres to the capsular ligament of the os humeri. Inserted, tendinous, into that part of the large protube- rance on the head of the os humeri that is next the groove) for lodging the tendon of the long head of the biceps. Use. To raise the arm upwards; and, at the same time, to pull the capsular ligament from between the bones, that it may not be pinched. 2. Infraspinatus, Arises, fleshy, from all that part of the base of the sca- pula that is between its spine and inferior angle; from the spine as far as the cervix of the scapula. The fibres ascend and descend obliquely towards a tendon in the middle of the muscle, which runs forwards, and adheres to the cap- sular ligament. Inserted, by a thick and short tendon, into the upper and middle part of the large protuberance on the head of the os humeri. Vol. I. 2 F 226 Muscles situated on the Scapula. Use. To roll the humerus outwards; to assist in raising, and in supporting it when raised; and to pull the ligament from between the bones. N. B, These two muscles are covered with a tendinous membrane, from which a number of their fleshy fibres arise. It serves besides to strengthen their actions, and keeps them from swelling too much outwardly when in action. 3. Teres Minor, Arises, fleshy, from all the round edge of the inferior costa of the scapula, and runs forwards along the inferior edge of the infraspinatus muscle, and adheres to the liga- ment. Inserted, tendinous, into the back part of the large pro- tuberance on the head of the os humeri, a little behind and below the termination of the last named muscle. Use, To roll the humerus outwards; to draw the hu- merus backwards; and to prevent the ligament from being pinched between the bones. Along the inferior costa of the scapula is, Teres Major. Arises, fleshy, from the inferior angle of the scapula, and from all that portion of its inferior costa that is rough and jhicker than the rest; its fleshy fibres are continued over part of the infraspinatus muscle, to which they firmly adhere. Inserted, by a broad, short, and thin tendon, into the ridge at the inner side of the groove for lodging the ten- don of the long head of the biceps, along with the latissi- mus dorsi. Deltoides.—Coraco-Brachialis. 227 Use. To roll the humerus inwards, and to draw it back- wards and downwards. The two before the scapula are, 1. Deltoides, Arises, fleshy, from all the posterior part of the clavicle that the pectoralis major does not possess; tendinous and fleshy, from the acromion, and lower margin of almost the whole spine of the scapula opposite to the insertion of the cucullaris muscle: from these origins it runs in three dif- ferent directions, i. e. from the clavicle outwards and downwards; from the spine of the scapula outwards, for- wards, and downwards; and from the acromion straight downwards; and is composed of a number of fasciculi, which form a strong fleshy muscle that covers the anterior part of the joint of the os humeri. Inserted, tendinous, into a rough protuberance in the outer side of the os humeri, near its middle, where the fibres of this muscle intermix with some part of the bra- chialis externus. Use. To pull the arm directly outwards and upwards, and a little forwards or backwards, according to the dif- ferent directions of its fibres. 2. Coraco-Brachialis, Arises, tendinous and fleshy, from the fore part of the coracoid process of the scapula; adhering, in its descent, to the short head of the biceps. Inserted, tendinous and fleshy, about the middle of the internal part of the os humeri, near the origin of the third head of the triceps, called brachialis externus, where it sends down a thin tendinous expansion to the internal condyle of the os humeri. 228 Subscapularis.—Biceps. Use. To raise the arm upwards and forwards. N. B. There passes a nerve through this muscle, called musculo cutaneus. The one beneath the scapula is, Subscapularis. Arises, fleshy, from all the base of the scapula internally, and from its superior and inferior costre, being composed of a number of tendinous and fleshy fasciculi, which make prints on the bone; they all join together, fill up the hol- low of the scapula, and pass over the joint, adhering to the capsular ligament. Inserted, tendinous, into the upper part of the internal protuberance at the head of the os humeri. Use. To roll the humerus inwards, and to draw it to the side of the body; and to prevent the capsular ligament from being pinched. Muscles situated on the Os Humeri. These are called Muscles of the Cubit or Fore Arm. They consist of two before, and two behind. Before are, 1. Biceps Flexor Cubiti. Arises, by two heads. The first and outermost, called longus, begins tendinous from the upper edge of the gle- noid cavity of the scapula; passes over the head of the os humeri within the joint; and, in its descent without the joint, is inclosed in a groove near the head of the os hu- meri, by a membranous ligament that proceeds from the Brachialis Internus. 229 capsular ligament and adjacent tendons. The second, or innermost head, called brevis, arises, tendinous and fleshy, from the coracoid process of the scapula, in common with the coraco-brachialis muscle. A little below the middle of the fore part of the os humeri, these heads unite. Inserted, by a strong roundish tendon, into the tubercle on the upper end of the radius internally. Use. To turn the hand supine, and to bend the fore arm. N. B. At the bending of the elbow, where it begins to grow tendinous, it sends off an aponeurosis, which covers all the muscles on the inside of the fore arm, and joins with another tendinous membrane, which is sent off from the triceps extensor cubiti, and covers all the muscles on the outside of the fore arm, and a number of the fibres, from opposite sides, decussate each other. It serves to strengthen the muscles, by keeping them from swelling too much outwardly, when in action; and a number of their fleshy fibres take their origin from it. 2. Brachialis Internus, Arises, fleshy, from the middle of theos humeri, at each side of the insertion of the deltoid muscle, covering all the inferior and fore part of this bone, runs over the joint, and adheres firmly to the ligament. Inserted, by a strong short tendon, into the coronoid process of the ulna. Use. To bend the fore arm, and to prevent the capsular ligament of the joint from being pinched. 230 Triceps Extensor.—Anconeus. Behind, are 1. Triceps Extensor Cubiti. Arises, by three heads; the first called longus, somewhat broad and tendinous, from the inferior costa of the scapu- la, near its cervix. The second head, called brevis, arises by an acute, tendinous, and fleshy beginning, from the back part of the os humeri, a little below its head, out- wardly. The third, called brachialis externus, arises by an acute beginning, from the back part of the os humeri. These three heads unite lower than the insertion of the teres major, and cover the whole posterior part of the hu- merus, from which they receive addition in their descent. Inserted into the upper and external part of the process of the ulna, called olecranon, and partly into the condyles of the os humeri, adhering firmly to the ligament. Use. To extend the fore arm. 2. Anconeus, Arises, tendinous, from the posterior part of the exter- nal condyle of the os humeri; it soon grows fleshy, and is continued from the third head of the triceps. Inserted, fleshy, and thin, into a ridge on the outer and posterior edge of the ulna, being continued some way be- low the olecranon, and covered with a tendinous mem- brane. Use. To assist in extending the fore arm. Palmaris Longus. 231 Muscles situated on the Fore Arm. These may be divided into three classes, viz. 1. The muscles which bend and extend the wrist, and of course the whole hand. 2. Those which bend and extend the fingers exclusively. 3. Those which act on the radius so as to roll it backwards and forwards on the ulna; which are called supinators and pronators. The flexors both of the wrist and fingers, and the prona- tors, lie on the front of the fore arm. The extensors and the supinators on the back. The flexors generally originate from the internal condyle of the os humeri, and the parts adjacent to it: the exten- sors from the external condyle of the same bone, and the parts which are near it. In the following description they are arranged in the order in which they occur in the dissection of the arm; begin- ning with those which originate from the internal con- dyle, without regard to their particular functions. Muscles on the Anterior part of the Fore Arm. 1. Palmaris Longus, Arises, tendinous, from the internal condyle of the os humeri, soon grows fleshy, and after a short progress, sends off a long slender tendon. Inserted into the ligamentum carpi annulare, and into a tendinous membrane that is expanded on the palm of the hand, named aponeurosis palmaris; which, above, begins at the transverse or annular ligament of the wrist, and, be- low, is fixed to the roots of the fingers. 232 Muscles on the Anterior Use. To bend the hand, and to stretch the membrane that is expanded on the palm. N. B. This muscle is sometimes wanting; but the apo- neurosis-palmaris is always to be found. 2. Pronator Radii Teres, Arises, fleshy, from the internal condyle of the os hu- meri, and tendinous from the coronoid process of the ulna. Inserted, thin, tendinous, and fleshy, into the middle of the posterior part of the radius. Use, To roll the radius, together with the hand, inwards. 3. Flexor Carpi Radialis, Arises, tendinous and fleshy, from the internal condyle of the os humeri, and from the anterior part of the upper end of the ulna, where it firmly adheres to the pronator radii teres. Inserted, by a flat tendon, into the fore and upper part of the metacarpal bone that sustains the fore finger, after running through a fossa in the os trapezium. Use. To bend the hand, and to assist in its pronation. 4. Flexor Carpi Ulnaris, Arises, tendinous, from the internal condyle of the os humeri. It has likewise a small fleshy beginning from the outer side of the olecranon; between which and the con- dyle the ulnar nerve passes to the fore arm; and a number of its fleshy fibres arise from the tendinous membrane which covers the fore arm. Inserted, by a short strong tendon, into the os pisiforme. At a little distance from its insertion, a small ligament is sent off to the metacarpal bone that sustains the little finger. Use. To assist the former in bending the arm. i part of the Fore Arm. 23:3 5. Flexor Sublimis Perforatus, Arises, tendinous and fleshy, from the internal condyle of the os humeri; tendinous from the coronoid process of the ulna, near the edge of the cavity that receives the head of the radius; fleshy from the tubercle of the radius; and membranous and fleshy from the middle of the fore part of the radius, where the flexor pollicis longus arises. Its fleshy belly sends off four round tendons before it passes under the ligament of the wrist. Inserted, into the anterior and upper part of the second bone of each finger, being, near the extremity of the first bone, divided for the passage of the perforans. Use. To bend the second joint or phalanx of the fingers. 6. Flexor Profundus Perforans, Arises, fleshy, from the external side, and upper part of the ulna, for some way downwards, and from a large share of the interosseous ligament. It splits into four tendons, a little before it passes under the ligamentum carpi annu- lare; and these pass through the slits in the tendons of the flexor sublimis. Inserted into the fore and upper part of the third or last v bone of all the four fingers. Use. To bend the'last joint of the fingers. 7. Flexor Longus Pollicis Manus, Arises, by an acute fleshy beginning, from the upper part of the radius, immediately below its tubercle, and is continued down for some space on the fore part of this bone. It has likewise generally another origin from the internal condyle of the os humeri, which forms a distinct fleshy slip that terminates near the upper part of the origin from the radius. Vol. I. 2 G 234 Muscles on the back part Inserted into the last joint of the thumb, after having passed its tendon under the ligament of the wrist. Use. To bend the last joint of the thumb.* 8. Pronator Radii §>uadratus, Arises, broad, tendinous, and fleshy, from the lower and inner part of the ulna; the fibres run transversely, to be Inserted into the lower and anterior part of the radius, opposite to its origin. Use. To turn the radius, together with the hand, in^ wards. Muscles of the External Side and Back of the Arm. 1. Supinator Radii Longus, Arises, by an acute and fleshy origin, from the external ridge of the os humeri, above the external condyle, nearly as far up as the middle of that bone. Inserted into the outer side of the inferior extremity of the radius. Use. To roll the radius outwards, and consequently the palm of the hand upwards. 2. Extensor Carpi Radialis Longior, Arises, broad, thin, and fleshy, immediately below the supinator radii longus, from the lower part of the external ridge of the os humeri, above its external condyle. * The thumb has but one flexor muscle on the front of the arm, although it has three extensors on the back part. of the Fore Arm. 235 Inserted, by a round tendon, into the posterior and upper part of the metacarpal bone that sustains the fore finger. Use. To extend and bring the hand backwards. 3. Extensor Carpi Radialis Brevior, Arises, tendinous, from the external condyle of the os humeri, and from the ligament that connects the radius to it, and runs along the outside of the radius. Inserted, by a round tendon, into the upper and back part of the metacarpal bone that sustains the middle finger. Use. To assist the last mentioned muscle. 4. Extensor Carpi Ulnaris, Arises, tendinous, from the external condyle of the os humeri; and, in its progress, fleshy from the middle of the ulna, where it passes over the ulna. Its round tendon is inclosed by a membranous sheath, in a groove which is situated at the extremity of the ulna. Inserted, by its round tendon, into the posterior and upper part of the metacarpal bone that sustains the little finger. Use. To assist the former in extending the hand. 5. Extensor Digitorum Communis, Arises, by an acute, tendinous, and fleshy beginning, from the external condyle of the os humeri, where it ad- heres to the supinator radii brevis. Before it passes under the ligamentum carpi annulare externum, it splits into four tendons; some of which may be divided into several smaller; and about the fore part of the metacarpal bones they remit tendinous filaments to each other. 236 Muscles on the back part Inserted into the posterior part of all the bones of the four fingers, by a tendinous expansion. Use. To extend all the joints of the fingers. 6. Supinator Radii Brevis, Arises, tendinous, from the external condyle of the os humeri; tendinous and fleshy, from the external and up- per part of the ulna, and adheres firmly to the ligament that joins these two bones. Inserted into the head, neck, and tubercle of the radius, near the insertion of the biceps, and ridge running from that downwards and outwards. Use. To roll the radius outwards, and so bring the hand supine. 7. Indicator, Arises, by an acute fleshy beginning, from the middle of the posterior part of the ulna; its tendon passes under the same ligament with the extensor digitorum communis, with part of which it is Inserted into the posterior part of the fore finger. Use. To extend the fore finger separately. 8. Extensor Ossis Metacarpi Pollicis Manus, Arises, fleshy, from the middle and posterior part of the ulna, immediately below the insertion of the anconeus muscle, from the posterior part of the middle of the radi- us, and from the interosseous ligament. Inserted, generally by two tendons, into the os trapezi- um, and upper back part of the metacarpal bone of the thumb, and often joins with the abductor pollicis. Use, To extend the metacarpal bone of the thumb out- wardly. of the Fore Arm. 237 9. Extensor Primi Internodii, Arises, fleshy, from the posterior part of the ulna near the former muscle, and from the interosseous ligament. Inserted, tendinous, into the posterior part of the first bone of the thumb; and part of it may be traced as far as the second bone. Use. To extend the first bone of the thumb obliquely outwards. 10. Extensor Secundi Internodii, Arises, by an acute, tendinous, and fleshy beginning, from the middle back part of the ulna, and from the inter- osseous ligament; its tendon runs through a small groove at the inner and back part of the lower end of the radius. Inserted into the last bone of the thumb. Use. To extend the last joint of the thumb obliquely backwards. Muscles on the Palm of the Hand. To obtain a full view of the muscles situated on the palm of the hand, it will be necessary to remove the annular or transverse ligament, which is stretched across from the projecting points of the pisiform and unciform bones on the inside of the wrist to the scaphoid and trapezium on the outside; for the purpose of retaining the tendons of the flexor muscles in their proper situation. And also, to remove from the palm of the hand the aponeurosis pal- maris, which has been described with the palmaris lon- gus muscle. 1. Palmaris Brevis, Arises from the ligamentum carpi annulare, and tendi- nous membrane that is expanded on the palm of the hand. 238 • Muscles on the Palm Inserted, by small bundles of fleshy fibres, into the skin and fat that covers the abductor minimi digiti, and into the os pisiforme. Use. To assist in contracting the palm of the hand. 2. Abductor Pollicis Manus, Arises, by a broad tendinous and fleshy beginning, from the ligamentum carpi annulare, and from the os trape- zium. Inserted, tendinous, into the outer side of the root of the first bone of the thumb. Use. To draw the thumb from the fingers. 3. Flexor Ossis Metacarpi Pollicis, or Opponens Pollicis, Arises, fleshy, from the os trapezium and ligamentum carpi annulare, lying under the abductor pollicis. Inserted, tendinous and fleshy, into the under and ante- rior part of the metacarpal bone of the thumb. Use. To bring the thumb inwards, opposite to the other fingers. 4. Flexor Brevis Pollicis Manus, Arises from the os trapezoides, magnum, and unciforme of the carpus, and is divided into two portions by the ten- don of the flexor pollicis longus. Inserted into the ossa sesamoidea and first bone of the thumb. Use. To bend the first joint of the thumb. 5. Adductor Pollicis Manus, Arises, fleshy, from almost the whole length of the me- tacarpal bone that sustains the middle finger; from thence its fibres are collected together. of the Hand. 239 Inserted, tendinous, into the inner part of the root of the first bone. Use. To pull the thumb towards the fingers. There are four small flexors, called, from their form, 6. Lumbricales, Which arise, thin and fleshy, from the outside of the tendons of the flexor profundus, a little above the lower edge of the ligamentum carpi annulare. Inserted, by long slender tendons, into the outer sides of the broad tendons of the interossei muscles, about the middle of the first joint. Use. To increase the flexion of the fingers while the long flexors are in full action. 7. Adductor Metacarpi Minimi Digiti Manus, Arises, fleshy, from the thin edge of the os unciforme, and from that part of the ligament of the wrist next to it. Inserted, tendinous, into the inner side and anterior part of the metacarpal bone of this finger. Use. To bend and bring the metacarpal bone of this finger towards the wrist. 8. Flexor Parvus Minimi Digiti, Arises, fleshy, from the outer side of the os unciforme, and from the ligament of the wrist which joins with that bone. Inserted, by a roundish tendon, into the inner and ante- rior part of the upper end of the first bone of this finger. Use. To bend the little finger, and assist the adductor. 9. Abductor Minimi Digiti Manus, Arises, fleshy, from the os pisiforme, and from that part of the ligamentum carpi annulare next it. 240 Muscles on the Palm Inserted, tendinous, into the inner side of the tipper end of the first bone of the little finger. Use. To draw this finger from the rest. The spaces between the metacarpal bones are occupied by muscles called, from their situation, interosseous. The four following are to be seen on the palm of the hand. Anterior Interosseous Muscles. 1. Prior Indicis, Arises, tendinous and fleshy, from the upper and outer .part of the metacarpal bone that sustains the fore finger. Inserted into the outside of that part of the tendinous expansion from the extensor digitorum communis, which covers the posterior part of the fore finger. Use. To draw the fore finger outwards towards the thumb, and extend it obliquely. 2. Posterior Indicis, Arises, tendinous and fleshy, from the root and inner part of the metacarpal bone that sustains the fore finger. Inserted into the inner side of the tendinous expansion which is sent off from the extensor digitorum communis, along the posterior part of the fore finger. Use. To extend the fore finger obliquely, and to draw it inwards. 3. Prior Annularis, Arises, from the root of the outside of the metacarpal bone that sustains the ring finger. Inserted into the outside of the tendinous expansion of the extensor digitorum communis which covers the ring finger. of the Hand. 241 Use. To extend and pull the ring finger towards the thumb. 4. Interosseous Auricularis, Arises from the root and outer side of the metacarpal bone of the little finger; and is Inserted into the outside of the tendinous expansion of the extensor digitorum communis, which covers the pos- terior part of the little finger. Use. To extend and draw the little finger outwards. On the back of the hand three muscles of the same kind are to be seen, which also appear on the palm. Posterior Interosseous Muscles. 1. Prior Medii, Arises, by two origins, from the roots of the metacar- pal bones that sustain the fore and middle fingers exter- nally, and next each other: runs along the outside of the middle finger; and, being conspicuous on both sides of the hand, is Inserted into the outside of the tendinous expansion from the extensor digitorum communis, which covers the posterior part of the middle finger. Use. To extend, and to draw the middle finger out- wards. 2. Posterior Medii, Arises, by two origins, from the roots of the metacarpal bones, next each other, that sustain the middle and ring fingers. Vol. I. 2 II 242 Muscles of the back of the Hand. Inserted into the inside of the tendinous expansion from the extensor digitorum communis, which runs along the posterior part of the middle finger. Lse. To extend and draw the middle finger inwards. 3. Posterior Annularis, Arises, by two origins, from the roots of the metacarpal bones that sustain the ring and little fingers, next each other. Inserted into the inside of the tendon on the back of the ring finger. Use. To draw the ring finger inward. The following muscle also appears on the back of the hand. Abductor Indicis Manus, Arises, from the os trapezium, and from the superior part and inner side of the metacarpal bone of the thumb. Inserted, by a short tendon, into the outer and back part of the first bone of the fore finger. Use. To bring the fore finger towards the thumb. 243 Muscles of tlie Inferior Extremities. THESE may be divided into the muscles situated ojp. the outside of the pelvis, on the thigh, on the leg', and on the foot. Muscles on the outside of the pelvis, which are called muscles of the thigh. These are composed of one layer before and three layers aehind. The layer before consists, of five muscles: I. Psoas Magnus. *) „._ ° J. see page 212. 2. Iliacus Internus. J 3. Pectinalis, Arises, broad and fleshy, from the upper and anterior part of the os pubis or pectinis, immediately above the fo- ramen thyroideum. Inserted into the anterior and upper part of the linea aspera of the os femoris, a little below the trochanter mi- nor, by a flat and short tendon. Use. To bring the thigh upwards and inwards, and to give it a degree of rotation outwards. 4. Triceps Adductor Femoris, Under this appellation are comprehended three distinct muscles: a. Adductor Longus Femoris, Arises, by a strong roundish tendon, from the upper and anterior part of the os pubis, and ligament of its synchondrosis, on the inner side of the pectinalis. 244 Muscles of the Os Femoris. Inserted, tendinous, near the middle of the posterior part of the linea aspera, being continued for some way down. b. Adductor Brevis Femoris, Arises, tendinous, from the os pubis near its joining with the opposite os pubis, below and behind the former. Inserted, tendinous and fleshy, into the inner and upper part of the linea aspera, from a little below the trochanter minor, to the beginning of the insertion of the adductor longus. c. Adductor Magnus Femoris, Arises, a little lower down than the former, near the symphysis of the ossa pubis; tendinous and fleshy, from the tuberosity of the os ischium; the fibres run outwards and downwards. Inserted, into almost the whole length of the linea aspe- ra; into a ridge above the internal condyle of the os femoris; and, by a roundish long tendon, into the upper part of that condyle, a little above which the femoral ar- tery takes a spiral turn towards the ham, passing between this muscle and the bone. Use of these three muscles, or triceps. To bring the thigh inwards and upwards, according to the different di- rections of their fibres; and, in some degree, to roll the thigh outwards. 5. Obturator Externus, Arises, fleshy, from the lower fore part of the os pubis, and fore part of the inner crus of the ischium; surrounds the foramen thyroideum; a number of its fibres, arising from the membrane which fills up that foramen, are coi- Muscles of the Os Femoris. 245 lected like rays towards a center, and pass outwards around the root of the back part of the cervix of the os femoris. Inserted, by a strong tendon, into the cavity at the in- ner and back part of the root of the trochanter major, adhering in its course to the capsular ligament of the thigh bone. Use. To roll the thigh bone obliquely outwards, and to prevent the capsular ligament from being pinched. Behind are, First layer, Gluteus Maximus, Arises, fleshy, from the posterior part of the spine of the os ilium, a little higher up than the joining of the ilium with the os sacrum, from the whole external side of the os sacrum, below the posterior spinous process of the os ilium; from the posterior sacro ischiatic ligament, over which part of the inferior edge of this muscle hangs in a folded manner; from the os coccygis. All the fleshy fibres run obliquely forwards, and a little downwards, to form a thick broad muscle, which is divided into a number of strong fasciculi. The upper part of it covers almost the whole of the trochanter major, between which and the tendon of this muscle there is a large bursa mucosa, and where it is inseparably joined to the broad tendon of the tensor vagina femoris. Inserted, by a strong, thick, and broad tendon into the upper and outer part of the linea aspera, which is con- tinued from the trochanter major, for some way down- wards. Use. To extend the thigh, by pulling it directly back- wards, and a little outwards. 246 Muscles of the Os Femoris. Second layer, Gluteus Medius, Arises, fleshy, from the anterior superior spinous pro- cess of the os ilium, and from all the outer edge of the spine of the ilium, except its posterior part, where it arises from the dorsum of that bone. Inserted, by a broad tendon, into the outer and poste- rior part of the trochanter major. Use. To draw the thigh bone outwards, and a little backwards; to roll the thigh bone outwards, especially when it is bended. N. B. The anterior and upper part of this- muscle is covered by a tendinous membrane, from which a number of its fleshy fibres arise, and which joins with the broad tendons of the gluteus maximus, tensor vaginae femoris, and latissimus dorsi. Third layer consists of four muscles. 1. Gluteus Minimus, Arises, fleshy, from a ridge that is continued from the superior anterior spinous process of the os ilium, and from the middle of the dorsum of that bone, as far back as its great niche. • Inserted, by a strong tendon, into the fore and upper part of the trochanter major. Use. To assist the former in pulling the thigh outwards and backwards, and in rolling it. 2. Pyriformis, Arises, within the pelvis, by three tendinous and fleshy origins, from the second, third, and fourth pieces of the os sacrum; from thence growing gradually narrower, it Muscles of the Os Femoris* 247 passes out of the pelvis along with the posterior crural nerve, below the niche in the posterior part of the os ilium, where it receives a few fleshy fibres. Inserted, by a roundish tendon, into the upper part of the cavity at the inner side of the root of the trochanter major. Use. To move the thigh a little upwards, and roll it outwards. 3. Gemini, Arise, by two distinct origins; the superior from the spinous process, and the inferior from the tuberosity of the os ischium; also, from the posterior sacro ischiatic liga- ment. They are both united by a tendinous and fleshy membrane, and form a purse for the tendon of the obtura- tor internus muscle, which was formerly described. Inserted, tendinous and fleshy, into the cavity at the inner side of the root of the trochanter major, on each side of the tendon of the obturator internus, to which they firmly adhere. Use. To roll the thigh outwards, and to preserve the tendon of the obturator internus from being hurt by the hardness of that part of the ischium over which it passes; also, to hinder it from starting out of its place, while the muscle is in action. 4. ^uadratus Femoris, Arises, tendinous and fleshy, from the outside of the tuberosity of the os ischium; and, running transversely, is Inserted, fleshy, into a rough ridge, continued from the root of the large trochanter to the root of the small one. Use. To roll the thigh outwards. 248 Muscles of the Leg. Muscles situated on the Thigh. These are called muscles of the leg: and consist of one.t on the outside; two, on the inside; four, before; and four, behind. Previous to the description of the muscles that are situ- ated on the thigh and leg, it is necessary to take notice of a broad tendinous fascia or sheath, which is sent off from the back and from the tendons of the glutei and adjacent muscles. It is a strong thick membrane on the outside of the thigh and leg; but, towards the inside of both, it gradually turns thinner, and has rather the appearance of cellular substance, than a tendinous membrane. A little below the trochanter major, it is firmly fixed to the linea aspera; and, farther down, to that part of the head of the tibia that is next the fibula; where it sends off the tendinous expansion along the outside of the leg. It serves to strengthen the action of the muscles, by keeping them firm in their proper places while in action, particularly the tendons that pass over the joints, where this membrane is thickest; and it gives origin to a number of the fleshy fibres of the muscles. On the outside is, Tensor Vagina Femoris, Arises, by a narrow, tendinous, and fleshy beginning, from the external part of the anterior superior spinous process of the os ilium. Inserted, a little below the trochanter major, into the inner side of the membranous fascia which covers the out- side of the thigh. Muscles on the Thigh, 249 Use. To stretch the membranous fascia, to assist in the abduction of the thigh, and somewhat in its rotation in- wards. On the inside are, 1. Sartorius, Arises, tendinous, from the anterior superior spinous process of the os ilium, soon grows fleshy, runs down for some space upon the rectus, and going obliquely inwards, it passes over the vastus internus, and, about the middle of the os femoris, over part of the triceps, it runs down farther between the tendon of the adductor magnus and that of the gracilis muscle. Inserted, by abroad and thin tendon, into the inner side of the tibia, near the inferior part of its tubercle. Use. To bend the leg obliquely inwards, or to bring one leg across the other. 2. Gracilis, Arises, by a thin tendon, from the os pubis near the symphysis of these two bones: soon grows fleshy; and. descending by the inside of the thigh, is Inserted, tendinous, into the tibia under the sartorius, Use. To assist the sartorius. Before are, 1. Rectus. Arises, fleshy, from the inferior anterior spinous pro- cess of the os ilium, and tendinous from the dorsum of the ilium, a little above the acetabulum; runs down over the anterior part of the cervix of the os femoris; the fibres not being straight, but running down like the plumage of a feather obliquely outwards and inwards, from a tendon in the middle. Vol. I. 2 I £50 Muscles on the Thigh. Inserted, tendinous, into the upper part of the patella, from which a thin tendon rutis down, on the fore part of this bone, to terminate in a thick strong ligament, which is sent off from the inferior part of the patella, and inserted into the tubercle of the tibia. Use, To extend the leg, and in a powerful manner, by the intervention of the patella, like a pully. 2. Vastus Externus, Arises, broad, tendinous and fleshy, from the root of the trochanter major, and upper part of the linea aspera; its origin being continued from near the insertion of the glu- teus minimus, the whole length of the linea aspera, by fleshy fibres which run obliquely forwards to a middle tendon, where they terminate. Inserted into a large share of the upper part of the pa- tella; and part of it ends in an aponeurosis, which is con- tinued down to the leg, and in its passage is firmly fixed to the head of the tibia. Use, To extend the leg. 3. Vastus Internus, Arises, tendinous and fleshy, from between the forepart of the os femoris and root of the trochanter minor, and from almost all the inside of the linea aspera, by fibres running obliquely forwards and downwards. Inserted, tendinous, into the upper and inside of the patella, continuing fleshy lower than the vastus externus. Part of it likewise ends in an aponeurosis continued down to the leg, and fixed in its passage to the upper part of the tibia. Use, To extend the leg. Muscles qn the Thigh. 251 4. Cruralis, Arises, fleshy, from between the two trochanters of the os femoris, but nearer the minor, and firmly adhering to most of the fore part of the os femoris, and connected to both, vasti muscles. Inserted, tendinous, into the tipper part of the patella, behind the rectus. Use. To assist in the extension, of the leg. N. B. These four muscles before, being inserted, into the patella, have the same effect upon the leg as if they were immediately inserted into it, by means of the strong tendon, or rather ligament, which is sent off from the Inferior part of the patella to the tibia. Behind are, 1. Semitend.inosus, Arises, tendinous and fleshy, in common with the long head of the biceps, from the posterior part of the tubero- sity of the os ischium; and sending down a long roundish tendon, which ends flat} is Inserted into the inside of the ridge of the tibia, a little below its tubercle. Use. To be^d the leg backwards and a little inwards. 2. Sen^membranosus, Arises, tendinous, from the upper arid posterior part of the tuberosity of the ps Uph)u;ra.; sends d^wn a broad fiat teadop, whjch ends, m a fleshy belly, and, in its descent, ruqs at first on the fore pajt of the biceps, and, lower, be- tween it and the semiten4inasus. Inserted, tendinous, into the inner ajid. back part of the head of the tibia. Use, To bend the leg, and bring it directly backwards. 252 3Iuscles on the Thigh. N. B. The two last form what is called the inner ham- string. 3. Biceps Flexor Cruris, Arises, by two distinct heads. The first, called longus, arises, in common with the semitendinosus, from the up- per and posterior part of the tuberosity of the os ischium. The second, called brevis, arises from the linea aspera, a little below the termination of the gluteus maximus, by a fleshy acute beginning, which soon grows broader as it descends to join with the first head, a little above the ex- ternal condyle of the os femoris. Inserted, by a strong tendon, into the upper part of the head of the fibula. Use. To bend the leg. N. B. This muscle forms what is called the outer ham- string; and between it and the inner, the nervus popliteus, the arteria and vena poplitea, are situated. 4. Popliteus, Arises, by a round tendon, from the lower and back part of the external condyle of the os femoris; then runs over the ligament that involves the joint; firmly adhering to it, and part of the semilunar cartilage. As it runs over the joint, it becomes fleshy, and the fibres run obliquely in- wards, being covered'with a thin tendinous membrane. Inserted, broad, thin, and fleshy, into a ridge at the up- per and internal edge of the tibia, a little below its head. Use. To assist in bending the leg, and to prevent the capsular ligament from being pinched. After the leg is bent, this muscle serves to roll it inwards. Muscles on the Leg. 25S Muscles situated on the Leg. These muscles may be arranged in the two general class- es of flexors and extensors of the foot, and flexors and extensors of the toes; but several of them, viz. the tibi- ales and the peronei, produce effects which are differ- ent from flexion or extension. For the accommodation of the student of anatomy, they may be studied in the order of their position, as they lie on the front, on the outside, and on the back of the leg. Muscles on the Front of the Leg. 1. Tibialis Anticus, Arises, tendinous and fleshy, from the middle of that process of the tibia, to which the fibula is connected above; then it runs down fleshy on the outside of the tibia; from which, and the upper part of the interosseous ligament, it receives a number of distinct fleshy fibres; near the extre- mity of the tibia, it sends off a strong round tendon, which passes under part of the ligamentum tarsi annulare near the malleolus internus. Inserted, tendinous, into the inside of the os cuneiforme internum, and posterior end of the metatarsal bone that sustains the great toe. Use, To bend the foot, by drawing it upwards, and, at the same time, to turn the toes inwards. 2. Extensor Proprius' Pollicis Pedis, Arises, by an acute, tendinous, and fleshy beginning, some way below the head and anterior part of the fibula, along which it runs to near its lower extremity, connected to it by a number of fleshy fibres, which descend obliquely towards a tendon. 254 Muscle,* on the Leg. Inserted, tendinous, into the posterior part of the first and last joint of the great toe. Use. To extend the great toe. 3. Extensor Longus Digitorum Pedis, Arises, tendinous and fleshy, from the upper and outei part of the head of the tibia, and from the head of the fibula where it joins with the tibia, and from the interos- seous ligament; also from the tendinous fascia, which covers the upper and outside of the leg by a number of fleshy fibres; and tendinous and fleshy, from the anterior spine of the fibula, almost its whole lengthy where it is in- separable from the peroneus tertius. It splits into four round tendons, under the ligamentum tarsi annulare. Inserted, by a flat tendon, into the root of the first joint of each of the four small toes; and is expanded over the upper side of the toes, as far as the root of the last joint. Use, To extend all the joints of the four small toes. N, B. A portion of this muscle, which is called 4. Peroneus Tertius, Arises from the middle of the fibula, continues down to near its inferior extremity, and sends its fleshy fibres for- wards to a tendon, which passes under the annular liga- ment, and is Inserted into the root of the metatarsal bone that sus- tains the little toe. Use. To assist in bending the foot. Muscles on the Leg. 255 Muscles on the Outside of the Leg, 1. Peroneus Longus, Arises, tendinous and fleshy, from the fore part of the head of the perone, or fibula, the fibres running straight down; also from the upper and external part of the fibula, where it begins to rise into a round edge; as also, from the hollow between that and its anterior edge, as far down as to reach within a hand's breadth of the ankle, by a num- ber of fleshy fibres, which run outwards towards a tendon, that sends off a long round one, which passes through a channel at the outer ankle, in the back part of the inferior extremity of the fibula; then, being reflected to the sinuo- sity of the os calcis, it runs along a groove in the os cu- boides, above the muscles in the sole of the foot. Inserted, tendinous, into the outside of the root of the metatarsal bone that sustains the great toe, and by some tendinous fibres in the os cuneiforme internum. Use, To move the foot outwards, and to extend it a little. 2. Peroneus Brevis, Arises, by an acute fleshy beginning, from above the middle of the external part of the fibula; from the outer side of the anterior spine of this bone; as also, from its round edge externally, the fibres running obliquely out- wards towards a tendon on its external side: it sends off a round tendon which passes through the groove at the outer ankle, being there included under the same ligament with that of the preceding muscle; and a little farther, it runs through a particular one of its own. Inserted, tendinous, into the root and external part of the metatarsal bone that sustains the little toe. 256 Muscles on the Leg. Use. To assist the former in pulling the foot outwards, and extending it a little. Muscles on the Back of the Leg. 1. Gastrocnemius Externus, seu Gemellus, Arises, by two distinct heads. The first head arises from the upper and back part of the internal condyle of the os femoris, and from that bone, a little above its condyle, by two distinct tendinous origins. The second head arises tendinous from the upper and back part of the external condyle of the os femoris. A little below the joint, their fleshy bellies unite in a middle tendon; and, below the middle of the tibia, it sends off a broad thin tendon, which joins a little above the extremity of the tibia with the ten- don of the following. 2. Soleus, seu Gastrocnemius Internus, Arises by two origins. The first is from the upper and back part of the head of the fibula, continuing to receive many of its fleshy fibres from the posterior part of that bone for some space below its head. The other origin be- gins from the posterior and upper part of the middle of the tibia; and runs inwards along the inferior edge of the popliteus towards the inner part of the tibia, from which it receives fleshy fibres for some way down. The flesh of this muscle, covered by the tendon of the gemellus, runs down near as far as the extremity of the tibia; a little above which the tendons of both gastrocnemii unite, and form a strong round chord, which is called tendo Achillis. Inserted into the upper and posterior part of the os cal- cis; by the projection of which the tendo Achillis > at a considerable distance from the tibia. Muscles on the Back of the Leg. 257 Use. To extend the foot, by bringing it backwards and downwards. 3. Plantaris, Arises, thin and fleshy, from the upper and back part of the root of the external condyle of the os femoris, near the inferior extremity of that bone, adhering to the ligament that involves the joint in its descent. It passes along the second origin of the soleus, and under the gemellus, where it sends off a long, slender, thin tendon, which comes from between the great extensors, where they join tendons; then runs down by the inside of the tendo Achillis. Inserted into the inside of the posterior part of the os calcis, below the tendo Achillis. Use. To assist the former, and to pull the capsular liga- ment of the knee from between the bones. It seems like- wise to assist in rolling the foot inwards. 4. Flexor Longus Digitorum Pedis, Profundus, Perforans, Arises, by an acute tendon, which soon becomes fleshy, from the back part of the tibia, some way below its head, near the entry of the medullary artery; which beginning »s continued down the inner edge of this bone by short fleshy fibres, ending in its tendon; also by tendinous and fleshy fibres, from the outer edge of the tibia; and between this double order of fibres, the tibialis posticus muscle lies inclosed. Having passed under two annular ligaments, it then passes through a sinuosity at the inside of the os cal- cis; and, about the middle of the "sole of the foot, divides into four tendons, which pass through the slits of the per foratus; and, just before its division, it receives a consi- derable tendon from that of the flexor pollicis longus. Vol. I. 2 K 258 Muscles on the Foot. Inserted into the extremity of the last joint of the four lesser toes. Use. To bend the last joint of the toes. 5. Tibialis Posticus, Arises, by a narrow fleshy beginning, from the fore and upper part of the tibia, just under the process which joins it to the fibula; then passing through a perforation in the upper part of the interosseous ligament, it continues its origin from the back part of the fibula next the tibia, and from near one half of the upper part of the last named bone; as also, from the interosseous ligament, the fibres running towards a middle tendon, which sends off a round one that passes in a groove behind the malleolus internus. Inserted, tendinous, into the upper and inner part of the os naviculare, being further continued to the os cuneiforme internum and medium; besides, it gives some tendinous filaments to the os calcis, os cuboides-, and to the root of the metatarsal bone that sustains the middle toe. Use. To extend the foot, and to turn the toes inwards. 6. Flexor Longus Pollicis Pedis. Arises, by an acute, tendinous, and fleshy beginning, from the posterior part of the fibula, some way below its head, being continued down the same bone, almost to its inferior extremity, by a double order of oblique fleshy fibres; its tendon passes under an annular ligament at the inner ankle. Inserted into the last joint of the great toe, and generally ■sends a small tendon to the os calcis. Use. To bend the last joint of this toe. On the upper surface of the foot, there is one muscle; •■viz. Muscles on the Sole of the Foot. 259 Extensor Brevis Digitorum Pedis. Arises, fleshy and tendinous, from the fore and upper part of the os calcis; and soon forms a fleshy belly, divisi- ble into four portions, which send off an equal number of tendons that pass over the upper part of the foot, under the tendons of the former. Inserted, by four slender tendons, into the tendinous ex- pansion from the extensor longus which covers the small toes, except the little one; also into the tendinous expan- sion from the extensor pollicis, that covers the upper part of the great toe. Use. To extend the toes. Muscles on the Sole of the Foot. On the sole of the foot there is a strong tendinous mem- brane called Aponeurosis Plantaris, which originates from the tuberosity of the os calcis, and proceeds for- ward to the toes, increasing gradually in breadth. It is divided into three portions. That in the middle is the largest; it protects and covers the short flexor muscles, and the tendons in the middle of the foot. That on the outside, which covers the abductor, and the flexor of the little toe, is next in size. The inter- nal portion, which covers the abductor of the great toe, is the smallest. « The edges of these portions dip down so as to separate the muscles they cover from each other. They are divided into five processes, corresponding with the heads of the metatarsal bones; each of these portions is divided into two bands, which are inserted into each side of the head of each metatarsal bone, and the ten- dons, nerves, and arteries, pass between them. Immediately under the middle portion of this aponeurosis are the common short flexors of the toes, viz. 260 Muscles on the Sole of the Foot. 1. Flexor Brevis Digitorum Pedis, Sublimis Perforatum, Arises, by a narrow fleshy beginning, from the inferior and posterior part of a protuberance of the os calcis, be- tween the abductors of the great and little toes, soon forms a thick fleshy belly, which sends off four tendons that split for the passage of the flexor longus. Inserted into the second phalanx of the four lesser toes. The tendon of the little toe is often wanting. Use. To bend the second joint of the toes. 2. Flexor Digitorum Accessorius, seu Massa Carnea Jacobii Sylvii, Arises, by a thin fleshy origin, from most part of the sinuosity at the inside of the os calcis, which is continued forwards, for some space on the same bone; also by a thin tendinous beginning, from before the tuberosity of the os calcis externally; and, soon becoming all fleshy, is Inserted into the tendon of the flexor longus, just at its division into four tendons. Use, To assist the flexor longus. 3. Lumbricales Pedis, Arise, by four tendinous and fleshy beginnings, from the tendon of the flexor profundus, just before its division, near the insertion of the massa carnea. Inserted, by four slender tendons, into the inside of the first joint of the four lesser toes, and are lost in the tendi- nous expansion that is sent from the extensors to cover the upper part of the toes. Use, To increase the flexion of the toes, and to draw them inwards. Muscles on the Sole of the Foot. 261 On the inside of the foot, and under the common flex- ors, are the muscles which are considered as exclusively appropriated to the great toe, viz. 1. Abductor Pollicis Pedis, Arises, from the internal side of the tuberosity of the os calcis, and from a ligament which extends from this tube- rosity to the sheath of the tendon of the tibialis posticus muscle, and also from the internal and inferior side of this sheath. It likewise arises from that portion of the aponeurosis plantaris, which separates it from the short flexor of the toes, and many of its fibres appear to be con- nected with the ligaments which pass from the posterior to the anterior bones of the foot: as it passes under the cuneiform bone a portion of its lower surface is tendi- nous. It is inseparably connected to the flexor of the great toe, and is inserted into the internal sesamoid bone, and the inferior and internal part of the root of the first bone of the great toe. This muscle not only separates the great toe from the other toes, but it must increase the curvature, or arched form of the foot. 2. Flexor Brevis Pollicis Pedis, Arises, tendinous, from the under and fore part of the os calcis where it joins with the os cuboides, from the os cuneiforme externum, and is inseparably united with the abductor and adductor pollicis. Inserted into the external os sesamoideilm and root of the first joint of the great toe. Use, To bend the first joint. 262 Muscles on the Sole of the Foot. 3. Adductor Pollicis Pedis, Arises, by a long thin tendon, from the os calcis, from the os cuboides, from the os cuneiforme externum, and from the root of the metatarsal bone of the second toe. Inserted into the external os sesamoideum, and root of the metatarsal bone of the great toe. Use, To bring this toe nearer the rest. Near the outer edge of the foot, under the second por- tion of the aponeurosis plantaris, are the muscles peculiar to the little toe, viz. 1. Abductor Minimi Digiti Pedis, Arises, tendinous and fleshy, from the semicircular edge of a cavity on the inferior part of the protuberance of the os calcis, and from the root; of the metatarsal bone of the little toe. Inserted into the root of the first joint of the little toe externally. Use, To draw the little toe outwards from the rest, and assist in preserving the arched form of the foot. 2. Flexor Brevis Minimi Digiti Pedis, Arises, tendinous, from the os cuboides, near the sulcus or furrow for lodging the tendon of the peroneus longus; fleshy from the outside of the metatarsal bone that sustains this toe, below its protuberant part. Inserted into the anterior extremity of the metatarsal bone, and root of the first joint of this toe. Use, To bend this toe. External Interosseous Muscles. 263 Between the metatarsal bones are four external and three internal interossei: and one muscle which is common to all the metatarsal bones. Interossei Pedis Externi, Bicipites. 1. Abductor Indicis Pedis, Arises, tendinous and fleshy, by two origins, from the> root of the inside of the metatarsal bone of the fore toe, from the outside of the root of the metatarsal bone of the great toe, and from the os cuneiforme internum. Inserted, tendinous, into the inside of the root of the first joint of the fore toe. Use. To pull the fore toe inwards from the rest of the small toes. 2. Adductor Indicis Pedis, Arises, tendinous and fleshy, from the roots of the me- tatarsal bones of the fore and second toe. Inserted, tendinous, into the outside of the root of the first joint of the fore toe. Use. To pull the fore toe outwards towards the rest. 3. Adductor Medii Digiti Pedis, Arises, tendinous and fleshy, from the roots of the me- tatarsal bones of the second and third toes. Inserted, tendinous, into the outside of the root of the first joint of the second toe. Use. To pull the second toe outwards. 4. Adductor Tertii Digiti Pedis, Arises, tendinous and fleshy, from the roots of the me- tatarsal bones of the third and little toe. Inserted, tendinous, into the outside of the root of the first joint of the third toe. Use. To pull the third toe outwards. 264 Internal Interosseous Muscles. Interossei Pedis Interni. 1. Abductor Medii Digiti Pedis, Arises, tendinous and fleshy, from the inside of the root of the metatarsal bone of the middle toe internally. Inserted, tendinous, into the inside of the root of the first joint of the middle toe. Use. To pull the middle toe inwards. 2. Abductor Tertii Digiti Pedis, Arises, tendinous and fleshy, from the inside and infe- rior part of the root of the metatarsal bone of the third toe. Inserted, tendinous, into the inside of the root of the first joint-of the third toe. Use. To pull the third toe inwards. 3. Abductor Minimi Digiti Pedis, Arises, tendinous and fleshy, from the inside of the root of the metatarsal bone of the little toe. Inserted, tendinous, into the inside of the root of the first joint of the little toe. Use. To pull the little toe inwards. The common muscle, Transversalis Pedis, Arises, tendinous, from the under part of the anterior extremity of the metatarsal bone of the great toe, and from the internal os sesamoideum of the first joint, adher- ing to the adductor pollicis. Inserted, tendinous, into the under and outer part of the anterior extremity of the metatarsal bone of the little toe, and ligament of the next toe. Use. To contract the foot, by bringing the great toe and the two outermost toes nearer each other. ALPHABETICAL ARRANGEMENT OF THE MUSCLES. Page ABDUCTOR indicis ma- nus, 242 indicis pedis, 263 minimi digiti ma- nus, 239 minimi digiti pe- dis, 262, 264 oculi, 173 pollicis manus, 238 pollicis pedis 261 medii digiti pedis, 264 tertii digiti pedis, 264 Accelerator urinae, 205 Adductor brevis femoris, 244 indicis pedis, 263 longus femoris, 243 medii digiti pedis, 263 magnus femoris, 244 metacarpi minimi digiti manus, 239 oculi, 173 pollicis manus, 238 pollicis pedis, 262 tertii digiti pedis, 263 Anconeus, 230 Ani sphincter, 206, 209 levator, 207, 209 Anterior auris, 1 70 Arytenoideus obliquus, 192 transversus, id. Arytseno-epiglottideus, id. Attollens aurem, 170 Auris retrahentes, 171 Azygos uvulae, 189 Biceps flexor cubiti, 228 flpxor cruris, 252 Brachialis internus. 229 Buccinator, 178 Vol. I. Page Capitis obliquus superior, 222 obliquus inferior, id. Cervicalis descendens, 218 Circumflexus, or Tensor pa- lati, 187 Clitoridis erector, 208 Coccygeus, 213 Colli longus, 193 transversalis, 221 semispinalis, id. interspinales, 223 intertransversales, 224 Complexus, 219 Compressor naris, 175 Constrictor isthmi faucium, 188 pharyngis, 189, 190 Coraco-brachialis, 227 Corrugator supercilii, 170 Cremaster, 204 Crico-arytenoideus lateralis, 191 arytenoideus posticus, id. thyroideus, 185 Cruralis, 251 Cubit,or fore-arm,muscles of, 233 Cuculearis, 214 Cutaneus, 181 Dartos, 204 Deltoides, 227 Depressor anguli oris, 177 labii inferioris, id. labii superioris a- laequenasi, 176 oculi, 173 Diaphragma, 209 Digastricus, 182 Dorsi interspinales, 224 latissimus. 215 spinalis, 217 2 I. 266 Alphabetical Arrangement of the Muscles. Page Dorsi longissimus, 2is scmispinalis, 220 Erector clitoridis, 208 penis, 204 Extensor brevis digitorum pedis, 259 carpi radialis brevior, 235 carpi radialis longior, 234 carpi ulnaris, 235 digitorum communis, id. longus digitorum pe- dis, 254 ossis metacarpi pol- licis manus, 236 primi internodii pol- licis manus, 237 proprius pollicis pe- dis, 253 secundi internodii pol- licis manus, 237 Extremities superior, mus- cles of, 225 inferior, mus- cles of, 243 Eyeball, muscles of, 172 Eyelids, muscles of, 171 Faucium isthmi constrictor, 188 Flexor accessorius digitorum pedis, 260 brevis digitorum pe- dis, id. brevis minimi digiti pedis, 262 brevis pollicis manus, 238 brevis pollicis pedis, 261 longus digitorum pe- i dis, 257 carpi radialis, 232 carpi ulnaris, id. longus pollicis manus, 233 longus pollicis pedis, 258 ossis metacarpi polli- cis, 238 parvus minimi digiti manus, 239 profundus perforans, c\ o n 400 sublimis perforatus, id. Gastrocnemius externus, Page 256 internus, id. Gemellus, 256 Gemini, 247 Genio-hyo-glossus, Genio hyoideus, Gluteus maximus, 183 id. 245 medius, 246 minimus, Gracilis, id. 249 Humeri os, muscles situated on 228 Hyo-glossus, 184 Jaw lower, muscles of, 179 Iliacus internus, 212 Indicator, 236 Infra-spinatus, 225 Intercostales externi, 196 interni, id. Interosseus auricularis, 241 Interspinales colli, 223 dorsi et lumborum, 224 Intertransversales colli, id. dorsi, id. lumborum, id. Isthmi faucium constrictor, 188 Labii inferioris depressor, 176 inferioris levator, id. superioris alaeque nasi levator, 176 Latissimus dorsi, 215 Leg, muscles situated on, 253 Levator anguli oris, 176 ani, 207, 209 labii inferioris, 177 labii superioris alae-v que nasi, 176 oculi, 173 palati, 187 palpebrae superioris, 172 scapulae, 220 Lingualis, 184 Lips, muscles of, 175 Longissimus dorsi, 218 Longus colli, 193 Lumborum intertransversa- les, 224 Alphabetical Arrangement of the Muscles. 267 Page Page Lumborum quadratus, 212 Palato-pharyngeus, 188 Lumbricales manus, 239 Palmaris, brevis, 237 pedis, 260 longus, 231 Palpebrae superioris levator, 172 Masseter, 180 Palpebrarum orbicularis, 171 Mouth, muscles of, 175 Pectinalis, 243 Multifidus spinae, 220 Pectoralis major, 194 Musculus cutaneus. 181 minor, 195 Mylo-hyoideus 183 Penis erector, 204 Perinei transversus, 205 , 208 Naris compressor, 175 Peroneus brevis, 255 Nose, muscle of, id. longus, id. tertius, 254 Obliquus ascendens internus 201 Pharynx, muscles on the pos- capitis inferior, 222 terior part of it, 189 capitis superior, id. constrictor, 189, 190 descendens externus, * 197 Plantaris, 257 inferior oculi, 174 Platysma myoides. 181 superior, seu troch- Popliteus, 252 Iearis, id. Posterior annularis, 242 Obturator internus, 213 indicis, 240 externus, 244 medii, 241 Occipito-frontalis, 169 Prior annularis, 240 Oculi abductor, 173 indicis, id. adductor, id. medii, 241 depressor, id. Pronator radii quadratus, 234 elevator, id. teres, 232 obliquus inferior, 174 Psoas magnus, 212 » superior, id. parvus, id. Omo-hyoideus, 184 Pterygoideus externus, 181 Opponens pollicis, 238 internus, 180 Orbicularis oris, 179 Pyramidalis, 203 palpebrarum, 171 Pyriformis, 246 Oris anguli depressor, 177 levator anguli, 176 Quadratus femoris, 247 orbicularis, 179 lumborum, 212 Palati circumflexus, 187 Rectus, 249 levator, id. abdominis, 202 tensor, id. capitis internus major, 193 * The following account of the use of the obliquus externus abdo- minis was by accident omitted, viz. These muscles compress the abdomen, and therefore contribute to the evacuation of its contents: if the diaphragm is in a passive state they force it upwards, by press- ing the abdominal viscera against it; and thus assist in producing expiration and its various modifications of coughing, sneezing, Sec. They bend the spine forwards, or approach the thorax tp the pelvis. When one acts separately, it bends the trunk obliquely to the side on which it is situated. 268 Alphabetical Arrangement of the Muscles. Page Rectus capitis internus minor, 193 lateralis, posticus major, posticus minor, Retrahentes auris, Rhomboideus, 194 221 222 171 216 Sacro-lumbalis, 218 Sartorius, 249 Scalenus anticus, 223 medius, id. posticus, id. Scapulae levator, 220 Semimembranosus, 251 Semispinalis colli, 221 dorsi, 220 Semitendinosus, 251 Seminis ejaculator, 205 Serratus magnus, 195 posticus inferior, 215 posticus superior, 217 Soleus, 256 Sphincter ani, 206, 209 vaginae, 208 Spinalis dorsi, 217 Spinae multifidus, 220 Splenius, 216 capitis, 217 colli, id. Sterno-cleido-mastoideus, 182 Sterno-hyoideus, 184 Sterno-thyroideus, 185 Stylo-glossus, 186 Stylo-hyoideus, id. Stylo-pharyngeus, id. Pag« Subclavius, 195 Subscapularis, 228 Supercilii corrugator, 170 Supinator radii brevis, 236 longus, 234 Supra-spinatus, 225 Temporalis, 179 Tensor palati, 187 vaginae femoris, 248 Teres major, 226 minor, id, Thigh, muscles situated on, 248 191 192 185 253 258 219 202 221 264 205, 208 Thyreo-arytenoideus, Thyreo-epiglottideus, Thyreo-hyoideus, Tibialis anticus, posticus, Trachelo-mastoideus, Transversalis, colli, pedis, Transversus perinei, Trapezius, seu Cucularis, 21 - Triangularis, 197 or Sterno-costalis, id. Triceps adductor femoris, 243 extensor cubiti, 230 Vastus externus, 250 internus, id. Urinae accelerator, 205 Uvulae azygos, 189 Zygomaticus major, 178 minor, id. SYSTEM OF ANATOMY. PART III OF THE LIGAMENTS AND MEMBRANES AVHICH CONNECT THE DIFFERENT PARTS OF THE BODY TO EACH OTHER--AND OF THE AR- TICULATIONS. CHAPTER I. OF LIGAMENTS IN GENERAL. THE tendons and the strong membranes connected with them called aponeuroses; the fascia which bind down some of the muscles and afford an origin to many of their fibres; and the membranes which confine the tendons, appear to be composed of the same substance. They consist of fibres which are flexible but extremely strong, and in general have but little elasticity; their sur- faces are smooth and polished; their colour is whitish and silverlike. The vessels which enter into their composition do not commonly carry red blood; and although it seems certain that they must have nerves, many very expert anatomists have declared that no nerves could be traced into them. 270 Of Ligaments in General. In a healthy state they are entirely void of sensibility, and can be cut and punctured, or corroded with caustic applications, without pain. When inflamed, they are ex- tremely painful. The ligaments, which connect the different bones to each other, have a very strong resemblance to these tendinous parts, not only in their structure but in their qualities also. Many of them appear rather more firm in their texture and more vascular. Their vessels are also larger: their colour sometimes inclines to a dull white, and when examined chemically, they appear to differ in some res- pects from tendons. They agree however with the tendinous parts as to their insensibility in a sound state, and the extreme pain which occurs when they are inflamed. No nerves have been traced into their structure. Notwithstanding the ordinary insensibility of these parts, it was asserted by M. Bichat that several animals who seemed to suffer no pain from cutting, puncturing, or corroding the ligaments of their joints, appeared to be in great agony when these parts were violently stretched or twisted; and he declared this to be the case when all the nerves which passed over the ligaments, and could have been affected by the process, were cut away. He explained by this the pain which sometimes occurs instan- taneously in sprains, in the reduction of luxations, and in other analogous processes. The ultimate structure of these parts is perhaps not perfectly understood. An anatomist of the highest authority, Haller, appears to have considered them as formed of membrane, while a late writer who has paid great attention to the subject, and is also of high authority, M. Bichat, has satisfied himself that their structure is essentially fibrous. Of Ligaments in General. 271 If a tendon, or a portion of tendinous membrane, be spread out, or forcibly extended, in a direction which is transverse with respect to its fibres, it will seem to be con- verted into a fine membrane, and the fibres will disappear to the naked eye. The same circumstances will occur when a ligament is treated in a similar way; but much more force is required. Thus constructed, these parts are perfectly passive por- tions of the animal fabric, and have not more power of motion than the bones with which many of them are connected. But notwithstanding their ordinary insensibility, they often induce a general violent affection of the system, when they are diseased. A high degree of fever, as well as severe pain, attends their acute inflammations; and hec- tical symptoms, in their greatest extent, are often induced by their suppurations. There is another circumstance in their history which is very difficult to reconcile with their ordinary insensibility. They are the most common seats of gouty painful affec- tions. In these cases, pain does not seem to be the simple effect of inflammation; it often occurs as the first symp- tom of the disease; it frequendy exists with great vio- lence for a short time and goes off without inflammation; and it is frequently vicarious with affections of the most sensible and irritable parts. Parts of a tendinous and ligamentous structure do not appear retentive of life, but lose their animation very readily, in consequence of the inflammation and other cir- cumstances which attend wounds. When thus deprived of life, they retain their usual appearance and their texture a long time. The dead 272 Of Ligaments in General. parts separate from the living in large portions, in a way which has a considerable analogy with the exfoliation of bones. The tendons and their expansions, and the various fascia, have the same chemical composition. If boiled a long time, they dissolve completely, and form the substance called by chemists, gelatine, or pure glue. The ligaments differ from them in some respects. When boiled they yield a portion of gelatine, and do not dissolve entirely; but are said to retain their form and even their strength, after very long boiling. The composi- tion of the part so insoluble in water, has not yet been ascertained. 273 CHAPTER II. A GENERAL ACCOUNT OF ARTICULATIONS AND OF BURS.E MUCOSA. SECTION I. Of Articulations. 1 HOSE surfaces of bones which form the movable arti- culations are covered with cartilaginous matter, which has been already described.* They are retained in their relative situations by ligaments, such as have been lately mentioned, which are exterior to the cavities of the arti- culations, and placed in such situations that they permit the motions the joints are calculated to perform, while they keep the respective bones in their proper places. They are invested in a particular manner by a thin deli- cate membrane, which in some joints, as those of the hip and shoulder, seems to be the internal lamen of a stronger , ligament called the capsular; and in other joints, the knee for example, appears to be independent of any other structure. In each case, this synovial membrane, as it has lately been called, forms a complete sack or bag, which covers the articular surface of one bone, and is reflected from it to the corresponding surface of the other; adhe- ring firmly to each of the articulating surfaces, and ex- tending loosely from the margin of one surface to that of the other. In this distribution it supplies the place of perichon- drium to the cartilages, and of periosteum to those surfa- ces of bone with which it is connected. « See page 4. Vol. I. 2 M 274 Of Articulations in general. It seems greatly to resemble the membranes which line the abdomen and thorax, and invest the parts contained in these cavities; and like them it may be termed a reflect- ed membrane. It is thin and very flexible, but dense and strong. It secretes, or effuses from its surface, a liquor called synovia; which is particularly calculated to lubricate parts that move upon each other. This fluid is nearly transparent: it has the consistence of a thin syrup, and is very tenacious or ropy. It mixes with cold water, and when heated becomes milky, and deposits some pellicles without losing its viscidity. It ap- pears to be composed of eighty parts in one hundred of water; above eleven parts of fibrous matter; and between four and five parts of albumen. It also contains a small portion of soda, of muriate of soda, and of phosphate of lime. There are in many of the joints masses of fat which appear to project into the cavity, but are exterior to the synovial membrane, and covered by it; as the viscera in the abdomen are covered by the peritoneum. They are generally situated so as to be pressed gendy, but not bruised, by the motions of the bones. In some joints they appear like portions of the common adipose membrane; in others they appear more vascular, and have a number of bloodvessels spread upon them. Small processes often project from their sides like fringe. ' These masses have been considered as synovial glands; but they do not appear like glands; and it is probable that the synovia is secreted by the whole internal surface of membrane. The synovial membrane, like the other parts of joints. Of Bursa Mucosa. 275 is insensible in a sound state, but extremely painful when inflamed. The synovia which is secreted, during the inflammation of this membrane, has a purulent appear- ance. SECTION II. Of Bursa Mucosa. THERE are certain membranous cavities called bursa mucosae, which are found between tendons and bones, near the joints, and in other places also; which have so strong a resemblance to the synovial membrane, and are so intimately connected with some of the articulations, that they ought tow to be mentioned. They are formed of a thin dense membrane, and are at- tached to the surrounding parts by cellular substance; they contain a fluid like the synovia; and sometimes there are masses of fat, which although exterior to them, appear to project into their cavities. There is commonly a thin cartilage, or tough membrane, between them and the bone on which they are placed. They often communicate with the cavities of joints, without inducing any change in the state of the part. As they are always situated between parts that move upon each other, there is the greatest reason to believe that they are intended to lessen friction.* These bursae mucosas are very numerous, as will appear from a subsequent account of them. Several of them are very interesting on account of their connexion with very important joints. * For further information respecting this subject, as well as joints in general, the reader is referred to a Description of the Bursae Mucosae of the Human Body, by Alexander Monro; to whom the world is so much indebted for the elucidation of many important point* in anatomy »nd physiology 276 CHAPTER III. OF PARTICULAR ARTICULATIONS. The Connexion of the Head zvith the Vertebra. 1 HE condyles of the occipital bone, and the corres- ponding cavities of the atlas, are covered with cartilage. The condyle and cavity on each side are invested with a synovial ligament, as described in the general account of articulations. An anterior ligament descends from the front part of the great occipital foramen, and is inserted into all the front part of the atlas, between its articulating processes. That portion of this ligament which is in the middle, and in- serted into the tubercle of the atlas, appears stronger, and is distinct from the rest of it. A posterior ligament passes from the posterior margin of the occipital foramen to the upper edge of the posterior arch of the atlas. From each side of the upper end of the toothlike pro- cess of the vertebra dentata, a ligament passes upwards and outwards, to be inserted into the internal side of the basis of each condyle of the occipital bone. From the anterior margin of the great occipital fora- men, a ligament passes down on the inside of the verte- bral cavity, over the toothlike process, which is inserted in the body of the vertebra dentata and the ligaments connected with it. There is also a ligament which runs across from one side of the atlas to the other, to confine the toothlike pro- Particular Articulations. 277 cess in its anterior cavity. This ligament adheres above to the occipital bone, and below to the body of the ver- tebra dentata. The anterior surface of the toothlike pro- cess plays on the anterior arch of the atlas; the posterior surface plays on this ligament. A synovial capsule is pla- ced on each surface of the toothlike process. The articulating surfaces of the oblique processes of the atlas and vertebra dentata on each side, are invested by a synovial membrane. There are also additional ligaments placed before and behind these processes, that have an ef- fect on their motions. The uses of these different ligaments are very obvious when they are dissected. The transverse ligament of the atlas, with the synovial membranes, forms an arti- culation for the toothlike process, which is of a peculiar kind. The ligaments that pass from this process, to the bones of the condyles of the occipital bone, must have an effect in restraining the rotation of the head and at- las on this process, and therefore have been called mo- derator ligaments. The Articulations oft/ie Vertebra with each other. To acquire a perfect idea of the construction of the Spine, it is necessary to examine at least two preparations of it; in one of which the bodies of the vertebrae should be sawed off from the processes, so that the spinal canal may be laid open. The bodies of all the vertebra, except the atlas, are connected to each other by the intervertebral cartilaginous matter described in page 72, which unites them very firm- ly, at the same time that it allows of some motion, in con- sequence of its elasticity and compressibility. This con- nexion is strengthened by two ligaments, which extend '278 Articulations of the Vertebra. the whole length of the spine, from the second cervical vertebra to the sacrum. The first of these, denominated the anterior vertebral ligament, covers a considerable part of the anterior sur- face of the bodies of the vertebrae; it is thickest in the middle, and varies in its breadth, in different parts of the vertebral column; it adheres very firmly to -the interver- tebral substance, and not so firmly to the bodies of the vertebrae. It has the shining silverlike appearance of ten- don, and seems to consist entirely of longitudinal fibres. There are many fibres which appear to be connected with it, that do not extend the whole length of the spine. On the posterior surface of the bodies of the vertebras, in the cavity which contains the spinal marrow, is the pos- terior or internal vertebral ligament, which like the ante- rior, extends from the upper part of the spine to the sa- crum. In its progress downwards it is broader where it is in contact with the intervertebral matter, and narrower about the middle of each of the bodies of the vertebrae. It ap- pears to consist of longitudinal tendinous fibres, which are similar to those of the anterior ligament. The oblique processes of the vertebras are covered with cartilage, and are invested with a synovial membrane, like the other movable articulations. In the neck and back these membranes are thin and delicate; but in the loins they are blended with ligamentous fibres, which give them additional strength. Some of the most curious and interesting ligaments of the spine, or indeed of the body, are those which are at- tached to the bony plates or arches that extend from the oblique to the spinous processes of each vertebra. These plates form a great portion of the posterior part of Articulations of the Vertebra. 279 the vertebral canal; and the vacant spaces between them are filled up by these ligaments, which extend from the plates of each upper vertebra to those of the next vertebra below. They are situated between the spinal process and the oblique processes on each side. There are, therefore, two distinct ligaments between the two vertebrae, one on each side of the spinal process,' and as they extend only from the plates or arches of one vertebra to those of the other, they must necessarily be very short. They are much more conspicuous on the in- ternal surface of the vertebral cavity than they are exter- nally. They are thick and substantial, and very elastic; their colour resembles that of yellowish adeps; and from that circumstance they are called by some anatomists the yellow ligaments. They complete the cavity for the spinal marrow. As the plates or arches to which they are connected must recede from each other, when the spine is bent for- wards, it seems necessary that they should be elastic. There are also ligaments between the spinous processes, which extend from the under surface of one spinous pro- cess to the upper surface of the spinous process below it. These are composed of tendinous shining fibres, and are sufficiently loose to permit the anterior flexure of the ver- tebral column. From their situation they are denominated interspinal ligaments. There is also a thin and narrow ligamentous band, which extends from the spinous process of the seventh cervi- cal vertebra to the spinous processes of the os sacrum, and adheres to the ends of the intermediate spinous pro- cesses. It is exterior to the tendinous origins of the tra- pexii and latissimi dorsi muscles. The upper portion is 280 Articulation of the Lower Jaxv. slightly connected to the trapezius the lower part adheres more firmly to the'latissimus dorsi. The ligamentum nucha, as it has been denominated, is a narrow but firm strip, which extends from the spinous process of the last cervical vertebra, to the occipital bone, at or near its protuberance. That portion of the trapezius muscle, which is between the occipital bone and the se- venth cervical vertebra, originates from it, or is intimately connected with it; and a portion of the splenius muscle is also connected with it. From the internal surface of this ligament, a thin ten- dinous membrane arises, whose fibres run obliquely up- wards and forwards, and are inserted into the spinous pro- cesses of each of the cervical vertebrae above the seventh, and also into the atlas and the os occipitis. Attached to the ligamentum nuchae and to the spine, this membrane seems like a partition between the muscles which lie on each side of the back of the neck. After inspecting the different ligaments of the spine, it will be obvious that the yellow ligaments are among the most important of them, in consequence of their posi- tion, their strength, and their elasticity. Articulation of the Lower Jaw. The glenoid cavity of the temporal bone with the tu- bercle before it, and the condyle of the lower jaw, are co- vered with cartilages. A cartilage is placed between them, which being flexible, is accommodated to the convexity of the condyle and hollowness of the glenoid cavity, and also to the figure of the aforesaid tubercle to which it is ex- tended. A synovial capsule or bag invests the glenoid cavity and the tubercle, and covers the upper surface of the cartilage. A second capsule of the same kind is attach- A rticulation of the Clavicle and Sternum. 281 ed to the condyle of the lower jaw and the lower surface of the cartilage. A few ligamentous fibres extend from the circumference of the cavity and tubercle of the temporal bone, over both synovial capsules and the cartilage between them, to the lower jaw below the condyle, and appear to be attached to the cartilage. These fibres are collected in such numbers, on the ex- ternal and internal sides of the articulation, that they have been called the external and internal lateral ligaments. Another ligament is mentioned which arises from the styloid process of the temporal bone, and is inserted into the lower jaw near its angle; but this seems rather appro- priated to the stylo-glossus muscle than to this articula- tion. In consequence of this structure, the condyle of the lower jaw moves out of the glenoid cavity upon the tubercle, when the mouth is opened widely. Articulation of the Clavicle and Sternum. The connexion of the clavicle and sternum resembles strongly that of the lower jaw and temporal bone. A movable cartilage is placed between the articulating sur- faces, with a distinct synovial capsule on each side of it, applied in the usual manner to the corresponding surface of the clavicle and of the sternum. Exterior to these cap* sules and the intervening cartilage, are many ligamentous fibres, which are most numerous on the anterior and pos- terior surfaces, but diverge from each other as they pro- ceed from the clavicle to the sternum, and are therefore called Radiated Ligaments. There is a strong ligament, called the Interclavicular, which passes across the sternum internally, from one cla- vicle to the other* Vol. I. 2 N 282 Articulation of the Shoulder. And another ligament, which arises from the interior rough surface of the clavicle, near the sternum, which is inserted into the cartilage of the first rib. This is called the Rhomboid, or Costo clavicular ligament. Articulation of the Clavicle and Scapula. The small surfaces of the clavicle and scapula which are in contact with each other, are furnished with the appara- tus of a movable articulation. They are covered with cartilage, and are invested with a small synovial capsule. The upper and lower surfaces of the extremities of the clavicle and acromion are covered by a ligamentous mem- brane, which is called, from its situation, the superior and inferior ligament of this articulation. But these bones are more firmly connected by the ligament which passes to the coracoid process of the scapula from the under side of the clavicle, and is very strong. Some of the fibres which compose this ligament are so arranged that they have the appearance of an inverted cone: the remaining fibres ap- pear like another ligament, and therefore they have been called the Trapezoid and Conoid ligaments. By their situation and strength they are enabled to re- tain the bones in their proper relative positions, at the same time that they permit a peculiar rotatory motion. Articulation of the Os Humeri and Scapula. The spherical portion of the upper extremity of the os humeri is the part of that bone which is principally con- cerned in the articulation, and is covered with cartilage; as is also the glenoid cavity of the scapula. The glenoid cavity of the scapula, which is so small in trie dried bone when compared with the head of the os humeri, is enlarged by the long tendon of the biceps mus- Articulation of the Shoulder. 283 tie, which is attached to the upper edge of its margin, and then divides and passes down on each side of the cavity, increasing the breadth of it considerably. It appears to be blended with the cartilage that lines the cavity, and also with the capsular ligament which is exterior to it. The articulating surface, thus composed, is perfectly regular and uniform. The synovial ligament in this articulation, is so blended with an external stronger ligament that it cannot be sepa- rated in the recent subject; but notwithstanding, it is ap- plied to the articulating surfaces in the same way that it is applied to the other joints forming a capsule. The stronger exterior lamen is of course only applied to that part of the synovial capsule which proceeds from the mar- gin of one cartilaginous articulating surface to the other; it appears to be most intimately connected with the peri- osteum, and is rendered more firm and thick in particular parts by the addition of fibres from the tendons of the su- pra and infraspinatus, and subscapularis muscles with which it is blended. It arises from the scapula at a small distance from the margin or edge of the glenoid cavity, as formed by the tendon of the biceps, and is inserted into the os humeri at a small distance from the edge of the cartilaginous articu- lating surface; and if dissected away from the bones, would appear like a cylindrical bag with both extremities open. The long tendon of the biceps muscle, in the groove at the head of the os humeri, appears to penetrate this liga- ment, but it is not within the cavity of the synovial mem- brane; for this membrane sends down a process like the finger of a glove, which lines the groove, and is reflected from its surface upon the surface of the tendon, and co- vers it during its whole extent, being reflected from the 284. Articulation of the Elbow. tendon, at its upper termination, to the adjoining surface: so that the tendon is in fact outside of the synovial capsule, which, xiiu'refore, confines the synovia completely. Tr is capsular ligament, which is one of the strongest, would not avail much in keeping the bones in their proper situations, if the muscles and their tendons were not dis- posed in such a manner that, when the muscles act, their power is excited to the same effect. In some cases of pa- ralytic affection, where the muscles exert no influence, the weight of the arm, when it is allowed to hang without sup- port, draws the head of the os humeri below the glenoid cavity, notwithstanding the capsular ligament. At the same time it ought to be observed, that this ligament must be lacerated in every case of complete luxation of the os humeri, as it cannot possibly distend sufficiently to permit the separation of the bones to the extent which then takes place. The Articulation of the Elbow. Those surfaces of the os humeri, ulna, and radius, which move upon each other, are covered with cartilage. The motion of the ulna and radius on the os humeri is that of simple flexion and extension. The cylindrical head of the radius performs a part of a revolution, nearly on its own axis, without moving from the depression in the side of the ulna, with which it is in contact. The synovial membrane adheres very firmly to the sui * fece covered with cartilage on each of the bones, and is reflected from the margin of this surface on one bone, to that of the others. As the principal motion performed is hingelike, the principal ligaments are on the sides. There is also a circular ligament, which arises from the ulna and invests the narrow part of the radius immediately below Articulation of the Wrist. 285 its cylindrical head, like a loop, to confine the radius in contact with the ulna, and at the same time permit its mo- tion. This ligament is so blended with the synovial membrane that it sometimes cannot be separated from it. The lateralligaments are denominated from their origin and insertion, Brachio radial, and Brachio cubital, or Inter- nal and External. The ligament which invests the neck of the radius is called Coronary, or Orbicular. There are also some ligamentous bands, which run up- on the front and back parts of the joint to strengthen it, which are called Anterior and Posterior accessary liga- ments. Within the synovial membrane, in the upper mar- gins of the depressions for the olecranon and coronoid pro- cesses of the ulna, are the adipose substances usually formed in joints. Articulation of the Wrist. The structure of the wrist is particularly complex, he- cause it consists of three articulations, which are con- tiguous to each other, viz. That of the ulna and radius; of the radius and first row of carpal bones; and of the first and second row of carpal bones with each other. An oblong convex head is formed by the upper surfaces of the scaphoides and lunare, and a portion of the upper surface of the cuneiform bone. This head is covered by one cartilage, which is so uniform that the different bones cannot be distinguished from each other. The lower end of the radius is articulated with this head, but does not cover the whole of it; a portion of it therefore is under the ulna, but not in contact with that bone; for the cartilage which lines the concavity of the radius is continued be- yond the radius, so as to cover the remainder of the head. 286 Articulation of the Wrist. The lower end of the ulna is in contact with the upper surface of this cartilage, and is articulated laterally with the semilunar cavity of the radius. This semilunar cavity is lined by a cartilaginous process, continued from the up- per surface of the aforesaid cartilage; so that the extremity and the side of the ulna play upon the cartilage continued from the radius. This articulation of the ulna and radius is distinct from that of the radius and carpus. A synovial membrane covers the articulating head form- ed by the three bones of the carpus, and is reflected from the margin of their cartilaginous surface, to the cartilage at the end of the radius. A plait or fold of this membrane passes from the head of the carpus, at the junction of the sca- phoides and lunare, to the opposite part of the cartilage of the radius, and has been called the Mucous ligament. A strong ligament is placed on the internal side of this articulation, which arises from the styloid process of the ulna, and is inserted into the anterior transverse ligament which confines the flexor tendons, and into the ligament of the pisiforme. Another ligament, on the external side, arises from the styloid process of the radius, and is inserted into the sca- phoides, some of its fibres being continued into the afore- said transverse ligament, and the trapezium. There are two broad irregular ligamentous membranes, one of which arises from the anterior margin of the articu- lating surface of the radius, and the other from the poste- rior margin; one of them is inserted anteriorly, and the other posteriorly, into the margin of the corresponding surface of the scaphoides, lunare, and cuneiforme. They adhere to the synovial membrane, but in some places this membrane appears through apertures which are in them. The surfaces bv which the first and second rows of Of the Carpal and Metacarpal Bones. 287 carpal bones are articulated with each other, are very ir- regular. The magnum and part of the unciforme form a prominent oblong head, on each side of which is a much lower surface, formed by the trapezium and trapezoides externally, and the remaining portion of the unciforme internally.* The scaphoides, lunare, and cuneiforme, form a cavity which correspond with this head, and also with the lower surface formed by the unciforme; while another surface of the scaphoides is articulated with the trapezium and tra- pezoides. These corresponding surfaces, formed by the two rows of carpal bones, irregular as they are, compose but one articulation, which is capable of a limited flexion and extension. It has a synovial membrane, with two late- ral ligaments, and an anterior and posterior ligament; these last, however, are short, and can be best examined from within, by cutting open the articulation. The bones of each row move laterally upon each other. Their lateral surfaces, which are in contact, are covered with cartilage; and the synovial sack which exists between the first and second row of bones, sends off processes be- tween these surfaces, which are disposed like the ordinary synovial membranes in other articulations; adhering to each of the cartilaginous surfaces, while they communicate with the larger cavity between the two rows. Articulation of the Carpal and Metacarpal Bones, The metacarpal bones are connected to the last row of the carpus by surfaces which are covered with cartilages, and supplied with synovial membranes as the most mova- ble articulations are; but the ligaments which connect these s The palm of the hand is supposed to present for^vd 288 Articulation of the Finger.*. bones do not permit much motion between them. The irregularity of the articulating surfaces of the metacarpal bones of the index and middle finger also contribute to re- strain their motion; and these bones accordingly move less than the other two metacarpal bones, whose surfaces are better calculated for motion. Articulation of the Fingers. The first joint of the fingers has a large synovial mem- brane, which invests the head of the metacarpal bone and the corresponding cavities of the bones of the first pha- lanx. On each side is a strong lateral ligament, which ari- ses from the side of the head of the metacarpal bone and is inserted into the side of the base of the first phalanx. Anteriorly there is also a ligament, which, although thick and strong, is very flexible: posteriorly the expansion of the tendons of the extensor muscle, and the tendons of the interossei, have the effect of a ligament. The different phalanges are articulated with each other in a similar manner. The lateral ligaments are very strong: the tendon of the extensor covers the articulation poste- riorly; and anteriorly, under the flexor tendons, there is a soft, but thick ligamentous substance. The metacarpal bone of the thumb differs greatly from the other metacar- pal bones in its articulation with the wrist, as respects its motions. The articulating surfaces are calculated for late- ral motion as well as flexion and extension; and there are no ligaments which prevent it. The first joint of the thumb resembles considerably that of the fingers, and the second joint resembles the last of the phalanges. Hip Joint. 289 Articulation of the Ribs. The ribs are connected to the bodies of the vertebrae and the intervertebral cartilages, by one articulation, and to the transverse processes of the vertebrae by another: these articulations have the ordinary apparatus for motion, with capsular ligaments, which in one case pass from the heads of the ribs to the bodies of the vertebrae, and in the other from the tubercles to the transverse processes. They are also secured in their positions by ligaments which arise from the transverse processes, and are called the Internal and External Transverse ligaments; and also by ligaments which arise from the oblique processes. These ligaments permit the motions necessary for respiration, and restrain all othe rs The connexion of the ribs anteriorly with their cartilages is such as admits of no motion whatever between them; but the extremities of the cartilages are articulated with the sternum, at the pits on the edges of that bone. In many instances there is no appearance of synovia between the ends of the cartilages and the sternum; but this fluid is mostly to be found in the pits, on the lower extremity of the sternum. The Hip Joint. The acetabulum is lined with cartilage; and the brim or margin of it is much enlarged, and the cavity deepened, by the addition of fibrous cartilaginous matter, which forms a regular smooth edge. This cartilaginous ring is continued across the upper part of the notch in the acetabulum; so that it completes the circular margin of the cavity, but leaves the under part of the notch open. The head of the os femoris is covered with cartilage; but the depression in it is still visible. From this depression a strong ligament Vol. I. 2 0 290 Hip Joint. arises, which appears to pass into the depression, near the center of the acetabulum; but actually terminates in the lower edge of the cartilaginous ring or margin, where it crosses over the notch, and not in the bone. The thin mem- brane with which this ligament is invested extends to the center of the acetabulum, and has given rise to the opi- nion that the ligament was inserted in the bottom of the acetabulum. There is therefore an analogy between the termination of this ligament and that of the long tendon of the biceps flexor cubiti, at the glenoid cavity of the scapula.* This ligament allows the head of the os femoris to rise out of the acetabulum; but it is probably torn in every luxation of the os femoris. The capsular ligament, which contains these articula- ting parts, is the strongest in the body. It arises around the acetabulum, near the basis of the cartilaginous brim; but it does not adhere to the cartilaginous edge; and it is inserted into the os femoris, near the roots of the trochan- ters, so that it includes a large portion of the neck of the bone. It is not every where of the same thickness and strength; for, in various places, there are additional liga- mentous fibres. The largest portion of these additional fibres appears to arise from the inferior anterior spinous process of the ilium. The synovial membrane forms the internal lamen of this ligament: it invests the articulating surfaces in the usual manner, and being reflected from the internal surface of the capsular ligament to the neck of the os femoris, it is in the place of periosteum to that part of the bone. It seems probable that this membrane is so reflected * See page 283. Knee Joint. 291 and arranged, that the internal ligament is covered by it also; and, of course, that this ligament is exterior to the synovial membrane. There is a considerable quantity of adipose matter, near the termination of the aforesaid internal ligament, which is also exterior to the synovial membrane :* some of this can be pressed out of the acetabulum, at the vacuity in the notch under the cartilaginous margin. Articulation of the Knee. The synovial membrane of the knee joint is, in some places, without the support of a proper capsular ligament, or external lamen; so that it is easier distinguished in this articulation than in many others. It adheres firmly to the cartilaginous surfaces of the os femoris, tibia, and patella, and is reflected in the usual manner, from one to the other of these surfaces. It arises closely from the edge of the cartilaginous surface at the top of the tibia; but on the anterior part of the os femoris, it is continued to some distance from the margin of the pulleylike surface, and the edges of the condyles. On each of the portions of the cartilaginous surface of the ti- bia is a cartilage of a semilunar form, so placed, that its convex edge rests on the margin of the cartilaginous sur- face; and its concave edge is internal. These cartilages are thick at their external, and very thin at their internal, edges; so that they form two superficial concavities on the top of the tibia. Their extremities are attached by ligaments to the cen- tral protuberance of the tibia; and their anterior extremi- ties are also connected by a ligament to each other. The synovial membrane is so reflected as to cover the whole surface of these cartilages, except the exterior 292 Knee Joint. edge, which is connected with the external ligaments of the articulation. The use of these cartilages is evidently to form conca- vities on the top of the tibia, for accommodating the con- dyles of the os femoris: and, upon examination, they will not appear so anomalous as they are at first view; for there is a considerable analogy between them and the cartilagi- nous edges of the glenoid cavity, and of the acetabulum. The patella appears to project into the cavity of the joint; and its internal surface is very prominent: around the margin of this surface, and especially at the under part of it, the adipose substance found in joints is very abundant. On each side of the adipose mass, under the patella, is a plait of the synovial membrane, called ligamentum alare minus, and majus; and a process of the membrane, called hgamentum mucosum, passes from the neighbourhood of the adipose mass to the os femoris between the condyles. These processes retain the adipose substance in its pro- per place, during the motions of the joint. There are two very strong ligaments, called the crucial, which arise from the middle protuberance of the tibia, one of which is inserted posteriorly into the external condyle, and the other into the internal. These ligaments decus- sate each other partially; on which account the name cru- cial is applied to them. They are in a state of tension when the leg is extended, and prevent it from moving further forward: when it is bended they are relaxed. They add greatly to the strength of the connexion between the os femoris and tibia. These ligaments are generally supposed to be in the cavity of the joint; but the synovial membrane is reflected round them in such manner that they are exterior to it. In addition to the crucial ligaments, this articulation has the following external supports. Articulation of the Tibia and Fibula. 293 1. Two strong lateral ligaments, one on each side of the knee. The external of which arises from the tubercle above the external condyle of the os femoris, and is attached to the fibula a little below its head; and the internal, from the upper part and tubercle of the internal condyle, and is inserted into the upper and inner part of the tibia. 2. The posterior ligament, whose fibres run obliquely from the external condyle to the back part of the internal side of the head of the tibia. This ligament also prevents the leg from being drawn too far forwards. When the leg is extended, these ligaments are tense: they therefore prevent rotation in the extended state. When the leg is bent, they are relaxed, and therefore ad- mit of that motion. 3. The connexion of the tendons of the extensor muscles of the leg, with this articulation, has a great effect upon it. Their insertion into the patella places them in the situ- ation of the upper part of an anterior ligament, of which the very strong ligament, that passes from the lower margin of the patella to the tubercle of the tibia, is only the lower portion; while the patella may be considered as an indura- ted part of the ligament. Articulation of the Tibia and Fibula. The surfaces of the upper extremities of the tibia and fibula, which are articulated with each other, are very small. When the bones are in their natural position, these sur- faces are nearly horizontal: that of the tibia looking down, and that of the fibula looking up: they are covered with caniLiges, and have a synovial membrane. This^articula- tion is supported by some ligamentous fibres, which have been called anterior and posterior ligaments; but it is Strengthened by the external lateral ligament of the knee, 294 Articulation of the Leg and Foot. and by the tendon of the biceps muscle which is inserted into the upper end of the fibula. Attheir lower extremities, the cartilaginous crust, which, on each of them, forms part of the articulating surface with the astragalus, is turned up on their lateral surfaces, which are in contact with each other; so that a small portion, (equal in breadth only to one-sixth of an inch) of the con- tiguous surfaces, is covered with cartilage: the other parts of these surfaces, which are very considerable, are attached to each other by the intervention of fibrous or membranous matter; and there is very little motion of the bones on each other. There are very strong external ligaments, anteriorly and posteriorly, which connect the fibula to the tibia; and from the posterior end of the fibula a small short ligament passes to the nearest part of the tibia, which resembles the margins of the glenoid cavity and acetabulum; for it en- larges the articulation with the astragalus, while it serves as a ligament of the tibia and fibula. Articulation of the Leg and Foot. It should be observed that the tibia and fibula are so firmly connected with each other below, that they may be considered as forming but one member of this articulation. The varied surfaces formed by the tibia and fibula, and by the astragalus, when it is contiguous to them, are in- vested with the usual apparatus of articulation. The syno- vial fluid is generally observed to be very redundant in this joint. A lateral ligament passes downwards from the tibia, at the internal malleolus, and is inserted into the inside of the astragalus, and also into the os calcis and naviculare. Some of the fibres are blinded with those of the sheath Of the Astragalus with Os Calcis, &fc. 295 for the tendon of the flexor communis; and some of them have a radiated arrangement, in consequence of which this has been called the deltoid ligament. From the fibula three ligaments arise. The middle liga- ment, which is strong and thick, passes downwards from the end of that bone, to be inserted into the outside of the os calcis. The anterior and posterior ligaments pass also from the external malleolus, and are inserted into the anterior and posterior portions of the astragalus. Articulation of the Astragalus and Os Calcis. The astragalus is attached firmly to the os calcis by very strong and short ligamentous fibres, which arise from the fossa on its under surface, and are inserted into the fossa between the upper articulating surfaces of the os calcis. This ligament separates the posterior articulations of the astragalus and os calcis from the anterior. The posterior articulation has a synovial membrane exclusively appro- priated to it. The anterior articulation is supplied by an extension of the membrane which also invests the articu- lating surfaces of the astragalus and naviculare. The connexion of the astragalus, with the os calcis, is supported by the lateral ligaments of the ankle joint, and also by many irregular ligamentous fibres. Articulation of the Astragalus with the Os Naviculare. This articulation appears calculated for considerable motion, as well from the form of the two surfaces concern- ed in it, as the perfect state of their articulating invest- ments. Their motions are restrained, to a certain degree, by ligaments, which are situated on the upper and internal surfaces of the foot. 296 Articulation of the Os Calcis and Cuboides. The ligaments which pass from the anterior internal ex- tremity of the os calcis to the os naviculare, and sup- port the head of the astragalus, ought to be observed with attention during the examination of this joint. Articulation of the Os Calcis and Cuboides. The articulating surfaces of this joint are arranged in the usual manner. There are two additional ligaments: one placed on the upper, and the other on the under, surfaces of the bones. The upper ligament is thin; but the under ligament is one of the strongest of the foot; and its fibres are blended with those which form the sheath for the tendon of the pero- neus longus, as it passes along the groove in the cuboides. 297 CHAPTER IV. OF PARTICULAR LIGAMENTS, AND OF THE SITUATION OF THE INDIVIDUAL BURSAE MUCOSiE, SECTION I. Enumeration of the most important ligaments, which have not been described. Ligaments proper to the Scapula. 1 ilE triangular ligament arises, broad, from the external surface of the coracoid process, and becomes narrower where it is fixed to the posterior margin of the acromion* It confines the tendon of the supraspinatus muscle, and assists in protecting the upper and inner part of the joint of the humerus. The posterior ligament of the scapula is sometimes dou- ble, and is stretched across the semilunar notch of the sca- pula, forming that notch into one or two holes for the pas- sage of the superior posterior scapulary vessels and nerves. It also gives rise to part of the omo-hyoideus muscle. The Interosseous Ligament of the Fore Arm Extends between the sharp ridges of the radius and ulna, filling up the greater part of the space between these two bones, and is composed of small fasciculi, or fibrous slips, which run obliquely downwards and inwards. Two or three of these, however, go in the opposite direction; and one of them, termed oblique ligament, and chorda trans- versalis cubiti, is stretched between the tubercle of the ulna and under part of the tubercle of the radius. In different parts of the ligament there are perforations for the pas- Vol. I. 2 P 298 Ligaments of the Hand. sage of bloodvessels from the fore to the back part of the bone; and a large opening is found at the upper part of it, which is filled up by muscles. It prevents the radius from rolling too much outwards, and furnishes a commodious attachment for muscles. Ligaments retaining the Tendons of the Muscles of the Hand and Fingers in their proper positions. The anterior annular ligament of the wrist is stretched across from the projecting points of the pisiform and un- ciform bones, to the os scaphoides and trapezium, and forms an arch which covers and preserves in their places the tendons of the flexor muscles of the fingers. The vaginal ligaments of the flexor tendons are fine membranes, connecting the tendons of the sublimis, first to each other, and then to those of the profundus; forming, at the same time, bursae mucosae which surround the tendons. The vaginal, or crucial ligaments of the phalanges, arise from the ridges on the concave side of the phalanges, and run over the tendons of the flexor muscles of the fin- gers. Upon the body of the phalanges, they are thick and strong, to bind down the tendons; but over the joints they are rhin, and have, in some parts, a crucial appearance, to allow the ready motion of the joints. The accessory ligaments of the flexor tendons of the fingers are small tendinous fraena, arising from the fiist and second phalanges of the fingers. They run ob- liqiu 1\ forwards within the vaginal ligaments, terminate in the tendons of the two flexor muscles of the fingers, and assist in keeping ihem in their places. The posteror annular ligament of thet wrist is part of the aponeurosis of the fore arm, extending across Ligaments of the Sternum. 299 the back of the wrist, from the extremity of the ulna and os pisiforme to the extremity of the radius. It is connect- ed with the small annular ligaments which tie down the tendons of the extensores ossis metacarpi etprimi interno- dii pollicis, and the extensor carpi ulnaris. The vaginal ligaments adhere to the last mentioned and serve as sheaths and bursae mucosae to the extensor tendons of the hand and fingers. The transverse ligaments, of the extensor tendons, are aponeurotic slips running between the tendons, near the heads of the metacarpal bones, and retaining them in their places. Ligaments on the Anterior part of the Thorax, The membrane proper to the sternum is a firm expan- sion, composed of tendinous fibres running in different di- rections, and covering the anterior and posterior surfaces of the bone, being confounded with the periosteum. The ligaments of the cartilago ensiformis are part of the proper membrane of the sternum, divided into strong bands, which run obliquely from the under and fore part of the second bone of the sternum, and from the cartilages of the seventh pair of ribs, to be fixed to the cartilago ensi- formis. The ligaments covering the sternum serve consi- derably to strengthen that bone. There are also thin tendinous expansions which run over the intercostal muscles at the fore part of the thorax, and connect the cartilages of the ribs to each other. Ligaments of the Bones of the Pelvis. The two transverse ligaments of the pelvis arise from the posterior part of the spine of the os ilium, and run transversely. The superior is fixed to the transverse 300 Ligaments of the Pelvis. process of the last vertebra of the loins; the inferior to the first transverse process of the os sacrum. The ilio-sacral ligaments arise from the posterior spinous process of the os ilium, descend obliquely, and are fixed to the first, third, and fourth, spurious trans- verse processes of the os sacrum. These, with the two transverse ligaments, assist in bind- ing the bones together, to which they are connected. The two sacro-ischiatic ligaments are situated in the uni der and back part of the pelvis. They arise in common from the transverse processes of the os sacrum, and likewise from the under and lateral part of that bone, and from the upper part of the os coccygis. The first, called the large, external, or posterior, descends obliquely, to be fixed to the tuberosity of the os ischium. The other, called the small, internal, or anterior, runs transversely to be fixed to the spinous process of the os ischium. These two liga- ments assist in binding the bones of the pelvis, in support- ing its contents, and in giving origin to part of its muscles. There are two membranous productions which are con- nected with the large sacro-ischiatic ligament, termed its superior and inferior appendices. The superior appendix, which is tendinous, arises from the back part of the spine of the os ilium, and is fixed along the outer edge of the ligament, which it increases m breadth. The inferior, or falciform appendix, situated within the eavity of the pelvis, the back part of which is connected with the middle of the large external ligament, and the rest of it is extended round the curvature of the os is- chium. These two productions assist the large sacro-ischiatic ligament in furnishing a more commodious situation for, Ligaments of the Pelvis, 301 and insertion of, part of the gluteus maximus, and ob- turator internus muscles. Besides the ilio-sacral, and sacro-ischiatic ligaments, several other slips are observed upon the back of the os sacrum, which descend in an irregular manner, and strengthen the connexion between that bone and the os ilium. The large holes upon the back part of the os sacrum are also surrounded with various ligamentous expansions, pro- jecting from one tubercle to another, and giving origin to muscular fibres, and protection to small vessels and nerves which creep under them. A general covering is sent down from the ligaments of the os sacrum, which spreads over and connects the dif- ferent pieces of the os coccygis together, allowing consi- derable motion, as already mentioned in the description of this bone. The longitudinal ligaments of the os coccygis de- scend from those upon the dorsum of the os sacrum, to be fixed to the back part of the os coccygis. The ligaments of this bone prevent it from being pulled too much forwards by the action of the coccygeus muscle, and they restore the bone to its natural situation, after the muscle has ceased to act. The ligamentous cartilage, which unites the two ossa pubis so firmly together as to admit of no motion, except- ing in the state of pregnancy, when it is frequently found to be so much softened as to yield a little in the time of delivery. The obturator membrane, or ligament of the foramen thyroideum, adheres to the margin of the foramen thyroi- deum, and fills the whole of that opening, excepting the oblique notch at its upper part for the passage of the ob- 302 Ligaments of the Foot. turator vessels and nerve. It assists in supporting the con- tents of the pelvis, and in giving origin to the obturator muscles. The interosseous ligament of the leg fills the space be- tween the tibia and fibula like the interosseous ligament of the fore arm, and is of a similar structure; being formed of oblique fibres, and perforated in various places for the passage of vessels and nerves. At the upper part of it there is a large opening, where? the muscles of the opposite sides are in contact; and where vessels and nerves pass to the fore part of the leg. It serves chiefly for the origin of part of the muscles which belong to the foot. Ligaments retaining the Tendons of the Muscles of the Foot and Toes, in their proper positions. The annular ligament of the tarsus is a thickened part of the aponeurosis of the leg, splitting into superior and inferior portions, which bind down the tendons of the extensors of the toes upon the fore part of the ankle. The vaginal ligament of the tendons of the peronei mus- cles behind the ankle is common to both, but divides at the outer part of the foot and becomes proper to each. They preserve the tendons in their places, and are the bursae of these tendons. The laciniated ligament*arises from the inner ankle, and spreads in a radiated manner, to be fixed partly in the cellular substance and fat, and partly to the os cal- cis, at the inner side of the heel. It incloses the tibialis posticus and flexor digitorum longus. The vaginal ligament of the tendon of the extensor pro- prius pollicis runs in a crucial direction. The vaginal ligament of the tendon of the flexor longus Bursa Mucosa, 303 pollicis surrounds this tendon in the hollow of the os cal- cis. The vaginal and crucial ligaments of the tendons of the flexors of the toes inclose these tendons on the surfaces of the phalanges, and form their bursae mucosae. The accessory ligaments of the flexor tendons of the toes, as in the fingers, arise from the phalanges, and are included in the sheaths of the tendons in which they ter- minate. The transverse ligaments of the extensor tendons run between them, and preserve them in their places be- hind the roots of the toes. SECTION II. . Enumeration of the most important Bursa Mucosa. Those about the articulation of the shoulder are situated 1. Under the clavicle, where it plays upon the coracoid process. 2. Between the triangular ligament of the scapula and the capsular ligament of the humerus. 3. Between the point of the coracoid process and cap- sular ligament of the humerus. 4. Between the tendon of the subscapularis muscle and capsular ligament of the humerus, frequently communi- cating with the cavity of that joint. 5. Between the origin of the coraco-brachialis and short head of the biceps muscle, and capsular ligament of the humerus. 6. Between the tendon of the teres major and the os humeri, and upper part of the tendon of the latissimus dorsi. 304 Bursa Mucosa of the Upper Extremity, 7. Between the tendon of the latissimus dorsi and os humeri. 8. Between the tendon of the long head of the biceps flexor cubiti and the humerus. The bursae marked 3 and 5 are sometimes absent. Near the articulation of the Elbow there are, 1. With apelaton of fat, between the tendon of the bi- ceps and tubercle of the radius. 2. Between the tendon common to the extensor carpi radialis brevior, extensor digitorum communis, and round head of the radius. 3. A small bursa, between the tendon of the triceps ex- tensor cubiti and olecranon. On the Fore Arm and Hand are situated, 1. A very large bursa surrounding the tendon of the flexor pollicis longus. 2. Four long bursae lining the sheaths which inclose the tendons of the flexors upon the fingers. 3. Four short bursae on the fore part of the tendons of the flexor digitorum sublimis in the palm of the hand. 4. A large bursa between the tendon of the flexor pol- licis longus, the fore part of the radius, and capsular liga- ment of the os trapezium. 5. A large bursa between the tendons of the flexor digi- torum profundus, and the fore part of the end of the radius and capsular ligament of the wrist. These two last mentioned bursae arc sometimes found to communicate with each other. 7, A bursa between the tendon of the flexor carpi radi- alis and os trapezium. 8. Between the tendon of the flexor carpi ulnaris and os pisiforme. Bursa Mucosa of the Thigh. 305 9. Between the tendon of the extensor ossis metacarpi pollicis and radius. 10. A large bursa common to the extensores carpi radi- ales, where they cross behind the extensor ossis metacarpi pollicis. 11. Another common to the extensores carpi radiales, where they cross behind the extensor secundi internodii pollicis. 12. A third at the insertion of the tendon of the exten- sor carpi radialis brevior. 13. A bursa for the tendon of the extensor secundi in- ternodii pollicis, which communicates with the second bursa common to the extensores carpi radiales. 14. Another bursa between the tendon of the extensor secundi internodii pollicis and metacarpal bone of the thumb. 15. A bursa between the tendons of the extensor of the fore, middle, and ring fingers, and ligament of the wrist. 16. For the tendons of the extensor of the little finger. 17. Between the tendon of the extensor carpi ulnaris and ligament of the wrist. Upon the Pelvis and upper part of the Thigh there are, 1. A very large bursa between the iliacus internus and psoas magnus muscle, and capsular ligament of the thigh bone. 2. One between the tendon of the pectinalis muscle and the thigh bone. 3. Between the gluteus medius and trochanter major, and before the insertion of the tendon of the pyriformis. 4. Between the-tendon of the gluteus minimus and tro- chanter major. Vol. I. 2 Q 306 Bursa Mucosa of the Thigh and Knee. 5, Between the gluteus maximus and vastus externus. 6. Between the gluteus medius and pyriformis. 7. Between the obturator internus and os ischium. 8. An oblong bursa continued a considerab e way be- tween the obturator internus, gemini, and capsular liga- ment of the thigh bone. 9. A small bursa at the head of the semimembranosus and biceps flexor cruris. 10. Between the origin of the semitendinosus and that of the two former muscles. 11. A large bursa between the tendon of the gluteus maximus and root of the trochanter major. 12. Two small bursae between the tendon of the gluteus maximus and thigh bone. About the joint of the Knee are, 1. A large bursa behind the tendon of the extensors of the leg, frequently found to communicate with the cavity of the knee joint. 2. Behind the ligament which joins the patella to the tibia, in the upper part of the cavity of which a fatty sub- stance projects. 3. Between the tendons of the sartorius, gracilis, semi- tendinosus, and tibia. 4. Between the tendons of the semimembranosus and gemellus, and ligament of the knee. This bursa contains a small one within it, from which a passage leads into the cavity of the joint of the knee. 5. Between the tendon of the semimembranosus and the lateral internal ligament of the knee, from which also there is a passage leading into the joint of the knee. 6. Under the popliteus muscle, likewise communicating with the cavity of the knee joint. About the Ankle and the Foot. 307 About the Ankle there are, 1. A bursa between the tendon of the tibialis anticus, and under part of the tibia and ligament of the ankle. 2. Between the tendon of the extensor proprius pollicis pedis, and the tibia and capsular ligament of the ankle. 3. Between the tendons of the extensor digitorum lon- gus, and ligament of the ankle. 4. Common to the tendons of the peronei muscles. S. Proper to the tendon of the peroneus brevis. 6. Between the tendo achillis and os calcis, into the ca- vity of which apeloton or mass of fat projects. *► 7. Between the os calcis and flexor pollicis longus. 8. Between the flexor digitorum longus and the tibia and os calcis. 9. A bursa between the tendon of the tibialis posticus and the tibia and astragalus. On the Sole of the Foot are also, 1. A second bursa for the tendon of the peroneus lon- gus, with an oblong peloton of fat within it. 2. One common to the tendon of the flexor pollicis lon- gus, and that of the flexor digitorum profundus, at the upper end of which a fatty substance projects. 3. Another for the tendon of the tibialis posticus* 4. Several for the tendons of the flexors of the toes. APPENDIX TO THE THREE PRECEDING PARTS Observations on the Motions of the Skeleton. THE falling down of the body during life, when mus- cular action is suspended, as well as the examination of the artificial skeleton, evinces that this machine is not con- structed to preserve the erect position of itself; but thi^* when unsupported, it bends at the joints, and invariably falls forward. It is retained in the erect position by the action of mus- cles; and that the muscles should produce this effect, it is necessary that they should have a fixed basis to act from. This basis is the feet, and they are fixed to the ground by the weight of the body. To keep the body from falling it is necessary that the center of gravity should be immediately over the center of the common basis. All our movements, both in walking, standing, and rising from our seats, are regulated by this principle; and whenever we move the body, so that the center of gravity is changed, we must change the position of the feet, that the center of the basis may be directly under it. If this proposition were not almost self-evident, it might be illustrated by several very easy experiments. If a person stands against a wall, with his heels and the back parts of his legs and thighs in contact with it, and in this situation attempts to stoop forward, he will fall upon his face; there is no power in his muscles, or in any other Adjustment of the Center of Gravity. 301> part of the body, when thus circumstanced, to prevent it; but a small movement forward of one foot, will enable him to stoop with ease by altering the basis of his body. When we sit in such a position that we cannot bring the center of gravity over the feet, the lower limbs are divested of all power of elevating the body: this is always the case when we sit with the thighs and legs at right angles with each other. Bend the knees to an acute angle, so that the feet are placed under the body, and we rise with ease. When we wish to stoop forward without advancing one of our feet, we acquire the power in a small degree, by placing our hands behind us, to preserve the equilibrium. Some old persons, whose spines curve forwards in con- sequence of age, bend their lower limbs, so that the pelvis may be projected backwards beyond the center of the base of the body, and form a counterpoise to the upper part of the trunk. Bending the knees alone without projecting the pelvis backwards, will not produce this effect; for a person who stands with his back to a wall will bend his knees without obtaining this advantage, while the heels and back part of the pelvis are in contact with the wall. When we stand with the toes pointing directly forwards, the base of the body is a square; of which the feet are two of the sides. As the positions of the feet are changed, the figure of the base and its center necessarily changes also. When the feet are placed one immediately before the other, the center is between the toes of the one and the heel of the other. When the position of the feet is such, that the toes point directly outwards, and the heels are opposed to each other, the center of the base is between the heels. 310 Adjustment of the Center of Gravity. In these cases, when the situation of the center of the base is changed, we immediately change the center of gra- vity. Thus, as we turn the toes outwards, the center of the base moves backwards; we therefore immediately make the body more erect; and by that means keep the center of gravity over the center of the base. We move the center of gravity laterally, as well as backwards and forwards, in conformity to this principle. Thus, when we raise one foot from the ground, the body inclines so much in the opposite direction, that the center of gravity is directly over the other. If the spine is dis- eased in one spot, and assumes a lateral curvature, placing the center of gravity on one side of the natural center of the base; another curve is formed by muscular action, in a sound part of the spine, to counteract the first, and keep the center of gravity in its natural position. The perception of a tendency to fall, when the center of gravity is in a wrong situation, first induces us to make efforts to resist this tendency; we learn by experience what these efforts ought to be; and by habit we at length make them without consciousness. As the natural tendency of the skeleton, when we stand, is to bend at the articulations, and therefore to fall for- wards; the muscles which have the principal effort in keep- ing the body erect, must be the extensors. Thus, the muscles on the back of the leg, and particu- larly the soleus, keep the tibia erect: while the muscles on the front of the thigh, the vasti and crureus, produce the same effect upon the os femoris: the bones being kept steady by the occasional counteraction of the antagonist muscles. The whole lower limb is thus made erect by an exertion Muscles which keep the Body erect. 311 which begins at the foot, while the foot is fixed to the ground by the weight and pressure of the body above it. The trunk of the body has a strong tendency to bend forward at the articulations of the thigh bones and the ossa innominata. This tendency is resisted by the muscles which lie on the back part of the ossa femoris, and extend the trunk on those bones, viz., the glutei maximi. The muscles which arise from the tuberosity of the ischium, and are inserted into the leg, the semitendinosus, semimembranosus, and the long head of the biceps flexor cruris, have also this effect. The flexure of the thoracic and lumbar portions of the spine is counteracted by the sacro-lumbalis, and longissi- mus dorsi, which act from the sacrum and back parts of the pelvis. The yellow ligaments, which are elastic, must also cooperate to this effect: so that with regard to the spine, there is an additional agent distinct from the mus- cular power. Indeed, respecting the vertebral articulations in general, it may be observed, that the connexion of the bodies of the vertebrae, by the intervertebral cartilaginous matter, and of the plates behind, by the elastic ligament, renders these articulations perfectly anomalous; and very different in their principles from the articulations in general. In no part of the skeleton is this tendency to bend for- ward more strongly perceived than in the head. When we are awake, and the muscles in a healthy situation, it is effec- tually restrained, and the head kept erect, by the splenius and complexus, and other muscles, which act from the spine below, upon the back part of the head and the ver- tebras of the neck. When we stand on one foot, some very different mus- cles are called into action; the tendency of the body is to 312 Muscles employed in rising from a Seat. fall sideways, towards the foot which is raised from the ground. To counteract this tendency, the two larger pero- nei muscles, which are situated on the outside of the leg, act from the foot, to keep the leg erect. The vastus externus acts upon the same principle from the leg upon the os fe- moris. The gluteus medius and minimus, and the muscle of the fascia lata, act from the os femoris upon the pelvis and trunk; while the quadratus lumborum, and those ab- dominal muscles which draw the spine to that side, con- tinue the operation: and so do likewise the muscles which act on the same side of the neck and head. In rising from a seat, the tibialis anticus acts very pow- erfully, to keep the tibia erect, and prevent it from incli- ning backwards. The two vasti,and the crureus, raise up the os femoris: while the gluteus maximus, the semiten- dinosus, and semimembranosus, and the long head of the biceps, extend the trunk of the body. There are several modes of walking, which are different from each other, in a small degree. We may walk, for example, with the knee of the hind limb straight or bent, as we bring it forward. This circum- stance is merely a matter of accommodation. But there are two essential processes in walking, viz. 1, Projecting one foot forward, and placing it on the ground while thus projected; and 2, Moving the body over that foot. The mode of projecting the foot requires no explana- tion; but the manner of bringing it to the ground when thus advanced, ought to be noticed. If, after standing with both feet on the same line, we move one foot forwards, suppose the right foot, it cannot be applied flat to the ground, unless we either incline the bod) forward: or move the pelvis on the left thigh, so that the right side may present obliquely forward: or lower the Motions necessary in Walking, 313 right side of the pelvis, so that it may be nearer the ground. When we incline the body forward, and thus bring the right foot to the ground, we perform the second essential process in walking along with the first; for we. move the body over the fore foot. The muscles on the front part of the hind leg, and particularly the tibialis anticus, seem to produce this effect, by bending, or inclining forward, the tibia on the foot. When the foot is brought to the ground by a rotation of the pelvis, it is likewise the tibialis anticus, and the muscles on the front of the hind leg, that move the body over it, or that begin the motion. The gastrocnemius and soleus, and the flexors of the toes, particularly that of the great toe, occasionally coope- rate with great effect. By raising the heel, and thus lengthening the hind limb, they push the body forward, and continue its motion in that direction after the effect of the tibialis anticus ceases. The length of the step appears therefore to require this elevation of the heel, and depression of the toes; but it should be observed, that when we take long steps, we also'turn the pelvis partly roundK presenting the side obliquely forward; and in this manner increase the anterior projection of the front leg. Although the action of the gastrocnemius, &c. seems necessary to walking with long steps; we can walk without their operation. This is proved incontestably by the act of walking on the heel: when the gastrocnemii and the flex- ors are so far from acting, that they are in a state of exten- sion. In this operation, the principal effort seems to be made by the tibialis anticus, and the muscles on the front" of the leg; and the extensor muscles on the front of the thigh. Vol. I. 2 R 314 Motions necessary in Jumping. Notwithstanding these facts, the action of the gastroc- nemius and soleus is essential whenever we raise the heel from the ground, while the weight of the body presses on the front part of the foot; and it then acts with a force which equals, if it does not exceed, the weight of the body. Jumping, at the first view of it, appears an extraordina- ry operation; but if a man who lies on the ground, with his feet against a wall, makes a muscular exertion, such as is necessary for jumping, the nature of the operation is very intelligible. It is a sudden extension of the feet and knees, and sometimes of the trunk of the body. The stroke is made against the wall; but as that does not yield, the whole motion is imprest upon the body; which is projected from the wall horizontally, in the same way that in jump- ing, it is projected from the ground vertically. SYSTEM OF ANATOMY. PART IV. OF THE BRAIN AND SPINAL MARROW: OF THE EYE AND THE EAR. CHAPTER I. OF THE BRAIN. 1 HE whole of the soft mass, which fills the cavity of the cranium, is called the brain. This mass is covered by three membranes: two of which were called meninges, or maters, by the ancient anatomists; who believed that all the other membranes of the body originated from them. These membranes are denominated the Dura Mater, Tunica Arachnoidea, and Pia Mater, SECTION I. Of the Membranes of the Brain, and Sinuses of the Dura Mater. The Dura Mater incloses the brain and all its appen- dages, and lines the different parts of the cranium. It con- sists of one membrane of a very dense texture, which in several places is composed of two or more laminae. It is the thickest and strongest membrane of the body, and is composed of tendinous fibres, which have a shining ap- pearance, particularly on its inner surface. In many parts these fibres run in a variety of directions, and decussate each other at different angles. 316 Dura Mater. The dura mater adheres every where to the surface of the cranium, in the same manner as the periosteum ad- heres to the bones iri the other parts of the body; but it is more firmly connected at the sutures and foramina than elsewhere; and so much-more firmly in children than in adults, that in separating it from the cranium, it is apt to bring along with it some of the fibres of the bone to which it is attached. In the adult, the separation of the bone from the membrane is less difficult, in consequence of many of the fibres being obliterated; although in old age the adhe- sion is sometimes very strong. The inner surface of the dura mater, which is remarka- bly smooth, is in close contact with the brain, but adheres only where the veins go into the sinuses; and is lubricated by a fluid discharged through its vessels, which guards the brain from danger, according as it may be affected by the different states of respiration. The dura mater serves as a defence to the brain, and supplies the place of a periosteum to the inside of the cra- nium; giving nourishment to them, as is evident from the numerous drops of blood which appear after removing a portion of it. The proper bloodvessels of the dura mater are not very numerous. Its arteries are derived partly from the exter- nal carotids, and partly from the internal carotids, and the vertebral arteries. Corresponding veins accompany these arteries; but the dura mater forms also reservoirs, that contain the venous blood, which is brought from the sub- stance of the brain. These are called sinuses, and are very different from common veins. Nerves have not been distinctly traced into the dura mater; and there have been disputes respecting its sensi- bility; but there is reason to believe that in a sound state it has very little sensibilitv. Tunica Arachnoidea—Pia Mater, 317 The Tunica Arachnoidea is an exceedingly thin, tender, and transparent, membrane, in which no vessels have been hitherto observed. It is spread uniformly over the surface of the brain, in- closing all its convolutions, without insinuating itself be- tween any of them. Atthe upper part of the brain, it adheres so closely to the subjacent coat by fine cellular substance, that it can scarcely be separated from it; but in different parts of the base of the brain, particularly about the tuber annulare and medulla oblongata, it is merely in contact withthe^mem- brane under it, and may readily be raised from it by the assistance of the blow-pipe. The Pia Mater, named from its tenderness, is some- what of the nature of the former covering, but is extremely yascular. It covers the brain in general, enters double between all its convolutions, and lines the different cavities called ventricles. It serves to conduct and support the vessels of the brain, and allows them to divide into sueh minute parts, as to prevent the blood from entering the tender substance of this viscus with too great force. The arteries of the pia mater are the same with those of the brain, and are derived from the internal carotids and vertebrals. The veins differ in no respect from those of the other viscera, excepting in this, that they do not accompany the arteries. From the dura mater certain membranous processes go off, forming incomplete partitions, which partially divide the cavity of the cranium; and in the same partial manner separate the parts of the brain from each other: thus pre- 318 Falx. venting them from pressing upon each other, and keeping them steady. They are formed of the internal lamen or layer of the dura mater, like a plait, and therefore each of them con- sists of a double membrane. The most conspicuous of these is denominated the falx, which extends from the anterior to the posterior part of the cranium, and divides the upper part of the brain into two hemispheres; but it is not sufficiently deep to divide the whole of the brain; for between the under edge of it, and the base of the cranium, there is a large space occu- pied by a portion of the brain, which is undivided; and, therefore, common to both hemispheres. The falx begins at the middle of the sphenoid bone, and, continuing its origin from the crista galli of the ethmoid bone, runs along the upper and middle part of the head; adhering first to the frontal, then to the joining of the parietal, and afterwards to the middle of the occipital, bone. In its passage it becomes gradually broader; and ter- minates behind, in the middle of the tentorium. It runs from before backwards in a straight direction, and has some resemblance in shape to a sickle or sithe, placed with its edge downwards; from which circumstance it has obtained the name of falx. After extending backwards as far as the center of the crucial ridge, on the internal surface of the occipital bone, it extends to each side, and forms a horizontal partition, which partially divides the lower part of the cavity from the upper; but it does not extend so far forward as to se- parate completely the mass which is under it, or the cere- bellumj from the upper part of the brain, or cerebrum. Tentorium—Sinuses. 319 This horizontal membrane is called the tentorium, and also the transverse septum: it is connected behind to the inner transverse ridges and grooves of the occipital bone, and, at the fore and outer edges, to the ridges and great angles of the temporal bone3, and terminates at the poste- rior clinoid process of the sphenoid bone. Between the inner edge of the tentorium and the poste- rior clinoid process of the sphenoid bone, there is a large notch, or foramen ovale, where the brain and cerebellum are united, or where the tuber annulare is chiefly situated. The tentorium keeps the falx tense, and forms a floor or vault over the cerebellum, which prevents the brain from pressing upon it. The falx minor, or septum cerebelli, is placed be- tween the lobes of the cerebellum. It descends from the under and back part of the falx in the middle of the ten- torium, adheres to the inferior longitudinal spine of theos occipitis, and terminates insensibly at the edge of the fo- ramen magnum of that bone. Besides the processes of the dura mater already descri- bed, there are four of inferior consideration: two of which are situated at the sides of the sella turcica, and two at the edges of the foramina lacera. As these partitions arise like plaits from the internal surface of the dura mater, there must necessarily be a ca- vity, larger or smaller, between the external layer of the dura mater, which lines the internal surface of the crani- um, and the basis of the partition which rises from it: this cavity must continue along the whole basis of the parti- tion; and a section of it will be triangular. This cavity is of considerable size at the upper edge of the falx, where it rises from the dura mater; and also where it forms the tentorium; and at the posterior edges of the tentorium, where it adheres to the occipital bone. 320 Sinuses. The cavity at the upper edge of the falx is called the longitudinal sinus; that at the posterior edge of the tento- rium forms two cavities, called the transverse sinuses; and that which is at the junction of the falx and tentorium has the name of the torcular or press of Herophilus. The veins of the brain open into these sinuses; and the blood flows through them into the internal jugular veins. They differ from veins principally in this: that they are triangular, and, by the tension of the dura mater, are pro- tected from pressure. The principal sinuses are, 1. The longitudinal sinus, which begins at the crista galli, and, running along the upper edge of the falx until it arises at the tentorium, increases gradually in size, and terminates in 2. The two lateral sinuses which run in depressions of the occipital and temporal bones, until they terminate in the internal jugular veins at the foramen lacerum. 3. The torcular Herophili which receives a large vein from the interior of the brain, and is situated at the junc- tion of the falx and tentorium, opening into the longitudi- nal sinus, where it divides into the lateral sinuses. These are the largest sinuses of the dura mater; but, in addition to these, there are several small sinuses: as, 4. The inferior longitudinal sinus, which is situated at the under edge of the falx, and receives blood from the central parts of the cerebrum: it terminates in the torcular Herophili, near the beginning. The other small sinuses are situated under the brain, viz. 5. The circular sinus of Ridley,* which frequently sur- * An English anatomist, who published near the end of the lrth ccnturv.' Cerebrum. 321 rounds the pituitary gland, and carries the blood from the contiguous parts to the 6. Cavernous sinuses, which are placed at the sides of the sella turcica, surrounding the carotid arteries and the sixth pair of nerves, and receive blood from the circular sinuses and several contiguous parts, and discharge it into the 7. Inferior petrous sinuses, which are placed at the bases of the partes petrosae, and discharge this blood into the ends of the lateral sinuses. To these, should be added 8. The superior petrous sinuses, which are situated on the upper edges of the petrous bones. They communicate both with the lateral and the cavernous sinuses, and re- ceive some small veins from the adjacent parts. There are also several small sinuses near the great oc- cipital foramen, which communicate with the lateral sinus- es, and also with the vertebral veins. They are called occipital sinuses. The brain, or the whole of the soft substance contained within these membranes, is composed of four portions, viz. cerebrum, cerebellum, tuber annulare or pons Varolii^ and medulla oblongata. SECTION II. Of the Cerebrum. The cerebrum completely fills the upper part of the cavi- ty of the cranium. It has some resemblance to the half of an egg which has been divided horizontally; and is composed of two equal parts, which are separated vertically from each other by the falx. This vertical separation does not extend through the center of the cerebrum, although it divides it completely before and behind. A portion of the central part of the cerebrum, which is situated deeper than the under edge of the falx, is not divided. Vol. I. 2 S 322 Cerebrum. The upper surface of the two hemispheres is convex. The under surface is rather irregular: it is divided in each hemisphere into three lobes: the anterior, the middle, and the posterior. The anterior lobes of the brain are situated on the front part of the base of the cranium, principally on the orbitar processes of the os frontis. The middle lobes are lodged in the fossae formed by the temporal and sphenoid bones. The posterior lobes rest chiefly upon the tentorium, over the cerebellum. Between the anterior and middle lobes is a deep furrow, corresponding to the base of the cranium on which they rest, which is called the fossa Sylvii. The surface of the brain resembles that of a mass of small intestines, or of a convoluted cylindrical tube: it is therefore said to be convoluted. The fissures between these convolutions do not extend very deep into the sub- stance of the brain. The whole surface of the brain, thus convoluted, is co- vered by pia mater; which is connected to every part of the surface by an infinite number of small vessels and pro- cesses, that appear when this membrane is peeled off from the surface- of the brain. The mass of brain consists of two substances of different colours: one of which is, for the most part, exterior to the other. The exterior substance is of a light brown colour, and is therefore called cineritious, or cortical, from its situation. The internal substance is white, and is denominated tht medullary. The proportion of this medullary part is much greater than that of the cortical. The cortical, however, surround? Cerebrum. 323 it, so as to form the whole of the surface of the cerebrum, that can be strictly said to be exterior. The colour of the cortical part appears to be derived from blood; as its intensity seems regularly proportioned to the quantity of blood in the head. In subjects who have been plethoric, or have had a determination of blood to the head, it is uniformly high-coloured; in pallid and ex- hausted subjects it is of a brighter colour. The medullary matter is uniformly white; but small red points appear upon its surface, when cut, which are the sections of vessels thatcarry red blood; and these points are larger and more numerous in plethoric, than in ex- hausted subjects. It is rather firmer than the cortical sub- stance. These two substances are most intimately connected in the cerebrum, and indeed seem to be a continuation of each other. In some parts they are blended together; and in other places, there are portions of cortical matter within the medullary. The division of the cerebrum into two hemispheres ex- tends to a considerable depth from above, and also to a considerable distance internally, from its anterior and pos- terior extremities; of course, the part which is undivided is in the center. The cortical part covers alsothe surfaces, which are inthe great fissure that forms the two hemispheres, and is occu- pied in a great degree by the falciform process of the dura mater. Towards the bottom of this fissure, and below the falx, these surfaces being opposed to each other, and in contact, are slightly united by adhesion of the membranes that cover them. The central part which is not divided, and which must appear at the bottom of the fissure, when the two hemis- 324 Lateral Ventricles. pheresare separated from each other, is medullary; being evidently a union of the medullary matter of each hemis- phere. This undivided medullary part is equal to about one half of the length of the hemispheres; the fissures at each extremity extending inwards about one-fourth of their length. On each side of it, a fissure, equal to it in length, extends horizontally into the medullary matter in each hemisphere, about half an inch; and the whole of this un- connected surface, the middle of which is directly at the bottom of the great fissure, is termed corpus callosum. When the hemispheres are cut away to the level of this surface, and the corpus callosum is examined, two raised lines appear in the middle, which extend from one end of it to the other; and between them is a small groove of the same length. This groove is called the raphe, or su- ture of the corpus callosum. From the raised lines or bands on each side of the raphe, small lines less eleva- ted pass across the corpus callosum, and are lost in the medullary matter. The hemispheres being thus cut off at the level of the corpus callosum, on the cut surface is to be seen the interior mass of medullary matter, with the corti- cal part exterior, its edge exhibiting the convoluted sur- face of the brain, and the pia mater following the convolu- tions. The medullary surface thus exhibited, with the,corpus callosum in the center, is denominated the centrum ovale. In the brain there are four cavities called ventricles: three of these are formed in the substance of the cere- brum; the fourth is situated between the cerebellum, the pons Varolii, and medulla oblongata. The two largest are called the lateral ventricles, from their situations; the others are named, from the order in which they occur, the third and the fourth ventricles. Lateral Ventricles. 325 The lateral ventricles are cavities of an extremely irre- gular figure: they are situated in each hemisphere a little below the level of the corpus callosum; and, with the ex- ception of the partition which separates them, are directly under it. They commence anteriorly, nearly on a line with the termination of the fissure that separates the two hemi- spheres anteriorly; and continue backwards almost as far as the commencement of the fissure that separates them posteriorly: when they have attained this length posteri- orly, they form a considerable curve, first outwards, then downwards, and afterwards forwards; so that they termi- nate almost as far forwards as they commenced; but much deeper. At the posterior part of their curve, when they incline outwards, previous to their turn downwards, a process or continuation of the cavity extends backwards, almost as far as the cerebrum itself. These elongations are called the posterior cornua or sinuses, or the digital cavities. Each ventricle may therefore be divided into three parts, viz. The portion under the corpus callosum; the portion which continues outwards and downwards, and terminates below it; and the posterior portion. It has been compared to a ram's horn, by some who have contemplated particularly the upper and lower por- tions of the cavity; and by others, who have had the whole extent in view, it has been called tricornis. The bottom, or lower surface of these cavities, is varied in almost every part of its extent. The front part of the bottom of each ventricle is a broad and convex eminence, which becomes narrower as it proceeds backwards; so that it resembles a portion of a pear. It inclines outwards as well as backwards, so that the narrow posterior extremi- 326 Fornix. ties of the two bodies are further from each other than the anterior broad extremities. The colour of these bodies is cineritious externally; but they are striated with medullary matter within, and there- fore are called corpora striata. Between their posterior extremities are two other emi« nences, which incline to the oval form, and have a white or medullary colour; although their substance, when cut into, is slightly striated: they are called the thalami nervo- rum opticorum. These bodies are very near each other; and, being convex in form, are in contact at the center: they adhere slightly to each other; and this adhesion is called the soft commissure. The corpora striata, and the thalami nervorum optico- rum, join each at the exterior sides of the thalami: where they are in contact, there is the appearance of a narrow me- dullary band, which continues during the whole extent of their connexion: it has been called, by some, tenia semicir- cularis, from its form; by others, centrum geminum semi- circulare. These surfaces constitute the bottom or floor of the first portion of the ventricles, which is under the corpus callo- sum: upon this floor is laid a thin lamen of the medullary matter, of a triangular form, called the fornix, which co- vers the thalami nervorum opticorum, and is attached to them by a membrane; so that when the ventricles are opened, the bottom appears to consist of the corpora striata, and the fornix. The upper surface or roof of the ventricles is concave; from the middle of it, immediately under the raphe of the corpus callosum, there proceeds downwards a partition of medullary matter, which separates the two ventricles from each other. This is called septum lucidum, from its being Plexus Choroides. 327 nearly transparent: below, it adheres to the fornix, posteri- orly; and anteriorly, it is continued into the medullary mat- ter, between the corpora striata. This septum lucidum is formed of two laminae or plates, which are separated from each other in the anterior portion of the septum, and thus form a small cavity, which has no communication with the other cavities of the bfain. The fornix is not perfectly flat; but, accommodating to the surface of the thalami nervorum opticorum, its under surface is rather concave, and its upper surface convex. The anterior angle passes down between the most anterior parts of the thalami nerv: optic: and is divided into two small portions called its crura, which can be traced some distance in that part of the brain. The body of the fornix is attached to the surfaces of the thalami nervorum opticorum; on which it rests, by a very vascular membrane, which is spread over the thalami, and called tela choroidea, and velum interpositum. At the edges of the fornix, there are many bloodvessels in this mem- brane, arranged close to each other, which are called the plexus choroides. The posterior side or edge of the triangular fornix ter- minates in the corpus callosum, or the medullary matter which is above it at that place; but the under surface is attached throughout to the parts on which it lies, by the aforesaid membrane. The two posterior angles of the fornix form what are called the crura; and they terminate in the following way. The surfaces of the inferior portions of the lateral ven- tricles are not uniformly concave; but at the bottom of each, there is a prominent body, which begins where this portion of the cavity winds outward and forward, and con- tinues its whole extent. This prominence has a curved 328 Third Ventricle. form, and is marked by transverse indentations, towards its extremity: hence it has been termed the hippocampus, or cornu ammonis. A similar prominence, but smaller, and without the transverse indentations, is to be found in the posterior portion of the ventricle: this has also been called hippocam- pus; but the terms minor and major are applied to distin- guish them. The posterior angles of the fornix terminate in the large hippocampi; and the margin, or thin edge of the two ante- rior sides of the fornix, is continued to form an edge to the hippocampus; and is called the tania hippocampi, or cor- pus flmbriatum. The word fornix was the ancient name of a vault or arch; and from its supposed resemblance to an arch, this part has been called by that name. When the fornix is raised up, (which must be done by dividing it at the anterior angle, and detaching it from the thalami nervorum opticorum, by dissecting the membrane which connects them*, called velum interpositum, or tela choroidea^) the thalami are brought fairly into view; and appear like oval bodies placed parallel to each other. They adhere slightly at their upper surfaces, and, when separa- ted, a fissure appears between them; which is the third ventricle. At the upper and front part of this third ventricle, near its commencement, before the anterior crura of the fornix, and very near them, is a white chord, like a nerve, which passes across the ventricle, and can be traced to some distance on each side into the medullary matter of the brain. * It is to be observed that the parts of the brain are here described in the order in which they occur, when successive layers of the organ are cut away: beginning with the upper surface. The fornix of course must be raised up, after it is brought into view. Commissures of the Brain. 329 This chord is called the anterior commissure of the brain. The thalami nerv: optic: being of an oval form, and touching each other in the middle, there must be a vacu- ity between them at their extremities. This vacuity is be- hind the anterior crura of the fornix, and has been called Vulva, Iter ad Infundibulum, and Iter ad Tertium Ventri- culum. It leads of course into the third ventricle; but a pas- sage continues from it downwards and rather forwards, to the infundibulum; which is a process somewhat resem- bling a funnel that is composed principally of cineritious substance, and passes from the lower and front part of the third ventricle, towards the sella turcica; in which is situ- ated the small body called the Pituitary Gland. The infundibulum is hollow at its commencement, and solid at its extremity near the gland. The adhesion of the thalami nerv: optic: to each other, at the upper part of the third ventricle, has been denomi- nated the Commissura Mollis. The recession from each other at their posterior extremities, in consequence of their oval figure, forms another opening into the third ventricle when the fornix and tela choroidea is raised, which is closed when they are in their natural situations upon it. In the back part of the third ventricle is another medul- lary chord called the posterior Commissure, which appears much like the anterior commissure; but does not extend into the substance of the brain in the same way. Under this chord, or posterior commissure, is a passage which This mode of examining the brain was employed by Vesalius, and has been adopted by a majority of anatomists; but Willis and Vieussens had each a different method Vol. I. 2 T 330 Tubercula ^uadrigemina—Pineal Gland. leads to the fourth ventricle, called Iter ad §>uartum Ven- triculum, or Aqueduct of Sylvius. Behind the third ventricle, and terminating it poste- riorly, are four convex bodies, called Tubercula §>uadri- gemina, or Nates and Testes: the nates are uppermost and most convex; the testes are immediately below, and some- what oval transversely. The nates and testes are situated so far backwards that they are near the anterior part of the upper surface of the cerebellum, and the anterior edge of the middle of the ten- torium. The posterior part of the fornix is directly over them, but it unites with the medullary matter of the cere- brum above it; there would therefore be a passage into the lateral ventricles from behind, between the back of the fornix which is above, and the nates and testes which are below; but a fine membrane passes in from behind, and attaches the lower surface of the fornix to all the parts on which it lies; and thus closes the ventricles at this place. The membrane, however, is continued in the ventricles, and is spread over the thalami nerv: optic: The fornix of course lies upon it, and is attached by it to the surfaces of the thalami; in this membrane, immediately over the pos- terior end of the fissure called the third Ventricle, and in contact with the nates, is the Pineal Gland. This body is not so large as a pea, and formed like a pine apple, or the cone of a pine tree. When the fornix is raised, by dissecting the membrane, it may be elevated with the membrane and fornix. The nature of this body is not understood: it resembles a small gland in its appearance; but it is very soft; and particles of sandlike matter are often found in it. There is a small chord on each edge of the third ven- tricle, which appears to proceed from the pineal gland and Communication of the Ventricles— Vena Galeni. 331 continues on the edge of the ventricle to the anterior crura of the fornix, which it unites to. These chords join each other under the pineal gland: they are called the pedun- xula, or footstalks of the pineal gland. The membrane connected with the pineal gland, is the tela choroidea, or velum interpositum, in which the plexus choroides is placed, at the edges of the fornix. This mem- brane is extended, somewhat thinner and less vascular, so as to line the surfaces of the ventricles. The plexus cho- roides appears to begin at the end of each of the inferior portions of the ventricles, where the pia mater penetrates from the basis of the brain: it proceeds into the upper por- tions of the ventricles, and, continuing along the edge of the fornix, passes under that body at its inferior angle, and meets the plexus of the opposite side. Between this meeting of the plexus, and the crura of the fornix, is a va- cuity of an oval figure, which forms a communication be- tween the ventricles of the brain. Under this vacuity or foramen, the thalami nerv: optic: recede from each other, and form the anterior passage into the third ventricle de- scribed at page 329, so that at this place the three ventri- cles communicate with each other. From the plexus choroides of each side, where it has passed under the fornix at the anterior angle, a large vein is turned backwards, so as to run nearly over the fissure of the third ventricle towards the pineal gland. Several veins from the surface of the ventricle join this vein near its commencement; thus formed it passes along with the corresponding vein from the opposite side, sometimes in contact, and sometimes separated to a small distance; near the pineal gland, these veins unite into one trunk, the great internal vein of the brain, called the Vena Galeni; which terminates soon after in the torcular Herophili. 332 Arbor Vita. SECTION III. Of the Cerebellum. THE cerebellum is situated in the lower and posterior part of the cavity of the cranium, in contact with that por- tion of the os occipitis which is below the grooves for the lateral sinuses. It is of course much less than the brain. It is covered above by the tentorium, and is divided be- low into two lobes, by the falx minor. The surface of the cerebellum differs in some respects from that of the cerebrum. Instead of the convolutions there are small superficial depressions, which are nearly horizontal, tending to divide the cerebellum into strata. The pia mater extends into these depressions; and the tunica arachnoidea passes over them, as in the cerebrum. The exterior part of the cerebellum is composed of cine- ritious or cortical, and the internal of medullary, matter, as is the case with the cerebrum: but the proportions of these substances in the cerebellum, are the reverse of what they are in the cerebrum. If sections are made in the cerebellum, the medullar)- matter is so arranged that it appears like the stem or trunk of a plant, with ramifications extending from it. This ap- pearance has been called the Arbor Vita. On the basis of the brain is a part called Tuber Annu- lare, or Pons Varolii, which is formed by processes from the cerebrum and cerebellum; and is in contact with the anterior and inferior portion of the cerebellum in the mid- dle. From this part the medulla oblongata proceeds down- wards and backwards, under the cerebellum: and between the cerebellum, the medulla oblongata, and the pons Varo- lii, is the vacuity, called tht fourth Ventricle of the brain. Valve of the Brain. 33S When the brain is in its natural situation this cavity is below and behind the nates and testes; and from the cere- bellum there passes up to the testes a lamen of medullary matter, which closes it above. This lamen is called the Valve of Vieussens, or the Valve of the Brain. Below, the ventricle is closed by a membrane which connects the me- dulla oblongata to the cerebellum. There is a passage into this cavity from the third ven- tricle, which passes under the posterior commissure and the nates and testes, and enters it below the testes. SECTION IV. Of the Basis of the Brain and the Nerves which proceed from it. WHEN the brain is detached from the basis of the cra- nium, and inverted; which can be readily done, if the nerves that proceed from it are divided, as it is inverted. The tunica arachnoidea appears more conspicuous on the basis than it is on the upper part; and the pia mater is disposed round the convolutions in the same manner that it is above; but the nerv«s and vessels connected with the sur- face of the brain are so much involved with these mem- branes, that considerable dissection is required to expose them properly. The anterior and middle lobes of the brain are very conspicuous on the inverted surface. The anterior lobes appear separated from each other by the extension of the great fissure which forms the two hemispheres. The mid- dle lobes appear at some distance from each other in the center; and the cerebellum forms the posterior and most prominent part of the surface. 334 Olfactory and Optic Nerves—Corpora Albicantia. When the brain has been carefully detached from the cranium and the nerves adhering to it are preserved, the olfactory or first pair,* of nerves, appear on the anterior lobes, running nearly parallel to each other at a small distance from the great fissure. They are flat and thin, and soft, in their texture: their breadth is rather more than one-sixth of an inch. They pass in three divisions from between the anterior and middle lobes of the cere- brum, which soon unite and run to the cribriform plate of the ethmoid, where they expand into soft bulbous lobes, from which proceed the fibres that perforate the cribri- form plate, and are spread upon the schneiderian mem- brane. Behind the olfactory nerves are the optic. Each of which comes out between the anterior and middle lobes of the cerebrum; and after bending so as to meet its fellow, turns off and passes through the optic foramen in the sphenoidal bone. These nerves can be traced in the brain to the tha- lami nerv: optic:. In the angle formed by the optic nerves posteriorly, is a mass of softish cineritious matter; and also the infundi- bulum which passes to the sella turcica. In this soft cineritious matter are two round white bo- dies that resemble peas; which are called the Corpora Albi- cantia of Willis, or the Eminentia Mammillares. Behind these bodies are two large medullary processes, called the Crura Cerebri, which are best seen if some of the cortical part of the adjoining middle lobes is dissected away. They come from the medulla of the opposite sides of the brain, and gradually approach each other until they arrive at the tuber annulare, or pons Varolii. * The nerves are numbered from before. Pons Varolii—Medulla Oblongata. 335 The Pons Variolii is a mass of considerable size, which has a medullary appearance externally, but is striated with- in: it is formed by the union of the two above mentioned crura cerebri, and of two similar processes derived from the cerebellum, called also its Crura. It lies over a part of the body of the sphenoid bone, and of the cuneiform process of the occipital bone, and under a portion of the middle lobes of the cerebrum and of the cerebellum. There is a longitu- dinal depression on its surface, made by the vertebral arte- ry; and there are also many transverse streaks on it. The crura of the cerebellum, which run into this sub- stance, are evidently continued from the arbor vitae, or me- dulla of the cerebellum. The anterior edge of the cerebellum, part of which is in contact with the pons varolii, is remarkably prominent on each side of it. These prominences are called the Vermes of the cerebellum. The medulla oblongata is continued backwards from the posterior side of the tuber; and somewhat resembles a truncated cone inverted. It lies on the cuneiform process of the occipital bone, and extends to the foramen magnum. It is indented length- ways, both anteriorly and posteriorly, by fissures which are very evident: it is composed of medullary matter ex- ternally, and cineritious matter within. On each side of the anterior fissure, which is in view when the brain is inverted, are two oblong convex bodies: those which are next to the fissure are called the Corpora Pyramidalia, and are the longest: the two exterior are called Corpora Olivaria, and are not so long. The third pair of nerves come from between the crura of the cerebrum, and pass forward, diverging from 336 Fourth, Fifth and Sixth pair of Nerves. each other. They proceed by the cavernous sinus, and, after penetrating the dura mater, go out of the cranium at the foramen lacerum. The fourth pair, the smallest nerves of the brain, resem- ble sewing thread in their size and appearance. They come out between the cerebellum and pons Varolii, but can be traced backwards as far as the testes. They proceed for- wards by the sides of the pons Varolii, and, after penetra- ting the dura mater near the posterior clinoid apophysis, pass through the foramen lacerum to the trqchlearis mus- cles of the eye. The fifth pair, the largest of the brain, arise from the crura of the cerebellum, where they unite with the pons Varolii: they pass forwards and downwards, and penetrate the dura mater near the point of the petrous portion of the temporal bone. This nerve appears like a bundle of fibres; and, under the dura mater, forms a plexus; from which its three great branches proceed to their destination. The sixth pair arise from the medulla oblongata, where it joins the pons Varolii. It is often composed of two chords on each side, one of which is very small; they pass under the pons Varolii, and through the cavernous sinus, with the carotid artery: after emerging from this sinus they proceed through the foramen lacerum to the abductor muscles of the eye. In this course a small twig passes from it, which accompanies the carotid artery through the ca- nal in the petrous portion of the temporal bone, and, with a twig from the fifth pair, is the origin of the intercostal nerve. The seventh pair appears at the side of the medulla ob- longata, near the pons Varolii. It is composed on each Eighth Pair of Nerves. 337 side of two chords, called Portio Dura and Portio Mollis, and of one or more small fibres between them, called Por- tio Media. The portio mollis can be traced to the fourth ventricle. The portio dura seems to arise from the place of union of the pons Varolii with the medulla oblongata and the crura cerebelli. The portio media appears to origi- nate in the same neighbourhood, and may be considered as an appurtenance of the portio dura. They all proceed to the meatus auditorius internus, as it has been called, in the temporal bone. The eighth pair of nerves arise from the corpora oliva- ria on the side of the medulla oblongata. They are com- posed on each side of one chord called the Glosso-Pharyn- geal, and of a considerable number of small filaments, which unite and form another chord called the Par Va- gum. With these nerves is associated a third chord called the Spinal Accessory Nerve of Willis, which passes up the spinal cavity, being composed of twigs from the posterior and anterior portions of almost all the cervical nerves. The par vagum, with this nerve and the glosso-pharyn- geal, proceeds from its origin to the foramen lacerum formed by the occipital and temporal bones; where they all pass out of the cranium; separated from each other and from the internal jugular vein, by small processes of the dura mater. Their destination is extremely different. The glosso- pharyngeal is spent upon the tongue and pharynx; the par vagum upon the contents of the thorax and abdomen, &c; while the accessory branch, which seems to have no connexion with them, perforates the sterno mastoid mus- cle, and is distributed among the muscles of the shoulder. Vol. I. 2 U 338 Ninth Pair of Nerves, fcrY. The ninth pair arise from the corpora pyramidalia by many filaments that are united on each side into three or four fasciculi, which perforate the dura mater separately, and then unite to pass out of the anterior condyloid fora- men of the occipital bone: this pair is spent upon the mus- cles of the tongue. Within the last three or four years, there have been many allusions in the public papers, to the discoveries of Dr. Gall, formerly of Vienna, respecting the brain. For information concerning these discoveries, the reader is referred to a very learned and judicious Memoir, pre- sented to the class of Mathematical and Physical Sciences of the National Institute of France, by Messrs* Tenon, Portal, Sabatier, Pinel, and Cuvier. A translation of this report has been published in the fifth volume of the Edinburgh Medical and Surgical Journal for 1809. 339 CHAPTER II. OF THE SPINAL MARROW. 1 HE medulla oblongata is continued from the cavity of the cranium, through the great foramen of the occipital bone, into the great canal of the spine; when it takes the name of Medulla Spinalis, or Spinal Marrow. The dura mater passes with it through the great fora- men, and incloses the whole of it. At the commencement of the spinal canal this membrane is attached to the sur- rounding bones, viz.: to the margin of the great occipital foramen, and to the atlas; but below this it is loosely con- nected by a membrane which sometimes appears to contain a little adeps. The tunica arachnoidea and the pia mater also invest the medulla spinalis. The arachnoidea appears unconnected with the dura mater; and it can easily be re- moved from the pia mater. The pia mater adheres rather firmly to the substance it incloses. The spinal marrow consists of medullary matter exter- nally, and cineritious or cortical matter internally. The fissures, which are observable, anteriorly, and pos- teriorly, in the medulla oblongata, are continued down the spinal marrow; dividing it partially into two lateral por- tions: these fissures penetrate to a considerable depth. Each of the lateral portions is marked on its external sur- face, by a more superficial fissure, which partially divides it into an anterior and posterior part; so that a transverse section of the spine has a cruciform appearance. The nerves go off in fasciculi from the anterior and pos- terior surfaces of each lateral portion of the spinal marrow; so that each nerve is formed of two fasciculi: one from 340 Ligamentum Denticulatum. Cauda Equina. before, and the other from behind. The fasciculi are of different sizes in different parts of the spine. The lower- most of the neck are large and broad; those of the back are slender; and those of the loins, and upper part of the sacrum are very large. The uppermost of the fasciculi of the spine proceed, almost at right angles with the medulla spinalis, to the fo- ramina through which they pass; those which are lower pass off in a direction obliquely downwards; and the low- ermost are almost perpendicular. Betv/een the anterior and posterior fasciculi, a fine ligamentous chord passes; which is attached above to the dura mater, as it passes through the foramen magnum, and continues to the os coccygis. It passes between the tunica arachnoidea and pia mater, attached to the pia mater by cellular membrane. It sends off a small process in a lateral direction, to be at- tached to the dura mater in the interstice between the places where the fasciculi pass through the dura mater, and nearly in the middle between the upper and lower fasciculi. The spinal marrow terminates in a point near the up- permost lumbar vertebrae. The ligamenta denticulata of the opposite sides join each other at this point, and form a small chord, which, continuing downwards, is inserted into the os coccygis. These ligaments may support, and keep fixed, the me- dulla and the nerves, as they originate from it. As the spinal marrow terminates at the upper lumbar vertebrae, the lumbar and sacral nerves go off above: they pass down like a bunch of straight twigs, and are called Cauda Equina; from a fancied resemblance to the tail of the horse. The sheath, formed by the dura mater for the Arteries of the Spinal Marrow. 341 spinal marrow, continues of its original size, and incloses them in one cavity. The posterior and anterior fasciculi pass out separately from the dura mater: after they are out, the posterior fas- ciculus forms a ganglion; from which one nerve passes that joins the anterior fasciculus, and thus forms the spi- nal nerves. When the nerves go off, either from the spinal canal, or the cavity of the cranium, the external lamen of the dura mater where they pass out, attaches itself to the bone or the periosteum; while an internal lamen, together with the pia mater, and perhaps the tunica arachnoidea, is continu- ed with the nerve. This process from the dura mater becomes so much changed that it has been considered as cellular membrane. The pia mater and tunica arachnoidea seem also to invest, not only the nerve in general, but the fibres of which it is composed. On this account, probably, the nerves are larger after passing through the dura mater, than they are when they leave the brain and spinal marrow. The arteries of the spinal marrow proceed from the head, and, with several additions, continue downwards to the lumbar vertebrae. There is generally one artery on the front surface of the medulla, which is formed by the union of two branches, that arise from the vertebral arteries within the cranium. This artery proceeds downwards and communicates with those of the neck, and with the intercostal arteries, by the intervertebral foramina, so that it preserves its size. It terminates with the spinal marrow; and the cauda equina below it is supplied by branches from the internal iliac, which enter through the foramina of the sacrum. 342 Veins of the Spinal Mdrrow. There are generally two arteries on the posterior sur- face of the medulla spinalis, which also pass out from the cranium; arising from the vertebral arteries, or inferior arteries of the cerebellum: they have a serpentine arrange- ment; and communicate with each other, and with the ramifications of the anterior spinal artery. All of these arteries are dispersed upon the spinal mar- row and its membrane, and the parts immediately conti- guous. The veins correspond with the ramifications of the arte- ries; but they are collected into two larger branches called the Sinus Venosi; which are situated exterior to the dura mater, on the front and lateral sides of the spinal canal. They extend the whole length of the canal, and, entering the great occipital foramen, communicate with the lateral and occipital sinuses. 343 CHAPTER III. OF THE EYE. It will be very proper to read the description of the orbit of the eye in page 54 of this volume, as an introduction to the following description of the organ. In addition to that account of the bones, it is to be ob- served, that processes of the dura mater pass through the foramina optica and lacera, which line the cavity of the orbit and unite with the periosteum at the margin of it. THE eye is an optical instrument of a spherical form, which lies in the orbit, in a bed of adipose membrane, more or less filled with adeps for the convenience of its motions. Connected with the ball of the eye, are several auxiliary parts, which are calculated for its motion and protection, as well as accommodation in other respects. SECTION I. Of the Parts auxiliary to the Eye. ABOVE the upper margin of the orbit, on the promi- nences of the os frontis, called the superciliary ridges, the adipose membrane is commonly more full than it is in the other contiguous places; and the skin which covers it is thereby rendered prominent. The supercilia or eyebrows grow out of this prominent skin. The hairs which compose them are placed obliquely, with their roots towards the nose. Their principal use seems to be to defend the eye from sweat, and other matters which roll down the forehead. They are moved by the corrugator muscle, and thus express certain passions; and 344 The Eyelids. they are also moved by the occipito-frontalis and orbicu- laris palpebrarum. The Eyelids or Palpebra, Are formed by a slit or orifice in the skin: immediately under the skin, surrounding this orifice, is a portion of the orbicularis muscle, (see page 171), in the upper eyelid: under this portion of the muscle, there is a plate of cartil- age; and under the cartilage, a portion of tunica conjunc- tiva, or membrane that covers the front part of the ball of the eye and lines the eyelids. The upper eyelid, therefore, is composed of the skin, of some fibres of the orbicularis muscle, of the cartilagin- ous plate, and of the tunica conjunctiva. The under eye- lid is formed in the same way, with the exception of the cartilage, which, in it, is confined to the margin. These cartilages form the margin of each eyelid, which is called Tarsus. The upper cartilage is broad in the mid- dle, and narrower at each extremity, and accommodated to the form of the eyeball and eyelid. The under cartilage is a narrow flat rim, which does not extend far from the margin of the eyelid. A thin delicate membrane is extend- ed from the upper and lower margins of the orbit to these cartilages, and has been considered as forming ligaments for them. Their edges are formed obliquely, and apply to each other in such way, that when the lids are closed, a groove is formed between them and the eye; by which the tears are conve)red towards the nose. The use of these cartilages is to keep the eyelids pro- perly expanded; and to form margins that apply accurately to each other. These cartilages are covered internally by the tunica conjunctiva. Tunica Conjunctiva. 345 The levator palpebrae muscle, which arises from the bot- tom of the orbit, at the upper part of the foramen opticum, and passes over the superior muscle of the eyeball; is insert- ed by a broad thin tendon or aponeurosis into the cartilage of the upper eyelid; and draws it upwards, within the up- per margin of the orbit, when the eyelids are opened. The tunica conjunctiva, that lines the eyelids, is conti- nued from them, without any interruption of the surface, over the anterior part of the ball of the eye; in the same manner that the reflected membranes are continued from one surface to another. Although this membrane is a continuation of the skin, it is essentially different from it in its structure; being ex- tremely thin, flexible and sensible, and also transparent. It abounds with vessels, which do not carry red blood in their natural state, but receive it largely when they are inflamed or much relaxed. It adheres firmly to the cartilage at the edges of the eyelids; and becomes more loose in its adhe- sion to the lids, as it proceeds backwards. It is so reflected to the ball of the eye, that it covers and adheres to about one third of it anteriorly. Where it first joins the eye, the adhesion is loose; but this adhesion becomes firmer as it advances over the eye; and it cannot be separated from the cornea without maceration, and a slight degree of pu- trefaction. The part immediately connected with the cor- nea is extremely thin and delicate. This membrane closes the orbit of the eye, and com- pletes the cavity which contains the muscles, lachrymal gland, &c. which are by this means precluded from contact with the external air. On the inside of each eyelid, apparently between the tunica conjunctiva and the cartilages, are a number of lines running inwards from the edge of the lid. These Vol. 1. 2 X 346 Glands of Mcibomius—Lachrymal Gland. lines are of various lengths, from one fourth to near hall an inch; the longest are in the middle of the upper lid. Some of them are straight; and others are serpentine: their colour is a yellowish white. There are generally more than thirty in the upper eyelid, and more than twenty in the lower. They are called the glands of Mcibomius. By pressure a sebaceous substance can be forced out of them, in the form of fine threads, from orifices on the edges of the eyelids. They are follicles into which the sebaceous substance is secreted. This substance appears to have a twofold effect: it prevents the tears from running over the eyelid, as any other unctuous matter would do; and it prevents the eyelids from adhering to each other, in con- sequence of their contact during sleep. The eyelashes or cilia are placed near the outer edge of the lower part of the cartilages of each eyelid. They are.always more or less curved; and their convexities are opposed to each other. By this arrangement the eye is defended from small external objects, and from light to a certain degree, without closing the lids completely. It is necessary for the perfection of the eye, that the whole surface covered by the tunica conjunctiva, viz. the anterior part of the eyeball and the internal surfaces of the eyelids, should be kept perfectly flexible and moist; for this purpose the lachrymal fluid is constantly secreted, in varying quantities, by t.he lachrymal gland, formerly called Glandula Innominata. \ This body is situated in a depression, in the upper sur- face of the orbit, near its external margin: it is exterior to, and above, the tunica conjunctiva. It is of an irregular ob- long form, and rather flat; but it has some thickness. The under surface is lobulated, and forms two principal lobes. From the anterior edge of the gland the excretory ducts, Tears—Lachrymal Sac. 347 to the number of six or seven, pass off. They terminate at a short distance from the gland, near the upper edge of the cartilage, and near the external angle of the eye. They do not communicate with each other. These ducts are so small that they are often not to be seen by the naked eye; but there is sometimes a chain of smaller glands, which lie between the gland and the eye- lid, nearly in the direction of the ducts. The fluid secreted by this gland, (viz. the tears) is trans- parent; but it is always salt to the taste. When evapora- ted,'by exposure to the air, some cubic crystals, and a small quantity of mucilaginous matter, remain. Chemistry has. ascertained that these crystals contain muriate of soda and soda uncombined; and that phosphate of lime and phos- phate of soda may be obtained by burning inspissated tears; but the whole of the saline matter does not amount to one hundredth part of the tears in which they are dis- solved. The tears therefore consist of these salts, and of mucus, dissolved in a large proportion of water. The tears are carried from the eye by two small canals, which commence, one on each eyelid, at the internal ex- tremities of the cartilages; opposite to each other. The orifices of these canals, being in the cartilages, are always open, and are called the Puncta Lachrymalia. Each of these canals runs, within the edge of each eye- lid, from the place of its commencement to the lachrymal sac; which is a larger membranous canal situated in the depression formed by the anterior portion of the os unguis and the corresponding portion of the upper maxillary bone; and extending thence along the bony canal, which continues from this depression into the nose, and termi- nates under the inferior spongy bone near its anterior ex- tremity. 348 Caruncula Lachrymalis-^-Valvula Semilunaris. These canals are very small at their commencement at the puncta lachrymalia; but this small portion is very short; it forms an angle with the remainder of the canal, which is considerably larger. The canals gradually approach each other as they pro- ceed towards the lachrymal sac; into which they enter, in contact with each other, but by distinct orifices, behind the tendon of the orbicularis muscle. About one fourth part of the lachrymal sac is situated above the junction of the two eyelids, or the tendon of the orbicularis muscle, and the remainder below. After it de- scends below the orbit of the eye, it sometimes contracts and takes the name of the Lachrymal Duct. The sac and duct have some resemblance to the schnei^ derian membrane in structure; and are defended with a similar mucus. The membrane, of which they are compo- sed, adheres to the periosteum of the bony canal. The tears appear to enter the puncta lachrymalia upon the principle of capillary attraction; and to be propelled forwards by the motion of the eyelids and the pressure of the orbicularis muscle. Between the puncta lachrymalia and the termination of the eyelids, at their junction with each other, is a small angular space, which is occupied by a body called the Ca- runcula Lachrymalis; which is of a reddish colour, with a few small hairs growing out of it: it is supposed to be glandular, and to secrete a sebaceous substance. It has an effect in directing the tears to the puncta lachrymalia. Between the caruncula lachrymalis and the cornea, the tunica conjunctiva, in many persons, forms a plait or fold; which is very obvious when the eye is directed inwards: this has some analogy with the membrana nictitans of cer- tain animals, and has been called the Valvula Semilunaris. The Ball of the Eye. 349 The muscles of the eye, described at page 171, &c. are to be ranked among the auxiliary appurtenances of that organ. The tendons of the four recti muscles, being spread up- on the anterior part of the ball of the eye, constitute a par- tial covering, which has been called Tunica Albuginea. This tendinous expansion does not extend to the edge of the cornea, but stops short of it by several lines.* SECTION II. Of the Ball of the Eye. THE spherical figure of the eyeball depends upon a strong and firm external coat called the Sclerotica; which has an aperture in its anterior part, filled up with a trans- parent substance, denominated Cornea, that closes it per- fectly. The sclerotica is lined by a thin and delicate membrane, the Choroides; which is in close contact with it, but does not extend over the whole internal surface of the eye, and is deficient in the whole of the part occupied by the cornea. On the internal surface of the choroides is spread the pulpy expansion of the optic nerve, called the Retina; the natural consistence of which is not much more firm than mucus. Within these coats is a cavity, that corresponds with the figure of the sclerotica and cornea, but is divided by an incomplete membranous partition that separates the an- terior part of it, which is covered by the cornea, from the remainder. This partition is called Iris; and it has a cir- cular vacuity in the center, directly opposite to the middle of the cornea, which is denominated the Pupil. Rays of * By line, is meant one twelfth of an inch. 350 Tunica Sclerotica. light, which penetrate the transparent cornea, pass through the pupil into the posterior part of the eye. The eye, thus formed, is filled with several transparent substances, called Humors. The greatest part of the cavity, posterior to the iris, is occupied by the vitreous Humor, which is thus denominated, from its apparent resemblance to melted glass. In front of the vitreous humor, and directly behind the pupil, is a small body with double convex surfaces, called the Crystalline Lens. In the space between the lens and cornea is a thin fluid, denominated from its consistency, the Aqueous Hu- mor. Several of the parts, above enumerated, have exquisite delicacy of structure, and require a very minute descrip- tion. Of the Tunica Sclerotica. The Tunica Sclerotica, or external coat of the ball of the eye, derives its name from a Greek word, which signi- fies to make hard. It is composed of opake white fibres, of great firmness; which form a membrane of very close tex- ture, that supports the globular figure of the eye. It is thicker behind than it is before; but the expansion of the tendons of the recti muscles give it a partial additional covering. It has been considered,by many anatomists, as continued into the cornea; but it can be separated from it by putre- faction; and it is essentially different from it in structure. The aperture in it, which is occupied by the cornea, is not perfectly circular; but inclines somewhat to* the oval form: the transverse diameter being rather longer than the vertical. Posteriorly, it is intimately connected with the optic nerve; which enters it, not directly at the extremity of the The Cornea. 35 \ axis that passes through the center of the cornea and pupil, but on the inside of this spot. The optic nerve has a firm coat, which invests it rather loosely: this coat seems to be continued into, or expanded upon, the sclerotica. Within it, the delicate nerve dimi- nishes considerably before it perforates the sclerotica, and appears to be composed of fibres. At the small place of its penetration, the sclerotica is very thin: and it seems that the nerve does not penetrate through one aperture, but these fibres pass separately through very small foramina, in this thin part. The foramina are necessarily very small, as the diameter of the whole perforated portion, in some eyes, does not exceed two lines or one sixth of an inch. The sclerotica, in its natural state, has few, if any, ves- sels, that carry red blood. The great vascularity, which is so evident in ophthalmia, is in the tunica conjunctiva. The Cornea. The transparent membrane, which fills up the vacuity in the anterior part of the sclerotica, is denominated Cornea; from its resemblance to horn. It is said to be superior in strength to the sclerotica; and it is also very firm. It is formed of lamina, that are separable from each other; which are supposed to be connected by a very delicate cellular membrane. This cellular membrane appears sometimes to contain a fluid; for if a section be made of the coats of the eye, and pressure be applied to the cornea, an exudation will be perceived, both upon its internal and external surfaces. The cornea is covered by the tunica conjunctiva; which adheres firmly to it, but may be separated after macera- tion. It can also be separated from the sclerotica after maceration, and a slight degree of putrefaction; especially 352 The Cornea. if the parts, when in this situation, are suspended a short time in boiling water. The cornea is lined internally by a fine membrane, called the Capsule of the aqueous humor; which will be evident if the coats of the eye are boiled. In this case, the cornea hardens; and the capsule of the aqueous humor appears detached from it, like the cuticle raised by vesication. In a sound state, the sensibility of the cornea varies con- siderably.* The vessels in its structure do not carry red blood: it is however much charged in its structure by in- flammation; and it is said that blood has been found between its laminae, in consequence of violent strokes upon the eye. It is the segment of a smaller sphere than the scleroti- ca; and therefore is more convex. The degree of convex- ity is very different in different persons: those in whom it is very great are necessarily short sighted. It is not per- fectly circular, but rather oval; the transverse diameter being the longest. The cornea and sclerotica are connected to each other by sloping surfaces. The edge of the sclerotica projects over the internal lamen of the cornea; and the edge of the cornea passes under the external surface of the sclerotica. The separation above mentioned proves the cornea and sclerotica to be distinct from each other. And although their structure is essentially different in the human spe- cies, it is much more so in some fishes and in some birds. From what has been already stated, it appears that there are pores in the cornea, through which the fluid situated between its laminae may be pressed. It is probable that an exudation through these pores takes place after death; as a pellicle then forms on the cor- * It is the opinion of Dr. Physick, whose numerous and successful operations on the eye have afforded him many opportunities of judging, that the incision of the cornea always occasions some pain; which is very different, as to its intensity, in different persons Tunica Choroides. 353 nea; which, upon examination, appears to originate from the drying of a fluid effused there. This has been consi- dered as a proof of death; but there are few practical phy- sicians who have not seen a similar pellicle, during life, in persons who were very weak. The Choroid Coat. The sclerotica and cornea compose a firm external shell for the eye; upon which its form depends. The sclerotica is lined by a thin flexible very vascular membrane, denominated Choroides; which is in contact with it nearly throughout its whole extent. The choroides has been supposed to be derived from the pia mater; but this sentiment is not confirmed by ob- servation: for the pia mater appears to be connected with the interior surface of the sclerotic coat. It is so delicate a membrane, and so vascular, that it has been considered by some anatomists as a texture com- posed entirely of vessels and nerves. It has three sets of arteries which are derived from the ophthalmic branch of the internal carotid, viz: 1st. The long ciliary arteries, which are generally two in number: they penetrate the sclerotic coat at the poste- rior part of the eye, and pass, one on each side of the ex- ternal surface of the choroides, dividing at the ciliary cir- cle, each into two branches, which inosculate with each other around the great circumference of the iris. 2d. The short ciliary arteries, which are very nume- rous: they penetrate the sclerotic coat near the optic nerve; are spread upon the choroides, and anastomose very frequently with each other: in their progress forwards they penetrate from the external to the internal surface of the choroides, and supply the iris and ciliary processes. Vol. I. 2 Y 354 Bloodvessels—Nerves—Pigmentum Nigrum. 3d. The anterior ciliary arteries, which are not very numerous; and penetrate the sclerotic coat no great way behind the cornea. These are distributed among the branches of the long ciliary arteries on the iris. The veins of the choroides are very peculiar: besides those, which accompany.the arteries above described, there are several veins which are situated more internally than the arteries and nerves, and about the middle of the eye: their branches are not arranged in the usual manner, but run from the main trunk, nearly in a semicircular curve, are almost parallel to each other, and very numerous: from this arrangement of their branches they are called the Vasa Vorticosa. The nerves, which appear on the choroides, come from the ophthalmic ganglion: they pass very obliquely through the sclerotica, and run forwards; having a very flat appear- ance in proportion to their size: five or six of them are of this description; some others are very small. The internal surface of this coat is covered with a black paste, denominated Pigmentum Nigrum. This appears to be spread most thickly on the anterior part of it, and thin- ner behind, near the optic nerve: it is said to be thinner and less dark in old persons. It seems to resemble the matter of the rete mucosum of negroes, when that is soft- ened by putrefaction. It is asserted that the colour of this pigment has never been changed by the ordinary chemical agenls, or by moderate heat. When it is washed away, the internal surface of the choroides appears to be villous. There is also a portion of this substance on the external , surface of the choroides. It is said that, in very recent sub- jects, this matter appears to be inherent in the structure of the internal surface, and does not come off when it is rub- Ciliary Ligament. 355 bed gently, or even when immersed in water; but it is certain, that a considerable quantity of it sometimes ap- pears on the exterior surface of the choroides, in eyes that are not very recent. The celebrated Ruysch, and many anatomists after him, have considered the choroides as composed of two lami- nae; but it seems to be the opinion of the anatomists of the present day, that it consists of but one membrane; and there are some who consider it as a mere tissue of ves- sels. In the white rabbit, and some other animals, the pig- mentum nigrum is entirely deficient: and the pupil of the eye appears of a red colour, owing to the bloodvessels of the choroides. The general connexion between the choroides and scle- rotica is very slight; depending upon a fine cellular sub- stance, and very small bloodvessels and nerves. But immediately around the margin of the cornea, the choroides and sclerotica are firmly connected to each other; by the intervention of a portion of cellular substance, which, although soft, is dense and compact, and of some thickness. As this substance extends round the circum- ference of the cornea, it necessarily forms a ring; which is between one and two lines broad. This substance, thus placed, constitutes the Ciliary Ligament; which has, to the great perplexity of students, been called by many differ- ent names; as Orbiculus Ciliaris, Annulus Gangliformis, &c It is very distinguishable from the choroides, the scle- rotica, and the iris; and appears generally of a gray colour; but contains the ciliary nerves and arteries, in great num- bers, as they pass to the iris. In this circular band is a small canal, discovered by the 356 The Iris. Abbe Fontana, which is moistened by a pellucid fluid; but its use is not known.* All of the choroides, which is posterior to the ciliary ligament, is in close contact with the internal surface of the sclerotica; but the choroides continues anterior to the ligament; and this anterior portion takes a different posi- tion. It no longer lines the internal surface of the shell of the eye; but is reflected inwards; and takes a transverse direction, as if it were to form a partition. This reflected part forms the ciliary body and the ciliary processes, which will soon be described. The choroides begins to take this reflection inwards, at the place where it is connected with the ciliary liga- ment. Immediately anterior to the ligament, the cornea is continued from the sclerotica; and being more convex than the sclerotica, it projects externally and anteriorly. As the ciliary ligament is situated between the sclerotica and choroides, very near to the place where the sclerotica unites to the cornea, and where the choroides is reflected inter- nally to form the ciliary processes, the edge of the ciliary ligament must lie in the angle formed by the cornea, which is anterior and external, and the reflexion of the choroides, which is internal. To this edge of the ciliary ligament is fixed the circumference of the circular membrane, called Iris, which is now to be described. The Iris Is a flat membrane, which does not partake of the sphe- rical figure of the sclerotica and cornea, or of the cho- roides; but extends across a portion of the cavity of the ' * See Fontana on the poison of the viper, vol. 2, page 310: and also Adolph Murray, Nova Acta Upsal, vol. 3. The Iris. 357 eye, and forms a septum. As it is circumscribed by the ciliary ligament, it is necessarily circular. It has a round foramen near its center, which is called the Pupil; which, in the healthy subject, varies continually in size, according to the degree of light to which the eye is exposed. The situation of the iris, in the cavity of the eye, is such, that, at its circumference, it is nearly in contact with the circumference of the cornea: it is actually in contact with the anterior edge of the ciliary ligament, and with the an- terior surface of that part of the tunica choroides, which is reflected inwards and forms the ciliary processes. As the iris is flat or plain, and the cornea is the seg- ment of a sphere, there must be a considerable vacuity between them. This vacuity constitutes the anterior cham- ber of the eye. The iris therefore is a septum, passing across the eye at its anterior part; and separating that portion of the cavity, which is bounded exteriorly by the cornea, from the larger portion formed by the sclerotica. The anterior, or external, surface of this membrane is very remarkable for its colour: in persons of a light com- plexion it is generally of a light blue intermixed with white; or of a gray, or light hazel, &c. In those whose complexions are dark, it is almost invariably dark also. These colours are so arranged, that frequently there is an appearance of lines, sometimes nearly radiated, more fre- quendy curved in various directions, but tending from the circumference towards the pupil or center: and when the general colour of the iris is blue, these lines are'often whitish. When this surface is examined in water with a magni- fying glass, it appears to be covered with very fine villi; which probably have the effect of increasing its lustre. 358 The Iris. From a supposed resemblance of these colours to those of the rainbow, this membrane has been called Iris; and is generally known by this name: but unfortunately another appellation has been given to its internal and posterior surface, which is covered with a black pigment like the choroides, and has been commonly called Uvea. This black pigment, on the internal or posterior surface of the iris, has been supposed to have a great effect in de- termining its general colour: and there is some reason for this opinion, as the iris is partly transparent; but there is a colour inherent in it, which is most evident when the black pigment is very carefully washed away. Thus, the light hazel colour remains unchanged after the black pigment is removed; and it is not probable that light blue is much influenced by the black pigment. The iris is capable of dilatation and contraction to a very considerable degree; by which the pupil, or central vacuity, is enlarged or diminished. By this means it regu- lates the quantity of light admitted into the eye. Upon the first exposure to strong light, the pupil of every healthy person is observed to be diminished; and, upon the dimi- nution of light, to be enlarged. It has been ascertained by experiment, that this motion of the iris is not excited by the action of light immediately upon it, but on the interior surface of the eye: and this cir- cumstance has occasioned the greatest attention to the structure of this extraordinary and important membrane. Anatomists of the greatest respectability hold opinions, extremely different indeed, respecting the structure of the iris. They agree that there are many radiated fibres on the posterior surface, commencing at its circumference; which can be readily seen when the black pigment is washed awav; but, while many respectable anatomists declare that The Iris. 359 they have not been able to see any circular fibres, Dr. Monro describes them minutely; and has published a plate of them. He considers them as forming a circle imme- diately round the pupil; which circle makes about one fifth part of the breadth of the iris.* In the human subject, the iris does not appear to be di- visible into laminae. It is abundantly supplied with arteries; which form two circles upon it! and it has also a large supply of nerves. The operations of the iris, in contracting the pupil upon exposure to light, and dilating it, when light is diminished or withdrawn, have been explained very differently by persons who have different sentiments respecting its struc- ture. Some of those persons, who do not believe in the ex- istence of muscular fibres in the iris, suppose its motions to depend upon the sudden turgescence or depletion of its bloodvessels: while others impute it to a peculiar quality, exclusively enjoyed by this membrane. Dr. Monro, who refers the contraction of the pupil to the action of the circular fibres, which is excited by the stimulus of light applied to the retina, considers this ope- ration as analogous to that of the abdominal muscles, in those cases of coughing, which are produced by irritation of the glottis. The action of the muscles, in these instances, being excited by substances applied to the glottis, which would have produced no irritation if they had been appli- ed to the muscles themselves. The dilatation of the pupil appears, in the human spe- cies, if not in other animals, to depend upon a different cause. It seems to be a passive state of the eye, which * See his publication, intitled Three Treatises. On the Brain, the Eye, and the Ear. 360 The Ciliary Processes. takes place when the stimulus is withdrawn, and the conse- quent action ceases. There is some elasticity in the iris; and a great many, perhaps all, of its natural or healthy motions in the human species, may be explained by con- traction of circular fibres, induced by irritation; and by the contraction of the rest of the membrane in conse- quence of its elasticity; which always exhibits its effects when the muscular action ceases. The elastic power in this case may have some analogy to the elasticity of the arteries, which certainly produces some of the contraction of those vessels; but there is ano- ther principle of contraction superadded to this, which produces effects different from those of mere elasticity. In the foetus, prior to the seventh month after concep- tion, the pupil is closed by a delicate vascular membrane, called the Membrana Pupilaris; which, after this period, completely disappears. The iris was formerly supposed to be a continuation of the choroid coat; but it is now generally agreed that it is a distinct membrane; for, when the eye is very slightly affected by putrefaction, it can be pulled off, and the cho- roides left entire. The choroides, on the contrary, cannot be separated from the ciliary processes without laceration; and when the pigment is washed away from its internal surface, it is very obvious that these processes are continued from the substance of the choroides, and reflected inwards, so as to form a projection into the cavity of the eye. Of the Ciliary Body, and the Ciliary Processes. This internal projection forms a ring or circle, which has the ciliary ligament before described, near its circum- ference, and anterior to it. The Ciliary Processes. 361 It is so disposed that the whole of its internal or poste- rior surface appears to be formed into radiated plaits which extend from the circumference to the central vacu- ity, where they terminate. The whole of this structure, or the membrane thus plaited, is called the ciliary body, while the plaits are called the ciliary processes. In a natural state of the eye the ciliary body is covered With a large portion of the pigmentum nigrum. It may be seen very well by placing the eye on the cornea, and re- moving the posterior part of all the coats, without deran- ging the humors. When thus viewed from behind, it has a black surface, and appears like a ring formed of radiated lines; it has been compared to a flower with radiated petals. When the black paint is washed off, and the humors of the eye removed with proper caution, the plaited struc- ture of these radiated processes will be very apparent; it then will appear that the membrane of the choroides is so arranged as to form radiated plaits; and that these plaits are the ciliary processes. These plaits or processes do not lie upon each other, but are placed on their edges, one edge looking into the cavity of the eye, and the other edge anteriorly towards the iris. Each ciliary process seems to originate from two lines or smaller plaits, called the ciliary stria, which soon unite and form it. The process or plait, thus formed, is rather larger at its central extremity than at any other part: and their central extremities are not of equal lengths, but alter- nately longer and shorter. , The ciliary processes do not extend to the center of the circle of which they are radii, but stop short of it; and Vol. I. 2 Z 362 The Ciliary Processes. thus include a circular vacuity or aperture; whioh is larger than the pupil of the iris, and situated a little way within or behind it. This aperture is occupied by the crystalline lens (to be hereafter described): but the central extremi- ties or terminations of the ciliary processes do not adhere to the lens, for they are loose and movable; but they arc in contact with its anterior surface, near the margin. It is however to be observed that the ciliary processes are loose only at their central extremities; for, towards their other extremities, they seem to adhere, anteriorly, to the iris; and posteriorly, to the retina, and to the capsule of the vitreous humor. ,The ciliary body, (or the radiated ring formed by all the processes) is about two lines broad in the human subject: but the part next to the nose is rather narrower than the rest of it. The black pigment is spread over the whole of the pos- terior, or internal surface, of the ciliary body; but it is much more abundant towards the circumference than towards the center. It is also more abundant in the furrows be- tween the ciliary processes than it is in the processes them- selves: and as the membrane, of which the processes or plaits are formed, is of a whitish colour, the processes, at their central extremities, have a whitish cast. The black pigment is also spread over the anterior sur- face of the processes; on all that part which is loose and not in contact with the posterior surface of the iris. When the pigment is completely washed away, the co- lour of the ciliary body appears grayish; but if the eye has been successfully injected, it will appear to be composed almost entirely of vessels; and to have a villous surface. 363 The Retina. Within the tunica choroides, and in contact with its in- ternal surface, is a third coat of the eye: the Retina. This coat is evidently derived from the optic nerve, although its texture appears somewhat different. Before the optic nerve arrives at the ball of the eye, a small branch of the ophthalmic artery penetrates its coats: and when the nerve perforates the tunica sclerotica, as de- scribed in page 351, this artery passes with it. On the internal surface of the choroides, the nerve forms a small prominence: and from this, the retina expands, with many ramifications of the aforesaid artery in it. The retina has the appearance of mucus, and the semi- transparency of a surface of ground glass; but when im- mersed in water, it floats like a membrane. By particular management, when the retina floats in water, a considerable quantity of the soft substance may be removed; and a delicate, soft, transparent membranous substance, with the vessels, will remain. The retina,itherefore, seems to consist of a delicate, vas- cular, membranous web, with a medullary pulpy matter spread upon it, and supported by it. Thus constituted, the retina extends from its origin, at the optic nerve, to the commencement of the ciliary pro- cesses. It lines the choroid coat, and is therefore in contact with the pigmentum nigrum, on its internal surface; but it is simply in contact with it, and is not tinged by it. No vessels pass from it to the choroides; and, when a poste- rior section of the eye is made, it slips from it without any appearance of adhesion; being attached only to the optic nerve. Most anatomists agree respecting these circumstances; but there is the greatest difference of opinion respecting 564 The Retina. the extent of the retina. Several distinguished anatomists, of the century lately past, thought very differently from each other on this subject: and two of the first anatomists of the present day have embraced opinions entirely oppo- site. Monro is convinced that it extends, under the ciliary processes, to the crystalline lens: and Soemmering asserts positively, that it terminates at the commencement of the ciliary processes. Some of the* anatomists of London in- cline to the opinion embraced by Soemmering: and the late ingenious Bichat adopted the opinion embraced by Monro. Both parties confide in their own observations, and refer to the eye. An object which appears so very different to different persons, who have good sight, cannot be very dis- tinct. The pulpy substance of the retina appears to termi- nate at the commencement of the ciliary processes; but a membrane, of a very different texture, seems continued from it to the crystalline lens. Although the eye and the retina have long been objects of anatomical attention, professor Soemmering made an interesting discovery respecting it, so lately as the year 1791, viz.: that at the posterior part of the retina, in the axis of the eye, and of course a little exterior to the en- trance of the optic nerve, there is a spot of a circular or oval figure, about one line or rather mare, in diameter, of a yellowish or saffron colour, which is brightest about the center. In the center of this spot is a round hole, equal, in di- ameter, to one fourth of the spot. There are generally se- veral small plaits of the retina about this place: one of these, which is very constant, extends from the optic nerve to this spot. * See the New Cyclopedia, by Dr. Rees and others; article Eye, The Vitreous Humor. 365 The colour of this spot is pale in children, bright in young persons of mature age, and pale at an advanced period of life. The foramen is to be found in the foetus, but not the yellow spot. The colour of the spot diminishes when vi- sion is obstructed; and the spot disappears when vision is lost. There is the greatest reason to believe that the retina is the seat of vision: but it has been ascertained, most de- cisively, that the extremity of the optic nerve, from which the retina originates, is insensible to the rays of light. SECTION III. Of the Humors of the Eye. THE three humors of the eye, viz. the Aqueous, the Crystalline, and the Vitreous, are separately invested with a membranous capsule; which is very delicate, and per- fectly transparent. The Vitreous Humor Occupies almost all the cavity of eye which is posterior to the iris. It of course possesses a spherical form. It has a depression in the center of its anterior surface, in which the posterior surface of the crystalline lens is received. It is covered by the retina as far as the retina extends. The peculiar consistence of this body, which resembles that of melted glass, is owing to its membrane; which is a spherical sac, divided by many septa, or partitions; that form small irregular cavities, in which a fluid is contained. This membranous sac is most perfectly transparent, and very flexible: it has, however, some strength; as it will support the weight of all the fluid it contains; and may be suspended from a hook or forceps. The fluid may be se- parated from the membranes, by beating the vitreous hu- 366 Tunica Hyaloidea. mors in a cup with a spoon; or by suspending the vitre- ous humor, and then puncturing it. In either case the fluid, escapes; but the internal arrangement of the mem- branes is not visible. It is said that when this body is frozen, the portions of fluid in the different cavities, have sometimes been distin- guishable from each other, as distinct pieces of ice: and that their form is that of small wedges; with their edges directed to the crystalline lens as a center, and their bases to the circumference of the vitreous humor, it is also said, that if this body be immersed in a solution of potash, the membranes will become opake, while the fluid continues transparent: and that the appearance of the cells, thus ex- hibited, agrees with the form of the pieces of ice above mentioned. This membrane is now generally called the Tunica Hy- aloidea, from two Greek words which imply a resemblance to glass. Its particular structure is not perfectly understood. Vessels are not generally seen on it in the adult; but, in the foetus, the artery in the optic nerve, (or the central ar- tery,) sends a branch through the vitreous humor to the crystalline lens: and some branches of this artery are to be seen on the tunica hyaloidea. Throughout the greatest part of its extent it appears to consist of but one lamen; but at the front part, near the ciliary processes, there are two lamina. The internal, or posterior, seems to be a portion of the proper tunica hya- loidea, and passes behind the crystalline lens. The exter- nal, or anterior, passes over and before the lens; or at least is attached to the anterior part of the capsule, in which the lens is contained. It is supposed by some anato- mists, that this membrane is continued completely over the crystalline lens; but it has not been separated. Fluid in the Tunica Hyaloidea. 367 As this lamen extends round the crystalline lens, it is in contact with the ciliary processes; or with a production from the retina, which is supposed by some anatomists to pass between them. It is impressed with radiated plaits by the ciliary processes, and some of the pigmentum ni- grum of these processes adheres to it. As these lamina of the tunica hyaloidea are separated by the crystalline lens, there is a vacuity between them, around the margin of the lens: and this vacuity must ne- cessarily be circular in its course. It is not probable that any considerable quantity of fluid is contained in this vacuity; for the two lamina concerned in its formation, appear in contact with each other. It can be readily demonstratedby puncturing the external lamen, and blowing into it through a small pipe. When thus dis- tended, there appear to be incomplete partitions in the canal; or partial adhesions of the lamina forming it, which have the effect of partitions; these partitions are placed in a radiated direction, at some distance from each other, and give the canal a peculiar indented appearance. It is called by the name of the celebrated Petit who discover- ed it. The anterior lamen of the tunica hyaloidea, which forms part of this canal, seems to adhere to the coats of the cili- ary processes; for if a section is made behind the ciliary ligament of all the coats of the eye, the vitreous humor will be found adhering to the anterior portion of the sec- tion, when it is lifted up. When the fluid contained in the tunica hyaloidea is dis- charged and collected, it appears to resemble, in all its properties, the fluid in the anterior chamber of the eye; which, from its consistence, has been called the Aqueous Humor. This circumstance proves that the particular 368 Crystalline Lens. consistence of the vitreous humor is derived from the arrangement of its internal membranes. The fluid, thus obtained, consists of water slightly im- pregnated; 1st, with albumen, 2d, with gelatine, and 3d, with muriate of soda. The vitreous humor appears necessary, to give the ball of the eye the necessary size, for the performance of its optical functions: to keep the retina properly distended: and to retain the crystalline lens at the proper focal di#- tance from the retina. The Crystalline Lens Is a solid body; although it is considered as one of the humors of the eye. It is of asoftish consistence; and has been compared to gum half dissolved; but is more firm in the center than about the circumference. When sound, it is perfectly transparent in young and middle aged persons, but is yellowish in old age. It is convex on both surfaces, but the convexity of the different surfaces is different. The anterior surface, which is the least convex, is the segment of a sphere,' whose diameter varies from six to nine lines. The posterior surface, which is most convex, is the segment of a sphere, whose diame- ter varies from four lines and a half, to five lines and a half. This lens is most convex in young subjects. It is invested with a tunic of the same lenticular form with itself; which has some firmness; but in a healthy state, is also perfectly transparent. The lens either does not adhere at all to this tunic, or so slightly, that it pro- jects from it with very slight pressure, as in the operation of extracting the cataract. The posterior surface of this tunic adheres firmly to the tunica hyaloidea, with which it is in contact: and this last membrane is generally ruptured in the attempt to separate Crystalline Lens, 369 them although the separation can sometimes be effected without rupture. The anterior surface is so intimately connected with that lamen of the hyaloid tunic which passes before it from the canal of Petit, that it is not separable without laceration; and it therefore has not been ascertained whether this lamen extends completely over the surface. As the capsule of the lens is thus connected with the tunica hyaloidea, and this last mentioned tunic adheres to the ciliary body, it necessarily follows that the lens is confined in its proper position behind the pupil. Notwithstanding the soft consistence of the exterior portion of the lens, there is reason to believe that it has a peculiar organization; for if it be immersed some time in a diluted solution of the nitrate or muriate of alumine, or in several other fluids, and then dried, it will be found to be composed of concentric lamellae, which resemble those of the bulbous roots; and these lamellae will appear to consist of fibres. It has been supposed that muscular fibres entered into the composition of the crystalline lens, but this opinion has never been generally adopted.* Between the lens and its capsule a small quantity of a transparent fluid is sometimes found, which escapes whenever the capsule is punctured. This fluid is supposed to be most abundant on the anterior surface. The lens is said by chemists to be composed of albumen and gelatine, with some phosphate of lime; and it is as- serted that the proportion of gelatine is much smaller in the center than in the exterior parts. No bloodvessels are to be seen in any of the humors • See Dr. Young's Memoir in the Philosophical Transactions for 370 Aqueous Humor, of the eye, or their capsules, excepting those of the foetus. In the fcetus a branch can be traced from the central artery of the retina, through the vitreous humor, to the posterior surface of the crystalline capsule, on which it ramifies. In similar subjects vessels have been seen passing from the ciliary processes to the anterior surface of the capsule; and it is also asserted by very respectable anatomists that they have seen vessels passing from the capsule into the sub- stance of the lens. The use of the crystalline lens is to concentrate the rays of light, so as to form a distinct image at the bottom of the eye. The Aqueous Humor Occupies the space which is between the crystalline lens and central extremities of the ciliary processes and the cornea. This space is divided by the iris into two cham- bers, very dissimilar in figure, which communicate with each other by means of the pupil. The posterior chamber is much smaller than the anterior, and its existence has been doubted; but it is easily proved by freezing the eye, when it is found filled with a part of the aqueous humor in a state of congelation. The aqueous humor, in a natural state, is perfectly transparent; but in the foetus, and for a short time after birth, it is reddish and turbid. It consists of water impreg- nated with albumen, gelatine, and muriate of soda; and of course resembles the fluid of the vitreous humor. It is probable that this fluid has also a capsule appropri- ated to it; for after boiling an eye, a delicate membrane can be found lining the internal surface of the cornea, and extending from it over the anterior surface of the iris. It has not yet been traced as far as the pupil, but it is proba- Dissection of the Eye. 371 ble that it extends through the pupil, and lines the poste- rior chamber also. The aqueous humor is quickly renewed after it has es- caped in consequence of wounds or operations: and blood accidentally effused into its cavity is often absorbed. This fluid preserves the convexity of the cornea, and admits the free motions of the iris; allowing at the same time a ready passage to the rays of light. It will be very beneficial to every student of anatomy to dis- sect this delicate organ himself, as he will thereby acquire more accurate and precise ideas of its structure than he can possibly obtain from the ordinary demonstrations. The eyes of sheep and oxen will serve very well for begin- ners, as many of them will be required, and they can be easily obtained. They resemble the human eye in many respects; and the circumstances in which they differ may be readily ascertained by the diligent student; especially if he dissects the human eye afterwards. For this purpose it will be requisite to have forceps, finely pointed; with Knives and scissors that are equally delicate in structure; and also a pair of strong scissors, bended like those in the cases of pocket instruments. When removing the exterior parts, it will be useful to pre- serve a portion of the optic nerve, to take hold of. The dissection of the coats of the eye can be best performed when the eye is placed in a vessel of water; which should be shallow or deep, according to the different stages of dissection. In some dissections it will be serviceable to form a bed of jelly, to support the eye, as proposed by Mr. C. Bell. As the first process in this operation, the sclerotica coat can be readily separated from the choroides, and the cornea with it. After examining the external surface of the choroides, the ciliary ligament, and the iris, the iris may be peeled off from the choroides. 372 Dissection of the Eye. The choroides, by a careful process, may then be separated from the retina, which it leaves surrounding the vitreous hu- mor. The ciliary processes, being a part of the choroides, come away with it. The preparation being in water, may now be suspended by the optic nerve. It consists of the vitreous and crystalline hu- mors, and the retina. The retina, originating from the optic nerve, adheres anteriorly also, with so much firmness that it will support the part inclosed. The retina may be removed without lacerating the tunics of either of the humors; and the pigmentum nigrum which often adheres to that part of the vitreous humor which was con- nected with the ciliary processes, may be washed away with a piece of soft sponge. By this washing, the radiated grooves in the vitreous humor, which were connected with the ciliary processes, will be very apparent. To examine the iris in its natural situation, an aperture may be made in the cornea of a fresh eye near its circumference, and the cornea cut out with the strong scissors. Tftis exposes the anterior surface of the iris. The iris may then be easily removed, and the crystalline lens, with the central extremities of the ciliary processes sur- rounding it, will be exposed to view. To examine the ciliary processes on the other side, a lancet may be plunged into another eye, somewhat anterior to the middle, and the eye divided by means of the strong scissors. The anterior section should then be laid on the cornea, and the ciliary processes will appear very distinctly, seen through a portion of the vitreous humor. The retina may also be seen at the same time, and a judgment may be formed of its extent. The view will be more distinct, if part of the vitreous humor should be cut away with the scissors and forceps, so as to lessen the quantity, without deranging the parts under it. Several methods have been proposed for rendering the structure of the vitreous humor, and the processes of the tunica Dissection of the Eye, 373 hyaloidea, more distinct. None of them have been very suc- cessful: but if the vitreous humor be suspended in water by means of a thread passed through that part which surrounds the crystalline lens, and some large incisions be made into the most depending part of it; after some days of suspension, its bulk will be diminished, by the discharge of the fluid. If, in this situation, it be immersed in a solution of nitrate of silver, the tunica hyaloidea will become very apparent; and by various degrees of exposure to light, may have its color varied from a whitish opacity to a dark brown. The appearance of the retina is also much changed by im- mersion in the solution of nitrate of silver. This effect of the solution,at first sight,appeared calculated to decide the question respecting the extent of the retina; but after several applica- tions it could only be said, that the pulpy substance of the reti- na appeared to terminate at the commencement of the ciliary processes, while a membrane of a different texture seemed continued from it to the crystalline lens. 374 CHAPTER IV. OF THE EAR. THE organ of hearing is composed of three distinct parts, viz. 1. The exterior portion, which, although merely auxiliary, is denominated the External Ear, 2. A cham- ber, situated in the petrous portion of the temporal bone, which is called the Cavity of the Tympanum. 3. A deeper seated cavity in the same bone, which, from its complica- ted form, is denominated the Labyrinth. SECTION I. Of the External Ear. THE External Ear consists of the expanded portion, exterior to the head, commonly called the Ear: and a wide tube passing from it to the cavity of the tympanum, called Meatus Auditorius Externus. The form of the Ear is so familiar to every one that it is not necessary to describe it. The uppermost and longest portion is denominated Pinna. The small pendulous part below is called Lobus. The form, as well as the firmness of the pinna, depends entirely upon a cartilage: the lobus consists of skin and cellular membrane. The skin which covers the pinna is particularly delicate; and when the cuticle is separated from it by maceration, it appears to be perforated with an unusual number of foramina, which are the orifices of the sebaceous glands. It is connected to the subjacent cartilage by a dense cellu- lar membrane, which, in most places, is free from adeps. The External Ear. 375 The cellular membrane of the lobus contains adeps very delicately arranged. The circumference of the ear is formed by a fold of the margin of the cartilage: and this folded edge is denomi- nated Helix, from its winding direction. It commences in the cavity called Concha, to be presently described; and after proceeding forwards and upwards, it turns round backwards and downwards. Within this prominent margin is a second prominence, called the Antihelix, which appears to be formed by a con- vexity of the surface of the cartilage, but is found in every ear. It is nearly semicircular, with its concavity towards the meatus; and it forms the margin of the cavity called Concha, above mentioned. The upper portion of the anti- helix consists of two superficial ridges which unite after forming between them a shallow depression, the Fossa Navicularis, or Scapha. The helix and antihelix form therefore three cavities or depressions, viz. 1. A sulcus, occasioned by the fold of the helix, which is sometimes called Fossa Innominata. 2. The Fossa Navicularis: and 3. The Concha, which may be considered as the proper orifice of the ear. Besides these prominences and fossae there are two other eminences, called Tragus and Antitragus, formed also by this cartilage. The Tragus is anterior to the mea- tus auditorius, and covered by a continuation of the skin of the face. It projects backwards, so as pardy to cover the meatus. On the inside of it there is commonly a tuft of hair, from which its name is derived. The Antitragus is opposite to the tragus, on the poste- rior margin of the concha. On the posterior surface of the ear the convexity of the concha is very conspicuous. 376 The External Ear. When the skin and cellular membrane is carefully re- moved, and the cartilaginous skeleton of this structure is examined, its form will not appear so regular as that of the undissected ear; for there are several deficiencies or vacuities in it; the most remarkable of which is between the tragus and the helix. The cartilage, thus formed, is attached to the bones of the face and cranium by three ligamentous membranes, which connect it to the zygomatic and the mastoid pro- cesses of the temporal bone, and to the aponeurosis on the squamous portion of the same bone. In addition to the common muscles of the ear, described at page 170, there are a few muscular fibres, which, in some subjects, may be discerned on particular parts of the cartilage, and therefore are considered as muscles proper to these portions of the ear. There are five of these portions of fibres: two on the helix, one on the tragus, one near the antitragus, and the fifth on the other side of the ear. The Major Helicis is on the anterior and most promi- nent part of the helix above the tragus. The Minor Helicis is lower down on the helix, in the concha. The Tragicus lies on the tragus, and the Antitragicus behind the antitragus, while the Transversus Auris is on the other side of the ear, on the prominence formed by the concha near its circumference. It may be observed respecting the muscles of the ear, that even the common muscles, which are by far the largest, are unable, in many persons, to move that organ; and the proper muscles, in a large majority of mankind, cannot be perceived to produce any effect at all. There are difficulties in explaining the form of the car- tilage; for, notwithstanding the well known opinion of the The Meatus Auditorius Externus. 377 Geometrician, said to have been employed by Boerhaave, that the line of reverberation is directed to the meatus auditorius externus from every part of the ear; in many persons the cartilage is so situated, that the concha appears to be the principal, if not the only part, from which sound can be reverberated into the meatus. It may be questioned whether the backward position of the ear, which is commonly observed at the present day, is altogether natural; as a more prominent position seems much better calculated for the reverberation of sound; but this position of the cartilage is observable in many infants at birth. It is asserted by some comparative anatomists that the cartilage is stronger and more elastic in proportion to its size, in man, than in any other animal; and that the lobus is peculiar to the human ear. The Meatus Auditorius Externus is a tube, extending from the concha to the membrana tympani; the external extremity of which is principally composed of cartilage, and the internal of bone. The cartilage in the external portion of the tube is a continuation of the substance of the concha. It does not solely form a complete tube, for when the meatus is opened longitudinally and spread out, this cartilage appears triangular in shape; a fibrous mem- brane being joined to it to complete the tube. In addition to the deficiency thus arising, there are two others, one of which, that has a transverse position, is of considerable size. These deficiencies are called Incisura; and the fibrous structure which closes that which is trans- verse, has been called by Santorini, who described it, Musculus Incisura Majoris. The cartilage is attached to the bony portion pf the me- Vol. I. " B 378 Meatus Auditorius Externus. atus at the lower part of its margin, and forms about one half of the length of the meatus. The skin covering the external ear is continued into the meatus, and lines it throughout: extending over the inci- surae, and also over the membrana tympani. It adheres more firmly to the periosteum of the bony part of the me- atus than to any other portion of the canal. Some fine hair is often observable growing out of it at the external extre- mity. As the skin advances deeper in the meatus it be- comes more and more delicate and sensible. The extreme pain excited by the penetration of an insect, or the intro- duction of an instrument, into the ear, evinces the great degree of sensibility with which it is indued. Exterior to the skin, in the cellular membrane which sur- rounds it, are many small glands of a yellowish colour, whose ducts open upon the surface of the meatus, and pour upon it the substance called cerumen or ear wax. These glands are most numerous about the middle of the meatus, and at those places in which the cartilage is deficient. The cerumen is fluid at first, and gradually thickens: in some diseases of the meatus it has the appearance of pus. The use of this secretion is probably to exclude insects, for which it is well calculated by its tenacity and bitterness; and also to defend the delicate surface upon which it is spread. The direction of the meatus is inwards and forwards: it is also curved, with the convexity upwards; but it is very easy, by management of the external ear, to admit the rays of the sun to the bottom of the meatus, and bring the membrana tympani into view. A vertical section of the meatus is of the oval figure. In the foetus the bony part of this duct is not formed, and the meatus consists entirely of cartilage. The Cavity of the Tympanum. 379 SECTION II. Of the Cavity of the Tympanum. THE meatus auditorius externus is terminated abruptly by the membrana tympani, which forms a septum that closes it completely. On the inner side of this septum is the Cavity of the Tympanum, which may be regarded as the continuation of the bony meatus. It differs however from the form of this canal, for its diameter is greater, and it is not so regularly cylindrical. It is not deep, as the distance from the membrana tym- pani, which constitutes the external side, to the opposite internal side, is seldom more than three or four lines. The breadth of the cavity of the tympanum is therefore greater than its depth. It is situated immediately between the membrana tym- pani, and the labyrinth, which is on the inside of it. In its natural state this cavity has two apertures: one which communicates with the fauces, by means of the Eustachian Tube; and another which leads into the cel- lular structure of the mastoid process. There are also two deficiencies in the bony plate, which separates it from the labyrinth, called the Foramen Ovale, and Foramen Rotundum; but in the natural state of the parts these deficiencies are closed, and there is no direct communication of the tympanum either with the labyrinth or the meatus auditorius externus. The only direct com- munications being those above mentioned, with the fauces and the mastoid cells. That deficiency in the bone between the tympanum and the labyrinth, which is called Foramen Ovale, is closed by 380 The Cavity of the Tympanum. one of the small bones of the ear, called the Stapes. The other deficiency called Foramen Rotundum, is covered by the membrane which lines the tympanum. In the cavity of the tympanum are four of the smallest bones of the body, which are articulated with each other so as to form one flexible piece. This piece is attached by one end to the Membrana Tympani, and the other to the Foramen Ovale; and it is moved by small muscles con- nected with it. The cavity of the tympanum is lined by a membrane which has been considered as similar to the periosteum; but it is asserted by Bichat that this membrane, when in- flamed, resembles the mucous membranes; and that in it6 natural state it secretes a mucus which passes into the fauces by the Eustachian tube. This lining membrane is continued over the internal surface of the membrana tympani, and was supposed by the aforesaid author to be reflected so as to cover the small bones of the ear. The opinion of Bichat, respecting the nature of the membrane, is rendered probable by analogical reasoning, viz. The internal surface of the cavity must be in want of mucus or cuticle, as it is in contact with the air. The membrane is very sensible in a sound state, and therefore is very different from periosteum, which it was supposed to resemble. The Membrana Tympani Appears to be fixed in a bony ring in the meatus audi- torius, and is nearly circular in form. It is not perpendicu- lar to the meatus, but has an oblique position in it; the inferior margin projecting further inwards than the supe- rior. The Membrana Tympani. 381 The external surface forms a conical concavity, being apparently drawn in by the malleus, one of the small bones of the ear, to which it is attached. It is asserted by several anatomists, that this membrane may be separated into four laminae, viz. 1. The Cuticle; and 2. The Cutis; which are continued from the skin of the meatus externus. 3. The proper Membrana Tympani: and 4. The Lining Membrane of the Tympanum; which is extended over the internal surface of the membrana tympani. Notwithstanding these laminae, it is almost trans- parent in the living subject, when in a healthy state, and appears highly polished on its external surface, when light is thrown upon it. It has been injected so as to appear vascular in every part; and one or two vessels are some- times seen even in common injected preparations. In some cases of inflammation it has been uniformly red. It was formerly asserted by Rivinus, a professor at Leipsic,that there was a natural aperture in this membrane; but although this opinion has had several votaries, it is certainly erroneous. At the same time it is to be observed, that an aperture, occasioned by accident or disease, is sometimes perceived in persons who enjoy the faculty of hearing to a considerable degree. Some persons thus cir- cumstanced, are accustomed to force tobacco smoke from the mouth through the ears. It is asserted by a respectable anatomist* that the mem- brana tympani, when viewed in a microscope, which will magnify twenty-three times, appears to be supplied with muscular fibres arranged in a radiated manner. These fibres are on the internal side of the membrane. He thinks * See Mr. Home's Memoir on the structure and uses of the Mem- brana Tympani, in the Philosophical Transactions for 1800. 382 Eustachian Tube—-Mastoid Cells. that under favourable circumstances they can be seen with the naked eye. The Eustachian Tube. The communication between the cavity of the tympa- num and the fauces is formed by the Eustachian Tube, a canal which consists of bone at one extremity, and of car- tilage at the other. It commences in the upper and ante- rior part of the cavity of the tympanum, and continues for- wards and inwards through a part of the petrous portion of the temporal bone, above the fissure in the cavity for receiving the condyle of the lower jaw. The bony canal terminates in a very rough and irregular orifice, which is united to a tube composed of cartilage and of membrane, that terminates by a large orifice behind the inferior tur- binated bone of the nose, and on a line with it. This tube is lined by a continuation of the membrane of the posterior nares, which becomes more thin as it pro- ceeds towards the cavity of the tympanum. Above the Eustachian Tube, and separated from it by a thin plate of bone, is a small canal which is nearly parallel to it, and continues so to the end of the petrous bone; this contains the internal muscle of the malleus. The Mastoid Cells. In the upper and back part of the cavity of the tympa- num, nearly opposite to the orifice of the Eustachian Tube, is an opening which communicates with the cells of the mastoid process. These cells do not appear different from those in other bones, and are lined with a membrane ap- parently similar. Their size is proportioned to that of the mastoid process, and consequently they do not exist in the foetus. The Cavity of the Tympanum. 383 Foramina and Protuberances of the Tympanum. That part of the surface of the cavity of the tympanum which is opposite to the membrana tympani, is very irre- gular; but it contributes to the formation of several parts which are very important in the structure of the ear. When the lining membrane is removed, the aperture called Foramen Ovale, appears in a conspicuous situation, rather above the middle of this surface. It would open directly into the Vestibule, or middle chamber of the Laby- rinth; but it is closed by the base of the stapes. Above the foramen ovale is a prominency of the sur- face, in which passes the canal for the Portio Dura of the seventh pair of nerves. Under the above mentioned foramen is a more striking protuberance called the Promontory; within which is a part of the Cochlea, the anterior division of the Labyrinth. At the under and posterior part of the promontory is the Foramen Rotundum, which opens into the Cochlea. This foramen is smaller than the foramen ovale; and in its recent state is covered by the membrane which lines the tympanum. Behind the foramen ovale is a protuberance with a per- foration in it. This is called the Pyramid. It is excavated, and contains the muscle of the stapes, which passes out through the perforation. The Bones of the Ear. The four small bones in the cavity of the tympanum are denominated Malleus, Incus, Orbiculare, and Stapes. The Malleus resembles a crooked club more than any of the hammers now in use. It consists of an irregular roundish head, a neck, and a long tapering body, called 384 The Bones of the Ear. the Manubrium or handle. It has also two processes; one arising from the neck which is long and slender, like a bony fibre, and therefore called Gracilis. The other called Brevis arises from the upper end of the handle. The Incus resembles a molar tooth, with two roots, widely separated from each other. On the body is a de- pression which is connected with the head of the malleus. One of the roots or crura is much longer than the other. The Os Orbiculare is equal in size to a small grain of sand. It is connected to the extremity of the long crus of the incus, and to the upper part of the stapes. The Stapes has a strong resemblance to the common stirrup. The upper part is called its head, the lower part its base, and the two lateral portions its crura. One of the crura is longer than the other. The base is applied to the Foramen Ovale. The situation of these bones is such, that the head of the malleus and the body of the incus are in an upper and anterior part of the cavity of the tympanum, which extends above the membrana tympani. The malleus is the most anterior of the two bones: its manubrium projects down- wards, and is included between the internal lamina of the membrana tympani and the membrane itself. Its long pro- cess extends horizontally, inwards and forwards, into the fissure of the glenoid cavity. The incus is so placed that the depression on its body receives the head of the malleus. The shortest leg projects backwards horizontally, and is attached by a ligament to a point in the opening into the mastoid cells. While the long leg projects downwards like the handle of the mal- leus, but behind it, and at a small distance inwards from the membrana tympani. i Muscles of the Bones of the Ear. 385 The situation of the stapes is almost at right angles with the long leg of the incus, projecting inwards. Between the bead of the stapes and the long leg of the incus, the os orbiculare intervenes. When these bones are viewed in their natural position, the short leg of the incus projects horizontally backwards, the long process of the malleus horizontally forwards; the handle of the malleus, and the long leg of the incus, di- rectly downwards, one connected with the membrana tympani, and the other with the foramen ovale, by the intervention of the os orbiculare and stapes. The Muscles of the Bones of the Ear. The aforesaid bones appear to be regularly articulated for motion with each other, and they are furnished with several muscles. One of these muscles runs in the bony canal above the Eustachian tube, and is inserted into the posterior side of the handle of the malleus, below the root of the long pro- cess. Its effect is to draw in the malleus and membrana tympani. It is therefore called Tensor Tympdni, or Inter- nus Auris. Another muscle, as it is supposed to be, runs in the fis- sure on the outside of the Eustachian tube, and is inserted into the long process of the malleus. This is supposed to draw the malleus obliquely for- ward, and of course to relax the tympanum. It is there- fore called Laxator Tympani, or Externus Mallei; but there are the strongest doubts respecting the muscularity of this organ. Morgagni, Haller, Lieutaud, and Meckel, could not satisfy themselves that it was muscular: and Sabatier also Vol. I. * 3 C 386 Chorda Tympani. doubts of it; while Bell, Fyfe, Hume, Bichat and Gavard, appear to adopt the opinion of its muscularity. Some anatomists describe a third muscle of the malleus as arising from the superior posterior margin of the meatus auditorius, where the membrana tympani adheres to it, and uniting to the neck of the malleus. It is supposed to draw the malleus and membrana tympani upwards and forwards. This however is noticed by few authors. The stapes is also supplied with one muscle called the Stapedius, which lies in the cavity of the pyramid. Its tendon pssses through the foramen in that protuberance, and is inserted into the posterior part of the head of the stapes. It pulls the head of the stapes upwards and back- wards. The Chorda Tympani. In the upper and posterior part of the cavity of the tym- panum is a small nervous cord which enters by a foramen near the basis of the pyramid, and passing downwards and forwards, goes out at an aperture in the fissure of the ca- vity for the head of the lower jaw. In this course it crosses the long leg of the incus, and the manubrium of the mal- leus, and passes between them, being in contact with the manubrium. This nerve comes off from the portio dura immediately before it emerges from the foramen stylo-mastoideum, and after passing a short distance through a small canal, enters the cavity of the tympanum as above described. After passing out of the cavity of the tympanum it joins the lingual branch of the fifth pair of nerves. It was believed by the late Mr. John Hunter that the chorda tympani is not simply a branch of the Portio Dura, but that it is the twig of*the reflected branch of the Observations on the Membrana Tympani, &?c. 387 spheno-palatine fierve, which entering the os petrosum by the Vidian Foramen joins the portio dura; and after pass- ing wit.h it a considerable distance, leaves it at the place above described. There is the greatest reason to believe that the principal object of the structure above described, is to transmit to the labyrinth those impulses of the air which occa- sion sound. The membrana tympani, with the small bones and their muscular appendages, seem to be the agents for this purpose. The Eustachian tube, and the mastoid cells, are subservient parts. The effect of the chorda tympani in the cavity, has not been ascertained. With this view of the subject it is surprising that persons in whom the membrana tympani has been destroyed, should enjoy the sense of hearing in a very considera- ble degree of perfection. Such, however, is the fact. It is necessary that there should be a communication be- tween the cavity of the tympanum and the external air, in order that the function may be duly performed. This is evinced by the deafness which results from the ob- struction of the Eustachian tube, and the cure of this deafness by relieving that obstruction; as well as by the salutary effects of opening the membrana tympani;* and even of instituting a communication through the mastoid cells, in cases where the obstruction of the Eustachian tube could not be removed. It has been supposed that the Eustachian tube has the effect of transmitting sound to the ear, and particularly the voice of the individual of whose structure it is a part: and it is certain that we hear our voices very distinctly, (although peculiarly modified,) when the external ori- • See Dr. Sim's paper on this subject in the first volume of Memoirs of the Medical Society of London: and Ashley Cooper's in the London Philosophical Transactions. 388 The Labyrinth. fices of the ears are closed: but the well known fact that a small watch, when placed in the mouth, and not in contact with any part of it, is scarcely heard, if heard at all, renders this opinion very doubtful. It ought, how- ever, to be remembered, during the investigation of this subject, that persons who hear with difficulty are almost invariably in the habit of opening their mouths when they listen. SECTION III. Of the Labyrinth. THOSE parts of the organ of hearing which have been already described seem calculated for concentrating the vibrations of air; and for communicating, with some modi- fication, the motion they occasion, to the Labyrinth. This important portion of the ear consists of three very dissimilar parts, which communicate with each other and form one general cavity. The central part of this cavity is a chamber of a form which approaches to the oval, and has been compared to that of a grain of barley. It is called the Vestibule. At one extremity of this chamber are three tubes, each of which is curved so as to form a large portion of a cir- cle. These tubes communicate with the vestibule by each extremity; but they form only five orifices, because two of their extremities are united before they open into the vestibule. These tubes are denominated the Semicircular Canals. At the other extremity, of the vestibule is a conical tube convoluted like the shell of a snail. This is called the Cochlea. It also communicates with the vestibule. The Vestibule. 389 The Ijabyrinth, thus complicated in its form, is situated on the inner side of the cavity of the tympanum. Its posi- tion is such that the Vestibule and the Cochlea are opposite to the Membrana Tympani, and the Semicircular Canals are posterior to it. The apex of the cochlea is on the side of the labyrinth which is next to the tympanum, the basis of the cochlea is next to the brain. The texture of the bone which immediately surrounds these cavities is much harder than that of the other parts of the os petrosum; and if the bone of a foetus be used, these softer parts may be cut away so as to leave behind the bony substance which surrounds these cavities, cor- responding exactly to their form. The Labyrinth, when thus prepared, may be consider- ed as a solid body which has been inveloped in a softer substance, and is brought into view by detaching the soft substance which surrounded it. In the foetus it is nearly as large as in the adult, so that the structure of the ear can be investigated with great ad- vantage in such subjects. The Vestibule Is situated within the Foramen Ovale. There are two remarkable depressions of its internal surface; one, which is in the superior part, is called Semi Elliptical; the other, which is below, has the name of Hemispherical. When the dried preparation is examined there are several foramina in this cavity, viz. The Foramen Ovale, already mentioned. A round aperture, by which it com- municates with one of the cavities of the cochlea, and the five openings, of the semicircular canals. Besides these, there are several small perforations from the Meatus Au- ditorius Internus for the transmission of nerves. 390 The Cochlea. The vestibule, as well as the other parts of the laby- rinth, is lined by a fine membrane which has been consi- dered as the periosteum. The Cochlea Commences at the anterior part of the vestibule. It is a conical tube, so convoluted that it has the form of the shell of a snail, making two circuits and an half round a center. It may be considered as wound in a spiral direction round a pillar of bone. To this central pillar the name of Modiolus is applied. It commences at the cavity called Meatus Auditorius Internus; and its base is somewhat excavated. It gradually diminishes in diameter as it pro- ceeds towards the apex of the cochlea, and is therefore conical in form; but it does not preserve this form to its termination, for near the apex it gradually becomes broad- errand thus forms a second cone inverted. This last por- tion of the central pillar is called the Infundibulum. It is hollow; and the portion of bone which covers its cavity, constitutes the basis of the inverted cone and the apex of the cochlea. It is called the Cupola. The tube thus wound round the modiolus, or the coch- lea, is divided from the beginning to the end by a partition. The cavities are called Scala, and the partition Lamina Spiralis. The Lamina Spiralis is made up of four parallel strips, which compose its breadth. 1st. A plate of bone. 2d. Outside of this, a softer plate which appears carti- laginous. 3d. A cellular portion, which appears to contain a pellu- cid fluid. Lamina Spiralis and Scala of the Cochlea. 391 4th. A thin membranous strip, which completes the septum or partition. These parts may be distinguished from each other when magnifying glasses of sufficient power are used. The bony plate is composed of two laminae with small cancelli'between them, in which are canals for the trans- mission of the fibres of nerves. These canals are extended into the cartilaginous part. The membranous part which completes the septum is continued into the lining membrane on the surface of the cochlea. The bony portion of the scala does not extend towards the apex of the cochlea so far as the cartilaginous and membranous portions; and none of them continue to the apex, for the lamina spiralis terminates in the infundibu- lum before it has arrived at the apex. Its extremity has the form of a hook, and is therefore termed Hamulus. As this septum does not extend to the apex of the coch- lea, the two scalae necessarily communicate with each other at the apex. These different bands or strips which compose the lamina spiralis, are called its Zones, and are termed Zona Ossea, Zona Coriacea, Zona Vesicularis, and Zona Mem- branacea. From what has been stated it follows that there are two cavities in the cochlea, each of which continues throughout its whole extent. One begins at the vestibule; the other at the tympanum. But a membrane extended over the fo- ramen rotundum separates this last from the tympanum. They communicate with each other at their terminations. * From their origins they are denominated Scala Vestibuli, and Scala Tympani. 392 Meatus Auditorius Internus. As both the cochlea and the vestibule are filled with a fluid, it is evident that a vibration produced on the mem- brane of the foramen rotundum may be communicated through the two scalse to the vestibule. The three Semicircular Canals Are placed obliquely behind the vestibule. Their posi- tion is such that one is Superior, another Posterior, and the third Exterior. The superior and posterior are so placed that one extremity of each may be considered as internal and the other external. They unite at their inter- nal extremities, which therefore form but one orifice in the vestibule. Their other extremities being separated to a considerable distance, form each one orifice. While the external canal, which is smaller than the others, opens by two orifices. Each of these canals is nearly of the same diameter, viz. rather more than two lines. At one of their extremities each of them has an en- largement, which is called Ampulla; and there is no other variation of their diameters. The cavity of the labyrinth, thus complicated, is perfo- rated by many small foramina, through which various nerves are transmitted. These foramina communicate with the large canal on the posterior side of the petrous bone called Meatus Audi- torius Internus; which continues very near to the basis of the cochlea, and transmits the seventh pair of nerves. The bottom of this cavity is divided by a ridge into two unequal fossae; the uppermost of which is the least. In the Upper Fossa are two foramina; the anterior, which is the largest, serves to transmit the Portio Dura, a. part of the seventh pair, to be hereafter described, which Contents of the Labyrinth. 393 passes through the petrous bone to the face. The poste- rior foramen forms a pit with a cribriform bottom, which admits nervous fibrillae to the vestibule. The anterior part of the Inferior Fossa is also cribri- form; its perforations lead to the cochlea; one of them which passes through the modiolus to the infundibulum is larger than the rest. The posterior part of this fossa is occupied by foramina, which pass to the vestibule and semicircular canals; but they are not so numerous as those "which lead to the cochlea. Contents of the Labyrinth. This interesting cavity is lined throughout by a delicate membrane. It contains sacs and tubes, and a plexus of delicate nerves, which constitute a soft labyrinth within that which is composed pf bone. In the cavity of the Vestibule are two sacs distinct from each other, and also from the lining membrane. One of these, denominated Sacculus Sphericus, or Sao cuius Vestibuii, is situated partly in the Hemispherical Ca- vity of the vestibule, and has no direct communication with any other part. It contains a limpid fluid, and is said to have so much -firmness, that when opened with the point of a lancet it will retain its form. The other sac is situated partly in the depression called Semi-elliptical, nearly opposite to the foramen ovale; it is so transparent that it is sometimes seen with difficulty, and appears like a bubble of air in a fluid. All the mem- branous semicircular canals, which are soon to be descri- bed, communicate with it by each of their extremities; it has been called Alveus Communis by Scarpa, and Utriculus by Soemmering. Vol. I. 3D 394 The Auditory Nerve. "> In the bony canals already described are three Membra- nous Semicircular Canals, which resemble them in form. They have an ampulla at one extremity, as they arise from the sac above mentioned, and are cylindrical during the remainder of their course; they are transparent, and have smaller diameters than the bony canals, although they are rather larger. The Auditory Nerve. Upon this structure, viz. the Sacs in the Vestibule, the Membranous Tubes in the Semicircular Canals, and the Lamina Spiralis of the Cochlea, are expanded the fibres of the Auditory Nerve. This nerve, with the Portio Dura, and its appendage, the Portio Media, composes the seventh pair of nerves of the brain. It is called Portio Mollis, and is very distinct from the Portio Dura, although they pass together along the Mea- tus Auditorius Internus. Corresponding to the foramina and the cribriform struc- ture of the bottom of the Meatus Auditorius Internus, the Auditory Nerve passes into the Labyrinth in branches, or fibrillae, of various sizes. One portion of them enters the Vestibule, and has been traced upon the Alveus or Utriculus, and its internal surface; and also upon the Semicircular membranous tubes. Another portion seems exclusively appropriated to a part of these tubes. And a Third is spent upon the Sacculus Sphericus. These nervous fibres seem to terminate in a pulpy ex- pansion on the internal surface of the aforesaid sacs and canals, in a way which has some analogy with the termi- nation of the optic nerve. The AqueduSts. 395 A large bundle of these fibrillae enters the Cochlea at its base; and the largest of them passes through a foramen, mentioned before, along the center of the Modiolus to the Infundibulum. These fibrillae divide most minutely, and passing be* tween the plates of the Lamina Spiralis, as well as the other parts of the bony structure of the Cochlea, at length form a plexus, which has the appearance of a pulpy menir brane, that is extended over the whole of the Lamina Spi- ralis. Thus is the Auditory Nerve distributed. To complete the account of the Labyrinth, it is to be observed that a pellucid fluid certainly exists in it; exterior to the sacs in the Vestibule, and to the Membranous Semi- circular Canals. So that the Membranous Labyrinth may be said to be immersed in a fluid. This fluid fills also the Cochlea. The Aqueducts. It is probably on account of this fluid, that two small canals exist; which are called, after the anatomist who first suggested their use, the Aqueducts of Cotunnius.* One of these commences in the Scala Tympani of the Cochlea, near the foramen ovale; and terminates in the jugular fossa, by a small orifice, situated before the spine that separates the eighth pair of nerves from the internal jugular vein. It is called the Aqueduct of the Cochlea. The other originates in the Vestibule, under the com- mon orifice of the two canals; and terminates on the pos- terior surface of the petrous bone, by a small orifice, which is situated at some distance behind the Meatus Audito- rius Internus. It is called the Aqueduct of the Vestibule. • See Sandifort's Thesaurus Dissertatlonum, Stc. vol. I. 396 Portio Dura. To this account of the ear it ought to be added, that the Portio Dura, after entering into the petrous bone by the foramen in the upper fossa of the Meatus Auditorius In- ternus, proceeds, in a canal which is called the Aqueduct of Fallopius, through the bone, to the Foramen Stylo Mas- toideum, on its inferior surface; where it emerges, and is distributed to the face. It is therefore called the Facial Nerve by some anatomists. In the course of this nerve from the Meatus through the solid bone, it forms a remarkable angle, and then passes, between the Cochlea and the Semicircular Canals, to the foramen Stylo Mastoideum, In this course it soon receives the Vidian Nerve already mentioned; and it sends-off the Chorda Tympani, imme- diately before it passes out at the Foramen Stylo Mastoi- deum. It also sends off small fibrils to the muscles of the bones of the ear. It has not been ascertained whether the Portio Dura, the Vidian Nerve, and the Chorda Tympani, have any effect upon the function of hearing. The situation of those branches of the Auditory Nerve which are expanded in the Vestibule and the Semicir- cular Canals, is somewhat different from the situation of those which are in the Cochlea; but it has not yet been ascertained how far their functions are different. The information on this subject derived from compara- tive anatomy, is very interesting; but for want of more acquaintance with the state of this function in the dif- ferent animals, no very decisive inferences have been drawn from it. The Vestibule and Semicircular Canals occur much more frequently than the Cochlea, which is to be found in few animals, if any, besides those of the classes of Mam- Functions of the different parts of the Labyrinth. 397 maha and of Birds. It is therefore supposed necessary to that perfect state of hearing which the animals of these classes enjoy. But there remains a considerable diffi- culty on this subject; the Cochlea is not by any means so perfect in birds as in quadrupeds; yet many birds appear to have clear perceptions of musical sounds, and some birds imitate articulate sounds with consider* able accuracy. That the impression which produces hearing is made on the nervous expansions in the Labyrinth, does not ap- pear to be doubted by any one. The structure of the whole organ, and the analogy between it and the eye. induce a strong belief that this is the case. This belief is confirmed by a dissection recorded by Mr. Haighton,* in which Original Deafness was found to de- pend upon a quantity of cheese-like matter, which filled the whole Labyrinth, and was attended with a considerable diminution of the size of the Auditory Nerve: while all the other parts of the organ were in a perfectly natural state. "See Memoirs of the Medical Society of London, vol. 3 SYSTEM OF ANATOMY. PART V. OF THE GENERAL INTEGUMENTS: OR THE CEL- LULAR MEMBRANE AND THE SKIN. CHAPTER I. OF THE CELLULAR MEMBRANE. A. HAT substance which is situated between the skin and the muscles, which is insinuated between the different muscles, and between the fibres which compose these; which also connects the different parts of the body to each other, is denominated the Cellular Membrane, or Tela Cel- lulosa. As it extends over the whole of the body, and is most intimately connected with the skin, it is considered as one of the integuments, although it is found in great quantities in some of the internal parts. It appears to be composed of membranous laminae, ex- quisitely fine and delicate in their structure, which are so connected to each other that they compose cells or cavi- ties of various forms and sizes. The Cellular Membrane. 399 When these cavities are empty, this arrangement of the cellular membrane is not apparent; but when they are dis- tended by water or air it is very evident. The laminae which pass from one contiguous part to another are of different lengths, according to the motions performed by the different parts; thus, about the muscles and their tendons they are of considerable length, and be- tween the coats of the eye they are very short. In some places these laminae are compressed together, and form a dense membrane somewhat resembling ten- don; but whenever they are separated from each other they appear pellucid, and extremely delicate. These laminae, when in a healthy state, appear to have no sensibility; but so many nerves pass through them, that pain is generally felt when incisions are made in the cellu- lar membrane. No vessels can be seen in their composition when they are free from disease, although many pass through them. On this account they have been considered by some very respectable physiologists as inorganic; but there are good reasons for regarding this sentiment as erroneous. If a portion of cellular membrane, in the living subject, be brought into view by a surgical operation or a wound, and be allowed to remain sometime, covered by an emollient cataplasm, or a soft plaster, a complete change of color will gradually take place; it will become uniformly red, in consequence of the great number of minute vessels into which blood has penetrated during inflammation; and gra- nulations will form on its surface. These vessels must have existed previously in the sound state of the membrane, and conveyed a transparent fluid; although no structure of this kind was visible. This single fact therefore proves completely its organization. 400 The Cellular Membrane. In some parts of the body, this cellular membrane ap- pears to be moistened by a small quantity of fluid, or hali- tus, in its cells; which seems merely sufficient to keep it soft and flexible. In other places it is loaded with fat. There is great reason to believe that the fat is contained in cavities which are somewhat different from the ordina- ry cavities of the cellular membrane. The cells or cavities which contain the moisture or ha- litus communicate with each other, over the whole body. Thus, air insinuated into the cellular membrane exterior to the pleura, in consequence of a fractured rib, will be dif- fused over the whole body; and produce the disease called emphysema. In a patient who is affected with that spe- ties of dropsy called anasarca, a portion of the fluid will be effused in the head and upper parts of the body, after he has passed a night in bed in a horizontal position; but af- ter Jie has been in an erect position for some time, the fluid will be accumulated in the legs and feet, or most depend- ing parts of the body, in consequence of its gravity. It is well known in dissecting rooms, that the effused water may be completely discharged from anasarcous sub- jects, by making incisions in the feet, and placing the sub- ject erect. Blood effused in the cellular membrane is sometimes dispersed in the same way: an ecchymosis often appears in the eyelids, in consequence of a coatusion on the upper part of the head; and similar appearances occur in almost every part of the body, in consequence of effusions of blood at a distance from them. The fat or adipose matter is not diffused in this man- ner: wherever it is first effused, it remains; uninfluenced by gravity, or the ordinary pressure. Fat is not observed in every part of the body; it is ne- The Reticular and Adipose Membranes. 401 ver seen in the cellular membrane of the eye lids; of the penis; of the lungs; or of the parts within the cranium; as well as of several other places. The inconvenience which would result from the accumulation of fat in these places is very obvious: and it is equally certain that the cellular membrane, in them, must be different from that in which fat is produced. From these peculiar circumstances, relative to the adeps, it has been inferred, that there was a peculiar apparatus for the production and retention of fat, superadded to the cel- lular membrane; and some anatomists, with a view to precision, have called the part containing fat, Adipose Membrane, and the other part Reticular Membrane.* They state that in dropsical subjects, who are much emaciated, the membrane, which in a healthy state contained adeps, is more ligamentous than the ordinary cellular membrane. It seems to be proved, by reasoning, that there must be a considerable difference between these different parts of the cellular membrane; but it ought to be observed that those parts of the omentum which are especially appropri- ated to the production of adeps, do not exhibit any pecu- liarity of structure. This adipose substance is distributed in unequal pro- portions in different parts of the body. In corpulent per- sons there is a considerable quantity of it immediately under the skin, and especially under the skin of the abdo^ men. It is also between the muscles; in the orbits of the eyes; in the omentum and mesentery; in the joints and the bones; as well as about the kidneys, and the heart also, in elderly persons. In the foetus, and for some time after • See remarks on the cellular membrane, &c. by Dr. W. Hunter, in the London Medical Observations and Inquiries, vol. JI Vol. I. 3 E 402 Connexion of the Cellular Membrane and Skin. birth, it appears to be confined to the parts immediately under the skin, but it soon becomes more diffused. It is observed by dissectors that there are no subjects, however emaciated, which are entirely free from fat; ex- cept those which have been affected with anasarca. The cellular membrane has been already observed to form granulations very promptly; and it has been asserted that the granulations, which arise from all the different parts of the body, when wounded, originate from the cel- lular membrane in those parts. Whether this proposition be true or not, to the extent above stated, it is a fact that granulations, in some instan- ces, seem to have a cellular structure; as the following case will prove. A patient with a compound fracture of the leg, which was attended with a large wound, covered with luxuriant granulations, was attacked with an oedematous swelling of the limb, which increased suddenly to a great degree. While this was going on, the granulations on the surface of the wound tumified with the limb; and upon examina- tion, appeared somewhat pellucid with an effused fluid; indenting by pressure, precisely as the skin was indented. The cellular memSrane appears to have a most intimate connexion with the skin; and cannot be completely sepa- rated from it by dissection. It is said that in certain cases of disease, where it is reduced to a slough, while the tex- ture of the skin remains unchanged, as in some species of anthrax or carbuncle, this separation may be completely effected; In such cases the under surface of the skin will appear to be composed of pits or excavations, which pene- trate very deep into its substance, and were occupied by the cellular or adipose membrane while it was in its natu- ral condition. 403 CHAPTER II. OF THE SKIN. The skin is composed of three dissimilar laminae, which are denominated the Cutis Vera, the Rete Mucosum, and the Cuticula. SECTION I. Of the Cutis Vera. JL HE innermost of the above mentioned laminae is much more substantial than the others, and therefore is called Cutis Vera. It is an elastic, dense, and strong membrane; which con- tains in its texture a large proportion of fibres that appear to be tendinous, and are woven together in an intricate manner. Blended with these fibres is an immense number of ves- sels which enter into the texture of the skin; these ves- sels do not generally convey red blood, and therefore they are not very visible; yet they may be readily brought into view, by the application of rubefacients during life; and, by fine injections, in the dead subject. Their existence is also demonstrated in the vigorous infant, at birth, by the uni- versal redness of the skin, which is observable at that time. Nerves are also distributed to every part of the skin. They can be traced to it very easily; and as there is no part of the skin, into which the finest needle can be pushed without pain, it is certain that their distribution must ex- tend to every part. 404 Cutis Vera. It is highly probable that the processes of absorption and exhalation are effected by small vessels which origi- nate or terminate on the surface of the skin, and of course form a part of its texture. The skin, thus constructed, extends over the whole of the body, and is continued into those cavities which open upon the surface, as the. mouth, nose, &c, although its texture changes immediately upon its reflexion. It varies in thickness in different parts; thus, it is thicker on the back than the front of the body. It is thin on the insides of the arms and legs, where opposite surfaces touch each other. It is in general thinner in women than in men. The elasticity of the skin is made evident by its yielding to distension, and returning to its usual size; as in preg- nancy, dropsy, &c: but it is particularly demonstrated in some cases of parturition, when the skin of the perinaeum stretches immensely, and, after labour, very quickly reco- vers its natural size. The external surface of the skin is very generally divi- ded by superficial grooves or sulci, into small spaces of various angular forms; most commonly rhomboidal. On the palms of the hands and soles of the feet, instead of these figures, we perceive the whole surface composed of furrows and ridges, which in some places are rectilineal and in, others oval and spiral. There are also a number of depressions or grooves Which seem formed to accommodate the various articula- tions, particularly about the fingers and toes. There are other furrows, occasioned by muscles, as those on the forehead; and some depend on the subjacent cellular membrane. Pores, &?c. of the Cutis Vera. 405 On the external surface of the true skin, when the two exterior laminae are removed, many papillae are to be seen. They differ in size in different parts of the body: they are vascular, and, on the ends of the fingers, appear like villi, when examined with a magnifying glass. There are many perforations or pores to be seen on the skin with the naked eye, which are probably the ducts of sebaceous glands, and the passages which transmit hairs. Other pores, different from either of these, are to be seen when magnifying glasses are used; as those on the fingers: these probably are the exhaling or absorbing pores, but their connexion with the vessels which perform these func- tions has not yet been demonstrated. The internal surface of the skin, when carefully dissect- ed from the subjacent cellular membrane, in a subject of ordinary corpulency, appears to have some adipose sub- stance in its texture; but, as has been already mentioned, when the cellular membrane is destroyed, these portions of adipose matter disappear, and the surface of the skin ap- pears pitted. It is probable that this connexion of the cellular membrane and skin may occasion that delicacy of skin which appears in some hydropic patients. In some places on the under surface of the skin are small glands called miliary, from their resemblance to the millet seed; these glands are supposed to secrete a seba- ceous matter, but they are not so general as has been sup- posed. There are sebaceous follicles or ducts which open on the external surface of the skin, and contain an oily substance, which sometimes has the consistence of suet or tallow: when these ducts are filled with sebaceous matter, their orifices are often covered by a black substance which acci- dentally adheres to the surface of the matter,and forms very 406 Cutis Anserina. small black spots in the skin. These often occur on the nose and ears, and may be removed by pressing out the sebaceous substance, which rises up in the form of small worms. Sometimes this secretion accumulates in the ducts in such quantities that it forms small tumors in the skin. Muscular fibres have been supposed by some persons to exist in the skin, but such fibres have never been de- monstrated in it. The skin of the scrotum is often much contracted, but the fibres which produce this effect are ve- ry visible in the cellular membrane, and have a muscular appearance. Although the skin is not muscular, it sometimes chan- ges its appearance in a surprising manner. When the surface of the body is suddenly exposed to cold, or when the chill of fever exists to a considerable degree, the skin will contract very sensibly; and at the same time a great number of conical papillae will project from its surface. This constitutes the Cutis Anserina; and is supposed to be produced by a sudden contraction of the vessels in the skin, which forces out their contents, and of course diminishes its bulk; while the papillae do not con- tract in the same degree, and therefore are somewhat pro- jected. When the skin is free from disease, the two exterior laminae may be separated from it completely, after macera- tion or putrefaction, and the surface will appear smooth; but in an inflamed skin, a network of vessels has been injected; which is considered by Mr. Cruikshank* as an additional lamina. In this lamina the pustules of smallpox originate. When the skin is injected they appear to be formed at first * See Experiments on Insensible Perspiration, &c. by W. Cruikshank. Vessels on the External Surface of the Skin. 407 by very small vessels, arranged in a radiated manner, with a white uninjected substance in the center, which is sup- posed to be a slough, occasioned by the irritation of the variolous matter. Mr. Cruikshank after removing this lamella was able, by continued maceration of the same skin, to separate another, which was also vascular. It is to be observed that this skin had been preserved for some time m spirits, and was macerated in putrid water a week du- ring the heat of summer, before the first lamella was re- moved. The colour of the healthy skin is invariably white, when all the lamellae exterior to it are removed. This is the case not only with the European, but with the blackest African, and the people of all the intermediate colours. The variety of colours in the human species depends upon the lamella next to the cutis, which is now to be de- scribed. 408 SECTION II Of the Bete Mucosum. IMMEDIATELY in contact with the external sur- face of the cutis vera is a thin stratum, of a pulpy or mu- cilaginous consistence, which appears to be spread uni- formly over it, but cannot be detached without deranging its own texture.* It can be best examined after the cuticle is raised in a blister. In this case it appears like a pulpy substance, spread upon a membrane of a soft and delicate texture. This is the Rete or Corpus Mucosum. In this pulpy substance resides the pigmentum, or co- louring matter, which gives the peculiar complexion to the different races of men. The cutis vera is white, and the cuticle is nearly transparent in them all; but this substance is black in the negro; copper-coloured, yellow, or tawny, in many of the Asiatics; and yellow, with a tincture of red, in the aborigines of America; while it is transparent, or whitish, in the people of Europe and their descendants. It can therefore be best examined in the negroes; and if it be inspected immediately after the cuticle of a blister is removed it will appear as above described, with a black matter diffused through it. The particular structure of this substance has not been ascertained; although anatomists have paid a good deal of attention to it. It is generally believed by them that no vessels can be injected in it; but Dr. Baynham of Virginia, while he was engaged in anatomical pursuits in London, * It has been asserted that the rete mucosum of the scrotum can sometimes be exhibited in a separate state. Vessels between the Rete Mucosum and Cutis. 409 made a preparation which excited the attention of the Bri- tish anatomists, on account of its particular relation to this subject. He injected one of the lower extremities, the os femoris of which was diseased with an exostosis; and with a view to an examination of the lamina? of the skin, he removed a portion of it from the leg; and after immersing it a few seconds in boiling water, to thicken the laminae, he macer- ated it in cold water for some days. Upon separating the cuticle, after this treatment, he discovered a texture of ves- sels on the surface of the cutis vera, which was distinct from the cutis itself. This has often been mentioned as injection of the rete mucosum. It is to be regretted that Dr. Baynham, who is particu- larly qualified to decide, has not published his opinion on the subject. Mr. Cruikshank', to whom he afforded the most satisfactory opportunity of examining his prepara- tion, believes that the aforesaid vessels were not a part of the rete mucosum; but that the rete mucosum was to be seen on the epidermis (being raised with it when it was separated from the cutis), while this texture remained on the surface of the cutis. He considers these vessels as be- longing to the additional lamellae already mentioned, of which he says Dr. Baynham is the discoverer. There is therefore every reason to believe that there is a texture of vessels, either in the rete mucosum, or between the cutis vera and the rete mucosum. After putrefaction, or maceration for a long time, trie cuticle separates readily from the cutis vera; and the rete mucosum sometimes adheres to the skin, and sometimes to the cuticle. If the parts are much softened by putrefac- tion, the rete mucosum can be washed away, like the pig- 410 Change of Colour in Negroes. mentum nigrum of the eye; leaving the cutis white, and the cuticle nearly transparent. In the negroes the black colour of the rete mucosum is greatly diminished, on the palms of the hands, and soles of the feet, and under the nails; but it is perceptible. It is said that the black colour does not appear in the cicatrices of the blacks. This is the fact with respect to recent cica- trices; but those of long standing are often dark coloured, although not so black as the original skin. The pits of the smallpox in their skins, although white at first, become finally as dark as the original surface. In Europeans and their descendants the colour of the rete mucosum becomes darker, as they are more exposed to the air and the rays of the sun; and soon changes again to its original fairness, by confinement to the house. In negroes, the skin loses some of its deep glossy black colour during the winter season of cold climates; and re- covers it again in summer. The rete mucosum sometimes undergoes very impor- tant changes; there have been several instances in the United States, where large portions of the skin of the African have changed from black to white; owing proba- bly to an absorption of the black pigment from the rete mucosum; or perhaps to an absorption of the rete muco- sum itself. There is now in Philadelphia a female, between thirty and forty years of age, in whom this process is going on. One of her parents was a negro and the other a mulatto; and her original complexion accorded with her origin. But a change of colourbegan during her childhood, in small spots, which have gradually increased so much, that at this time the whole of her body and limbs are nearly white, with the exception of her hands and feet. A large Changes of the Rete Mucosum in White Persons. 411 proportion of her face is also white, and the remainder of it much lighter than it was originally. At this time, some part of her face has an unnatural whiteness; but the skin of her fore-arms appears like that of an European in a per- fectly healthy state. This change of colour is attended with no unusual sensation; so that if she did not see the altera- tion, she would not suspect that her skin was any way dif- ferent now from what it had originally been. She does not appear sensible that the white parts are more suscep- tible of irritation from the rays of the sun than they were originally; but they are so much covered by her dress that the experiment has not yet been fairly made. The first appearance of a change is slight diminution of the dark colour; this change goes on gradually, and then small spots appear, which are perfectty white. They gra- dually increase, and run into each other, and thus a large white spot is formed. In a former case, where this process had gone on to a great extent, it is said that the black pigment was again deposited, and the skin resumed its original blackness. These circumstances in negroes have been considered as great deviations from the ordinary course of nature, but a process very analogous to it sometimes goes on in per- sons who are white. Thus, there are some in whom the skin becomes much browner than natural in some parts of the body, particularly on the arms; and in these brown portions, spots are formed which are much more white than the natural colour of the skin. In such cases there appears to be a deposition of colour- ing matter in the rete mucosum of the brown places; while the white spots are rendered more white than natural, either by an absorption of the rete mucosum, or by a de- position of whiter matter in it. 412 Changes of the Rete Mucosum in White Persons. The colour of the rete mucosum sometimes undergoes a temporary change in particular places. Thus, at a certain period of pregnancy, a dark circle forms round the nipple. In some cases, where the peculiar whiteness occurs, the skin becomes very susceptible of irritation from the rays of the sun; so as to be blistered, if exposed to them for a short time: this circumstance renders it probable that the colouring matter in the rete mucosum of the. blacks, was originally designed to protect their skins from the very powerful rays of the sun to which they are exposed. There are some persons to be found, among most of the different races of men, who are born with this peculiar whiteness of the whole skin, which continues during life. In these persons, the hair has a remarkable white colour, and the eyes are without the pigmentum nigrum. They appear to be in a state of imperfection, and are unable to endure the ordinary light of day. They are generally designated by the epithet of Albinoes. The texture which exists between the cutis vera and the epidermis is probably the principal seat of several im- portant cutaneous diseases; as the Scarlatina, Pemphigus, &c.;* and from what has been stated, there is good reason to believe thnt the smallpox also commences in it. It is therefore much to be wished that its structure* was more precisely ascertained. * In severe cases of the scarlatina, at the termination of the disease, large portions of the cuticle are sometimes detached from the cutis, so that several practitioners have seen the whole cuticle of the hand come off like a glove. As the texture of the cutis does not appear to be alter- ed in these cases, and the cuticle is also unchanged, the cause of this separation must exist in the intervening-structure which connects them 413 SECTION III. The Cuticula or Epidermis HAS been examined with the greatest care by several of the most successful anatomists; but notwithstanding their labours, the structure of this substance is by no means un- derstood. It appears to have some resemblance to the matter of the nails, and of horn; but is rather more flexible, even af- ter allowing for the difference of thickness. In those parts where it is thinnest it is semitransparent. It is insensible, and no vessels can be seen in it.* It extends over the whole external surface of the body, except the parts covered by the nails; and is accommoda- ted to the surface of the skin, by forming ridges or furrows, corresponding to it. It adheres most closely to the cutis; and when abraded by mechanical violence, the surface of the skin appears moistened by effusion. It is not certain that its mode of union with the skin is perfectly understood; the adhesion of these membranes to each other is as uniform as that of two smooth surfaces glued together, but it.is generally said that the cuticle is attached to the cutis by very numerous and fine filaments. It has often been asserted that these filaments are the exhaling and absorbing, vessels, which pass through the * In the early part of the last century an anatomist of the name of St. Andre exhibited a preparation of the cuticle which appeared to be in- jected with mercury. Ruysh declared the thing impossible, and in- vited him to an investigation of the subject. This invitation was not ac- cepted, and the affair has been generally considered as a mistake or an imposition. 414 Pores of the Cuticle. cuticle, to and from the skin. This sentiment appears verj reasonable, but no vessels that pass in this way can be in- jected. There are innumerable processes which pass from the cuticle to the skin. Many of these are the linings of the cavities which contain the roots of the hairs; but they are reported by microscopical observers to be, like the fingers of a glove, closed at their extremities. There are also many processes which contain a seba- ceous substance that may be pressed out of them in the form of worms: these are the ducts of sebaceous glands. Besides these, there is an immense number of whitish filaments which are as fine as the most delicate thread of a spider's web. These filaments can be best seen while the cuticle is separating from the skin of the sole of the foot, as suggested by Dr. William Hunter.* They are suppo- sed to be vascular, but they have never been injected. When the cuticle is in its natural situation, in union with the skin, there appear to be three species of foramina, or pores, on its external surface: viz. 1. Those formed by the passage of the hairs; and 2. Those which are the orifices of the ducts of the sebaceous glands; each of which has been already mentioned. And 3. Such pores as exist on the ends of the fingers and the inside of the hands. It is sajd that these last are very visible, when magnified to, twice or thrice their original bulk, and drawings of them have accordingly been made by Dr. Grewf and by Mr. Cruikshank.:j: Small specks of fluid can be seen with the na- ked eye, in the same situations, in warm weather, or when the ends of the fingers are made turgid by a ligature. It is * See the London Medical Observations and Inquiries, vol. II. | In the Philosophical Transactions, vol. III. Lowlhrop's Abridgment. \ See his Experiments on Insensible Perspiration. Question respecting the Outlets of Perspiration. 415 probable that they are formed by the accumulation of fluid at these orifices. The above described pores are situated on the ridges at the ends of the fingers and not in the furrows; and it is probable that similar pores are distributed over the surface of the bodj-. Notwithstanding, the appearance of these foramina, when the cuticle is in its natural situation, several of the most successful investigators of the subject have declared that they could not discover any pores or foramina in the cuticle, when it was separated from the cutis. The late professer Meckel of Berlin, who was one of this number, was induced to believe that the matter of ex- halation, and of absorption, soaked through the cuticle, as the vapour of warm water passes through leather.* In support of this doctrine he states, that perspiration goes on through the cuticle on the palms of the hands and soles of the feet when it is very thick; and observes, that if it were transmitted by delicate vessels, the vessels in the feet must be torn by the weight of the body, in persons who walk; and those in the hands would experience the same fate, in labourers, who work with heavy hammers, &c. On the other hand, Mr. Cruikshank, who could likewise find no pores in the separated cuticle, contends strenuous- ly for their existence notwithstanding; and explains their non-appearance by the following facts, among others, viz: that no foramen will appear in the separated cuticle al- though it has been punctured by a needle; and that when the cuticle has been peeled off, from portions of the cutis on which were hairs, which must necessarily have perfora- ted it, no foramina have appeared in it. * Sec Memoirs of the Royal Academy of Sciences of Berlin, vol. XIII. for 175" 416 Peculiar Permeability of the Cuticle. M.-Bichat took very different ground: he asserted that the pores of the separated cuticle were to be seen distinctly, in large numbers, by looking through it towards the light; he also believed that the course of the exhalent vessels, through the cuticle, might be seen in the same manner; and that they passed obliquely. That the cuticle is pervious, is proved incontestably by the functions of perspiration and sweating, as well as of absorption; but there are good reasons for believing that the perforations of the cuticle have a peculiar structure; and are not simple foramina. Thus, when a vesicle is formed by the operation of cantharides or any other pro- cess, if the cuticle is not lacerated, it will confine the effu- sed fluid for a considerable time; without any appearance of its escape through these pores. This fact, which is strongly opposed to the hypothesis of Meckel, is explained by Cruikshank upon the supposi- tion that the pores of the skin are lined by processes of the cuticle, and that when the cuticle ,is separated from the cutis, these processes go with it, and act like valves in con- fining the fluid. Bichat supposes the oblique vessels to produce the same effect upon analogous principles; and compares their situ- ation to that of the ureters, which pass obliquely between the coats of the bladder. This peculiar quality of the cuticle, in admitting of per- spiration and sweat, and also absorption, while it prevents evaporation from the parts which it incloses, is of immense importance. If a portion of skin be deprived of cuticle a short time before death, by a blister for example, this portion will in a few days become perfectly dry and hard, like horn; Causes which produce Vesication. 417 While the other parts of the skin of the subject, covered by the cuticle, retain their moisture and flexibility. It may therefore be admitted, that the use of the cuticle is to keep the skin soft and flexible, by confining its mois- ture, as well as to defend it. And it is probable that the sebaceous matter is secreted for the purpose of preserving the cuticle in a state of flexibility. As the cuticle is capable of confining fluid, and resisting the action of chemical agents, it is surprising that epispas- ties and rubefacients should act through it, upon the skin, with so much certainty as we find they do; and that can- tharides should produce vesications, when applied dry. The thickness of the cuticle on every part of the body- is much increased by long continued pressure, forming corns and excrescences of its own nature. By this cause also it is rendered very thick on the palms of the hands and soles of the feet; although it is originally thicker there than in other parts. It is said that, after long boiling, these thick portions of cuticle may be separated into distinct laminae. In the living subject, the cuticle, when immersed in warm water, seems to absorb some of that fluid; as is evinced by the hands when they have been long in that situation; and also by those parts of the skin to which poul- tices have been applied. Notwithstanding the uniform adhesion of the cuticle to the cutis, it is observed, in the living subject, to be separa- ted, and formed into vesicles, by a variety of causes, viz. 1. Pinching of the skin, or violent mechanical irritation' such as labouring with hard instruments. 2. By the application of cantharides, and certain other substances which produce vesications. Sometimes these substances appear to inflame the skin; but on other occa- Vol. I. 3 G 418 Separation of the Cuticle. sions the vesication is produced while the skin appears unchanged in colour, and free from inflammation. The pro- cess appears different from that of simple inflammation; for certain rubefacients often inflame the skin considera- bly, without vesicating or blistering it. 3. The boiling heat will very generally produce vesica- tion. 4. Certain diseased processes seem to occasion vesica- tion in a manner which is not well understood, viz. ery- sipelas, zona or shingles, pemphigus, and some other eruptions which have no name. In erysipelas thefe is an obvious inflammation of the skin; but in some of the other diseases the vesication takes place without the appearance of inflammation. 5. Vesications often appear when there is a tendency to gangrene. 6. They also occur in some cases of simple fracture, where there is considerable injury. In these cases the fluid effused is often tinged with blood. After death the cuticle is separated from the cutis: 1. By putrefaction; in which case large vesicles are sometimes formed. 2. By long continued maceration. 3. By boiling, and 4. By violent dry heat. The cuticle appears to be least deranged when it is sepa- rated by putrefaction and maceration: in these cases the internal surface corresponds to the surface of the skin; and the processes which contain the hairs, as well as those which are the ducts of the sebaceous glands, are particu- larly obvious. The external surface of the cuticle varies in different places, according to the surface of the skin. In some places Chemical Qualities of the Cuticle. 419 it appears scaly at times, and has therefore been supposed to consist entirely of scales; but in other parts, when ex- amined attentively, it appears like a half transparent con- creted substance, with a rough surface. When the skin has continued dry for a long time, bran- like scales can be rubbed off from it. These are probably composed of the residuum of the secretion deposited on the skin, and of a portion of the external surface of the cuticle. The same substance appears upon the first wash- ing of the skin, after that process has been discontinued for any length of time. Many speculations have arisen respecting the manner in which the cuticle is originally formed, and reproduced; but none of these are perfectly satisfactory. It is also a question whether the cuticle is endued with vitality, or is merely an inanimate unorganized concrete. No decisive arguments have been adduced in favour of its vitality; and it has already been stated, that neither nerves, nor vessels, can be demonstrated in it. It appears particularly calculated for protecting the skin which it covers; for it is insoluble in water, and resists the action of several powerful chemical agents. Thus, it is not affected by immersion for a considerable time in the sul- phuric and muriatic acids; although the nitric acid acts upon it. It resists for a short time, but is at length dissolved, by the pure fixed alkalies, and by lime. It is supposed by the chemists to consist of albumen, in a peculiar state of modification. The Nails. The roots of the nails appear to originate in a fold of the cutis vera, from the epidermis which lines the fold; 420 The Nails. but the bodies of the nails adhere firmly to the cutis on which they lie; and appear to cover it, in the place of the cuticle. The papilla? of those parts of the cutis which are covered by the nails are very conspicuous when the nails are removed. It has been supposed that there was no rete mucosum between the nails and cutis; but this opinion is probably erroneous, as the black pigment is perceptible under the nails of some negroes. The nails can be separated from the cutis by all those processes which separate the cuticle from it. When this is effected, they remain connected with the cuticle, which appears to be continued into them: and on this account, as v/ell as their insensibility, and their resemblance to the horny excrescences of the cuticle, they are considered as appendages of it. The root is opake, and appears white. The body is transparent, and in health shows the florid colour of the cutis which it covers; but the colour of this portion of the 1 cutis depends upon the state of the circulation; and be- comes livid when the blood is disoxygenated, or when the circulation ceases there; and this colour also appears through the nails. The nails are unquestionably organized, although their ultimate structure is not known. They appear to be com- posed of lamellae, and these lamellae, of fibres. They grow rapidly, and when they are not pared or worn away, they sometimes acquire an immense size. As a remarkable instance of this, it is related, that a nail of the great toe was sent from Turin, to the academy of sciences at Paris, which measured four inches and one half, in length. ^ The growth seems to take place altogether at the roots. The Hair. 421 The nails, when chemically examined, appear to consist of a modification of albumen; and thus resemble cuticle and horn, in their composition. The Hairs Originate from bulbs which are situated at the bottom of pores or cavities in the skin. These pores appear to be lined by a production of the cuticle, and the extremities of the bulbs project beyond them into the cellular mem- brane. In some cases, where the cuticle is separated after putrefaction, it seems that these lining processes of the cuticle come away completely, and bring the hairs and their roots with them; but in other cases, the cuticle separates from the cutis, and leaves the hairs in their natural situa- tion. When viewed in a microscope, the bulb appears half transparent, and whitish; and of a softer consistence than the hair itself. The extremity of it is remarkably flexible, and sometimes much darker than the rest of the bulb. The hair does not appear to extend completely to the end of the bulb. Neither bloodvessels nor nerves have been tra- ced to these bulbs, although it is probable they extend there; for the operation of extracting hair by the roots is generally very painful; and blood sometimes appears in the pore, from which the hair is extracted. The body of the hair appears to be composed of smaller fibres, inclosed in a membrane which often is imperfect at the extremity; in consequence of which the fibres often separate from each other, or split. Within the hair is diffused the substance upon which its colour depends: this does not appear to be essential to the strticture, as in the advance of life, the hair is so gene- 422 Vicarious connexion of the Skin. rally without it, while its structure continues unchanged; although it becomes less flexible. The colour of the hair appears to have some connexion with the colour of the rete mucosum, as it is so generally black when the rete mucosum is dark coloured. The sudden change of colour in consequence of fright or grief, is a very rare occurrence indeed; but Bichat re- lates an instance which came under his observation, ia which the hair became perfectly white in one night, in con- sequence of grief. The SKIN, constructed as above described, answers a fourfold purpose in the animal economy. It is the or- gan of touch. It covers and protects the whole struc- ture. It is the outlet for a large proportion of the insen- sible perspiration, and it performs absorption. Many facts have been noticed by practitioners of medi- cine which prove that it has a connexion with the lungs and stomach, which is not yet explained by anatomy. As one of these, an effect of the urticaria or nettle rash may be mentioned. This eruption sometimes relieves completely the spasmodic croup; and in other cases, nausea and vomiting. Some children, when affected with this species of croup, are relieved by rubbing the skin with harsh woollen cloth. In some places the urticaria and the affection of respira- tion are so much regarded as symptoms of the same disease, that the term hives is used as the name for each of them. END OF THE FIRST VOLUME. 110 Wgl7s NATIONAL library of medicine NLM 01001535 a NLM010015358