*v*f-..; '*•- • '?-^v;.' ; .V.'-Vif.'rVV.- ..--.M* ' «▼}!" '¥*< '/^' «■■ HARVARD UNIVERSITY. librarjj of % Pass. HJebhl College. RULES AND REGULATIONS. 1. Students attending- any of the Lectures in the .Massachusetts .Medical College may take books from the Library during- the course, by depositing- Five Dollars with the Dean; and the Students of any of the Medical Professors may have the same privilege on the same terms throughout the year. 2. The Library shall he open on the afternoon of every Saturday, from .'> to 4 o'clock, for the dfiliivirv iiiui l'i^txirn x\t' Lun iL-sj nd 'a^^qrOQrCQ so far as a knowledge of their structure may tend to perfect our ideas of the human body. Introduction to Anatomy. 41 The term anatomy may also have another and more extensive signification: it may be employed to express not only a knowledge of the structure and disposition of the parts but likewise of their oeconomy and use. Consider- ed in this light, it will seldom fail to excite the curiosity of people of taste, as a branch of philosophy; since if it is pleasing to be ac- quainted with the structure of the body, it is certainly more so to discover all the springs which give life and motion to the machine, and to observe the admirable mechanism by which so many different functions are exe- cuted. Astronomy and anatomy, as Dr. Hunter, after Fontenelle, observes, are the studies which present us with the most striking view of the two greatest attributes of the Supreme Being. The first of these fills the mind with the idea of his immensity, in the largeness, distances, and number of the heavenly bodies ; the last, astonishes with his intelligence and art in the variety and delicacy of animal me- chanism. The human body has been commonly enough known by the name of microcosmus, or the little world; as if it did not differ so much from the universal system of nature in the symmetry and number of its parts as in their size. Galen's excellent treatise De usu partium, was composed as a prose hymn to the Crea- tor ; and abounds with as irresistible proofs of a supreme Cause and governing Provi- F 42 Introduction to Anatomy. dence, as we find in modern physico-theology. And Cicero dwells more on the structure and ceconomy of animals than on all the produc- tions of nature besides, when he wants to prove the existence of the gods from the or- der and beauty of the universe. He there takes a survey of the body of man in a most elegant synopsis of anatomy, and concludes thus: " Quibus rebus expositis, satis docuisse videor, hominis natura, quanto omnes anteiret animantes. Ex quo debet intelligi, nee figu- ram situmque membrorum, nee ingenii men- tisque vim talem effici potuisse fortuna." The satisfaction of mind which arises from the study of anatomy, and the influence which it must naturally have upon our minds as phi- losophers, cannot be better conveyed than by the following passage from the same author: " Quse contuens animus, accepit ab his cogni- tionem deorum, ex qua oritur pietas: cui con- juncta justitia est, reliquseque virtutes: ex quibus vita beata exsistit, par et similes deo- rum, nulla alia re nisi immortalitate, quse ni- hil ad bene vivendum pertinet, cedens cceles- tibus." It would be endless to quote the animated passages of this sort which are to be found in the physicians, philosophers, and theologists, who have considered the structure and func- tions of animals with a view towards the Cre- ator. It is a view which must strike one with a most awful conviction. Who can know and consider the thousand evident proofs of the as- tonishing art of the Creator, in forming and Introduction to Anatomy. 43 sustaining an animal body such as ours, with- out feeling the most pleasant enthusiasm ? Can we seriously reflect upon this awful subject, without being almost lost in adoration ? with- out longing for another life after this, in which we may be gratified with the highest enjoy- ment, which our faculties and nature seem ca- pable of, the seeing and comprehending the whole plan of the Creator, in forming the uni- verse and in directing all its operations ? But the more immediate purposes of anato- my concern those who are to be the guardians of health, as this study is necessary to lay a foundation for all the branches of medicine. The more we know of our fabric, the more reason we have to believe, that if our senses were more acute, and our judgment more en- larged, we should be able to trace many springs of life which are now hidden from us: by the same sagacity we should discover the true causes and nature of diseases ; and there- by be enabled to restore the health of many, who are now, from our more confined know- ledge, said to labour under incurable disor- ders. By such an intimate acquaintance with the ceconomy of our bodies, we should disco- ver even the seeds of diseases, and destroy them before they had taken root in the consti- tution. That anatomy is the very basis of surgery every body allows. It is dissection alone that can teach us, where we may cut the living body with freedom and dispatch ; and where we may venture with great circumspection and U Introduction to Anatomy. delicacy; and where we must not, upon any account, attempt it. This informs the head, gives dexterity to the hand, and familiarizes the heart with a sort of necessary inhumanity, the use of cutting-instruments upon our fel- low-creatures. Besides the knowledge of our body, through all the variety of its structure and operations in a sound state, it is by anatomy only that we can arrive at the knowledge of the true nature of most of the diseases which afflict humanity. The symptoms of many disorders are often equivocal; and diseases themselves are thence frequently mistaken, even by sensible, expe- rienced, and attentive physicians. But by ana- tomical examination after death, we can with certainty find out the mistake, and learn to avoid it in any similar case. This use of anatomy has been so generally adopted by the moderns, that the cases alrea- dy published are almost innumerable: Man- ge tus, Morgagni, indeed many of the best mo- dern writings in physic, are full of them. And if we look among the physicians of the best character, and observe those who have the art itself, rather than the craft of the profes- sion at heart; we shall find them constantly taking pains to procure leave to examine the bodies of their patients after death. After having considered the rise and pro- gress of anatomy; the various discoveries that have been made in it, from time to time; the great number of diligent observers who have applied themselves to this art; and the Introduction to Anatomy. 45 importance of the study, not only for the pre- vention and cure of diseases, but in furnish- ing the liveliest proofs of divine wisdom; the following questions seem naturally to arise: For what purpose is there such a variety of parts in the human body ? Why such a com- plication of nice and tender machinery ? Why was there not rather a more simple, less deli- cate, and less expensive frame ?* In order to acquire a satisfactory general idea of this subject, and find a solution of all such questions, let us, in our imaginations, make a man: in other words let us suppose that the mind, or immaterial part, is to be placed in a corporeal fabric, in order to hold a correspondence with other material beings by the intervention of the body; and then consider, a priori, what will be wanted for her accommodation. In this inquiry, we shall plainly see the necessity or advantage, and therefore the final cause, of most of the parts which we actually find in the human body. And if we consider that, in order to answer some of the requisites, human wit and inven- tion would be very insufficient; we need not be surprised if we meet with some parts of the body whose use we cannot yet perceive, and with some operations and functions which we cannot explain. We can see that the whole bears the most striking characters of excelling wisdom and ingenuity: but the imperfect sen- * The following beautiful representation is taken from the late Dr. Hunter's Introductory Lecture in Anatomy. 46 Introduction to Anatomy. ses and capacity of man cannot pretend to reach every part of a machine, which nothing less than the intelligence and power of the Supreme Being could contrive and execute. First, then, the mind, the thinking, imma- terial agent, must be provided with a place of immediate residence, which shall have all the requisites for the union of spirit and body ; accordingly she is provided with the brain, where she dwells as governor and superintend- ant of the whole fabric. In the next place, as she is to hold a corre- spondence with all the material beings around her, she must be supplied with organs fitted to receive the different kinds of impressions which they will make. In fact, therefore, we see that she is provided with the organs of sense, as we call them : the eye is adapted to light; the ear to sound ; the nose to smell; the mouth to taste; and the skin to touch. Further: She must be furnished with or- gans of communication between herself in the brain and those organs of sense, to give her information of all the impressions that are made upon them: and she must have organs between herself in the brain and every other part of the body, fitted to convey her com- mands and influence over the whole. For these purposes the nerves are actually given. They are chords, which rise from the brain, the immediate residence of the mind, and dis- perse themselves in branches through all parts of the body. They convey all the different kinds of sensations to the mind, in the brain; Introduction to Anatomy, 47 and likewise carry out from thence all her com- mands or influence to the other parts of the body. They are intended to be occasional mo- nitors against all such impressions as might en- danger the well-being of the whole, or of any particular part; which vindicates the Creator of all things, in having actually subjected us to those many disagreeable and painful sensa- tions which we are exposed to from a thousand accidents in life. Moreover, the mind, in this corporeal sys- tem, must be endued with the power of mov- ing from place to place, that she may have in- tercourse with a variety of objects ; that she may fly from such as are disagreeable, danger- ous or hurtful, and pursue such as are plea- sant or useful to her. And accordingly she is furnished with limbs, and with muscles and tendons, the instruments of motion, which are found in every part of the fabric where motion is necessary. But to support, to give firmness and shape to the fabric ; to keep the softer parts in their proper.places ; to give fixed points for, and the proper direction to its motions, as well as to protect some of the more important and tender organs from external injuries; there must be some firm prop-work interwoven through the whole. And, in fact, for such purposes the bones are given. The prop-work must not be made into one rigid fabric, for that would prevent motion. Therefore there are a number of bones. 48 Introduction to Anatomy. These pieces must all be firmly bound toge- ther, to prevent their dislocation. And this end is perfectly well answered by the liga- ments. The extremities of these bony pieces, where they move and rub upon one another, must have smooth and slippery surfaces for easy motion. This is most happily provided for, by the cartilages and mucus of the joints. The interstices of all those parts must be filled up with some soft and ductile matter, which shall keep them in their places, unite them, and at the same time allow them to move a little upon one another. And these purposes are answered by the cellular membrane or adi- pose substance. There must be an outward covering over the whole apparatus, both to give it compact- ness and to defend it from a thousand injuries: which, in fact, are the very purposes of the skin and other integuments. Lastly, the mind being formed for society and intercourse with beings of her own kind, she must be endued with powers of expressing and communicating her thoughts by some sen- sible marks or signs ; which shall be both easy to herself, and admit of great variety; and ac- cordingly she is provided with the organs and faculty of speech, by which she can throw out signs with amazing facility, and vary them with- out end. Thus we have built up an animal body which would seem to be pretty complete : but as it is the nature of matter to be altered and work- Introduction to Anatomy. 49 ed upon by matter; so in a very little time such a living creature must be destroyed, if there is no provision for repairing the injuries which she must commit upon herself, and those which she must be exposed to from without. There- fore a treasure of blood is actually provided in the heart and vascular system, full of nutritious and healing particles, fluid enough to penetrate into the minutest parts of the animal; impelled by the heart, and conveyed by the arteries, it washes every part, builds up what was broken down, and sweeps away the old and useless materials. Hence we see the necessity or ad- vantage of the heart and arterial system. What more there was of this blood than enough to repair the present damages of the machine, must not be lost, but should be re- turned again to the heart; and for this purpose the venous system is actually provided. These requisites in the animal explain, a priori, the circulation of the blood. The old materials which were become use- less, and are swept off by the current of blood, must be separated and thrown out of the sys- tem. Therefore glands, the organs of Secre- tion, are given for straining whatever is redun- dant, vapid, or noxious, from the mass of blood; and when strained, they are thrown out by emunctories, called organs of Excre- tion. But now, as the machine must be constant- ly wearing, the reparation must be carried on without intermission, and the strainers must always be employed. Therefore there is ac- G 50 Introduction to Anatomy. tually a perpetual circulation of the blood, and the secretions are always going on. Even all this provision, however, would not be sufficient; for that store of blood would soon be consumed, and the fabric would break down, if there were not a provision made for fresh supplies. These we observe, in fact, are profusely scattered round her in the animal and vegetable kingdoms ; and she is furnished with hands, the fittest instruments that could have been contrived, for gathering them, and for preparing them in a variety of ways for the mouth. But these supplies, which we call food, must be considerably changed; they must be converted into blood. Therefore she is pro- vided with teeth for cutting and bruising the food, and with a stomach for melting it down: In short, with all the organs subservient to di- gestion.—The finer parts of the aliments only can be useful in the constitution: these must be taken up and conveyed into the blood, and the dregs must be thrown off. With this view the intestinal canal is actually given. It sepa- rates the nutritious part, which we call chyle, to be conveyed into the blood by the system of absorbent vessels; and the fseces pass down- wards, to be conducted out of the body. Now we have got our animal not only fur- nished with what is wanted for its immediate existence, but also with the powers of protract- ing that existence to an indefinite length of time. But its duration, we may presume, must necessarily be limited: for as it is nourished, Introduction to Anatomy. 51 grows, and is raised up to its full strength and utmost perfection ; so it must, in time, in com- mon with all material beings, begin to decay, and then hurry on to final ruin. Hence we see the necessity of a scheme for renovation. Accordingly wise Providence, to perpetuate, as well as preserve his work, besides giving a strong appetite for life and self-preservation, has made animals male and female, and given them such organs and passions as will secure the propagation of the species to the end of time. Thus we see, that by the very imperfect survey which human reason is able to take of this subject, the animal man must necessarily be complex in his corporeal system, and in its operations. He must have one great and general system, the vascular, branching through the whole for circulation: Another the nervous, with its ap- pendages the organs of sense, for every kind of feeling: And a third, for the union and connection of all those parts. Besides these primary and general systems, he requires others which may be more local, or confined: One for strength, support, and protection; the bony compages : Another for the requisite motions of the parts among them- selves, as well as for moving from place to place ; the muscular part of the body: Ano- ther to prepare nourishment for the daily re- cruit of the body; the digestive organs : And one for propagating the species; the organs of generation. 52 Introduction to Anatomy, And in taking this general survey of what would appear, a priori, to be necessary for ' adapting an animal to the situations of life, we observe with great satisfaction, that man is accordingly made of such systems, and for ' such purposes. He has them all; and he has nothing more, except the organs of respira- tion. Breatning it seemed difficult to account for a priori: we only knew it to be in fact essentially necessary to life. Notwithstanding this, when we saw all the other parts of the bo- dy, and their functions, so well accounted for, and so wisely adapted to their several purpo- ses, there could be no doubt that respiration was so likewise: And accordingly, the disco- veries of Dr. Priestley have lately thrown light upon this function also, as will be shown in its proper place. Of all the different systems in the human body, the use and necessity are not more ap- parent, than the wisdom and contrivance which has been exerted in putting them all into the most compact and convenient form : in dispos- ing them so, that they shall mutually receive, and give helps to one another; and that all, or many of the parts, shall not only answer their principal end or purpose, but operate success- fully and usefully in a variety of secondary ways. If we consider the whole animal machine in this light, and compare it with any machine in which human art has exerted its utmost; suppose the best constructed ship that ever was built, we shall be convinced beyond the Introduction to Anatomy. S3 possibility of doubt, that there are intelligence and power far surpassing what humanity can boast of. One superiority in the natural machine is peculiarly striking.—In machines of human contrivance or art, there is no internal power, no principle in the machine itself, by which it can alter and accommodate itself to any injury which it may suffer, or make up any injury which admits of repair. But in the natural machine, the animal body, this is most won- derfully provided for, by internal powers in the machine itself; many of which are not more certain and obvious in their effects, than they are above all human comprehension as to the manner and means of their operation. Thus, a wound heals up of itself; a broken bone is made firm again by a callus ; a dead part is separated and thrown off; noxious juic- es are driven out by some of the emunctories; a redundancy is removed by some spontane- ous bleeding ; a bleeding naturally stops of it- self; and a great loss of blood, from any cause, is in some measure compensated, by a con- tracting power in the vascular system, which accommodates the capacity of the vessels to the quantity contained. The stomach gives information when the supplies have been ex- pended ; represents, with great exactness, the quantity and the quality of what is wanted in the present state of the machine ; and in pro- portion as she meets with neglect, rises in her demand, urges her petition in a louder tone, and with more forcible arguments. For its 54 Introduction to Anatotny.. protection, an animal body resists heat and cold in a very wonderful manner, and pre- serves an equal temperature in a burning and in a freezing atmosphere. A farther excellence or superiority in the natural machine, if possible, still more aston- ishing, more beyond all human comprehen- sion, than what we have been speaking of, is the following. Besides those internal powers of self-preservation in each individual, when two of them co-operate, or act in concert, they are endued with powers of making other ani- mals or machines like themselves, which again are possessed of the same powers of produc- ing others, and so of multiplying the species without end. These are powers which mock all human in- vention or imitation. They are characteris- tics of the divine Architect. Having premised this general account of the subject, we shall next consider the method to be observed in treating it. The study of the human body, as already noticed, is commonly divided into two parts. The first, which is called Anatomy, relates to the matter and structure of its parts; the se- cond, called Physiology and Animal ozconomy, relates to the principles and laws of its inter- nal operations and functions. As the body is a compound of solids and fluids, Anatomy is divided into, 1. The Anatomy of the solids, and 2. The Anatomy of the fluids. Introduction to Anatomy. 55 I. The Solids, by which we mean all parts of our body, which are not fluid, are general- ly divided into two classes, viz. 1. The hard solids or bones. This part of anatomy is called Osteology; which signifies the doctrine of the bones. 2. The softer solids; which part is called Sarcology, Viz. the doctrine of flesh. This division of the solids, we may observe, has probably taken its origin from the vulgar observation, that the body is made of bone and flesh. And as there are many different kinds of what are called soft or fleshy parts, Sarcology is subdivided into, (l.) Angeiology, or the doctrine of vessels; by which is commonly understood blood-vessels : (2.) Adenology, of glands: (3.) Neurology, of nerves : (4.) Myology, of muscles : and, (5.) Splanchnology, of the viscera or bow- els. There is, besides, that part which treats of the organs of sense and of the integuments. This division of the solids has been here mentioned, rather for the sake of explaining so many words, which are constantly used by anatomists, than for its importance or ac- curacy. For besides many other objections that might be urged, there are in the body three species of solids, viz. gristle or carti- lage, hair, and nails ; which are of an inter- mediate nature between bone and flesh; and therefore cannot so properly be brought into the osteology or the sarcology. The cartila- ges were classed with the bones ; because the 56 Introduction to Anatomy. greatest number of them are appendages to bones : and for the like reason the hair and the nails were classed with the integuments. II. The Fluids of the human body may be divided into three kinds, which Dr. Hunter calls the crude, the general or perfect, and the local or secreted fluid. 1. By the crude fluid is meant the chyle, and whatever is absorbed at the surfaces of the body; in other words, what is recently taken into the body, and is not yet mixed with or converted into blood. 2. The general or perfect fluid is the blood itself; to wit, what is contained in the heart, arteries, and veins, and is going on in the round of the circulation. 3. The local or secreted, are those fluids pe- culiar to particular parts of the body, which are strained off from the blood, and yet are very different in their properties from the blood. They are commonly called secretions; and some are useful, others excrementitious. In treating of the Physiology, it is very dif- ficult to say what plan should be followed; for every method which has been yet proposed, is attended with manifest inconvenience. The powers and operations of the machine have such a dependence upon one another, such connections and reciprocal influence, that they cannot well be understood or explained sepa- rately. In this sense our body may be com- pared to a circular chain of powers, in which nothing is first or last, nothing solitary or in- dependent ; so that wherever we begin, we Osteology, 57 find that there is something preceding which we ought to have known. If we begin with the brain and the nerves, for example, we shall find that these cannot exist, even in idea, without the heart: if we set out with the heart and vascular system, we shall presently be sensible, that the brain and nerves must be supposed: or, should we take up the mouth, and follow the course of the aliment, we should see that the very first organ which presents itself, supposed the existence of both the heart and brain: Wherefore we shall incorporate the Physiology with the Anatomy, by attempting to explain the functions after we have demon- strated the organs. Part I. OSTEOLOGY. E begin with the bones, which may be considered as the great support of the body, tending to give it shape and firmness.— But before we enter into the detail of each par- ticular bone, it will be necessary to describe their composition and connections, and to ex- plain the nature of the different parts which have an immediate relation to them; as the car- tilages, ligaments, periosteum, marrow, and synovial glands. H w 58 Osteology. Sect. I. Of the Bones in general, with their Appendages, &fc. The bones are of a firm and hard* sub- stance, of a white colour, and perfectly insen- sible. They are the most compact and solid parts of the body, and serve for the attachment and support of all the other parts. Three different substances are usually dis- tinguished in them; their exterior or bony part, properly so called; their spongy cells; and their reticular substance. The first of these is formed of many laminae or plates, compos- ing a firm hard substance—The spongy or cellular part is so called on account of its re- semblance to a sponge, from the little cells which compose it. This substance forms al- most the whole of the extremities of cylindrical bones. The reticular part is composed of fi- bres, which cross each other in different direc- tions. This net-work forms the internal sur- face of those bones which have cavities. The flat bones, as those of the head, are composed only of the laminae and the cellular substance. This last is usually found in the middle of the bone dividing it into two plates, and is there called diploe. * Mr. Scheele has lately difcovered that bones contain the phofphoric acid united with calcareous earth; and that to this combination they owe their firmnefs. Osteology. 59 Gagliardi, who pretended to have discover- ed an infinite number of claviculi* or bony processes, which he describes as traversing the laminae to unite them together, has endea- voured to support this pretended discovery by the analogy of bones to the bark of trees, in which certain woody nails have been remark- ed; but this opinion seems to be altogether fanciful. Some writers have supposed, that the bones are formed by layers of the perioste- um, which gradually ossify, in the same man- ner as the timber is formed in trees by the hardening of the white substance that is found between the inner bark and the wood. M. Duhamel, who has adopted this opinion, fed different animals with madder and their ordi- nary food alternately during a certain time; and he asserts, that in dissecting their bones, he constantly observed distinct layers of red and white, which corresponded with the length of time they had lived on madder or their usual aliment. But it has since been proved by Detleff, that M. Duhamel's experiments were inaccurate, and that neither the perios- teum nor the cartilages are tinged by the use of madder, which is known to affect the bones only. We usually consider in a bone, its body and its extremities. The ancients gave the * In his Anat. Ojfium tiov. invent, dlujlrat. he defcribes four kinds of thefe claviculi or nails, viz. the perpendicular, oblique, headed, and crooked. 60 Osteology. name of diaphysis to the body or middle part, and divided the extremities into apophysis and epiphysis. An apophysis, or process, as it is more commonly called, is an eminence con- tinued from the body of the bone, whereas an epiphysis is at first a sort of appendage to the bone, by means of an intermediate cartilage. Many epiphyses, which appear as distinct bones in tue foetus, afterwards become apophy- ses ; for they are at length so completely unit- ed to the body of the bone as not to be distin- guishable from it in the adult state. It is not unusual, however, at the age of 18 and even 20 years, to find the extremities of bones still in the state of epiphysis. The names given to the processes of bones are expressive of their shape, size, or use; thus if a process is large and of a spherical form, it is called caput, or head; if the head is flatted, it is termed condyle. Some proces- ses, from their resemblance to a stiletto, a breast, or the beak of a crow, are called styloid, mastoid, or coracoid: others are styled ridges or spines. The two processes of the os femo- ris derive their name of trochanters from their use. A bone has its cavities as well as processes. These cavities either extend quite through its substance, or appear only as depressions. The former are called foramina or holes, and theso foramina are sometimes termed canals or con- duits, according to their form and extent. Of the depressions, some are useful in articula- tion. These are called cotyloid when they are Osteology, 61 deep, as is the case with the os innominatum, where it receives the head of the os femoris; or glenoid when they are superficial, as in the scapula, where it receives the os humeri. Of the depressions that are not designed for arti- culation, those which have small apertures are called sinuses; others that are large, and not equally surrounded by high brims, are styled fosse; such as are long and narrow,furrows; or if broad and superficial without brims, si- nuosities. Some are called digital impressions, from their resemblance to the traces of a fin- ger on soft bodies. We shall abridge this article, which is ex- ceedingly diffuse in the generality of anatomi- cal books, and will endeavour to describe it with all the clearness it will allow. The bones composing the skeleton are so constructed, that the end of every bone is per- fectly adapted to the extremity of that with which it is connected, and this connection forms what is called their articulation. Articulation is divided into diarthrosis, sy- narthrosis, and amphiarthrosis, or moveable, immovable, and mixed articulation. Each of the two first has its subdivisions. Thus the Diarthrosis, or moveable articulation, includes, 1. The enarthrosis, as it is called, when a large head is admitted into a deep cavity, as in the articulation of the os femoris with the os innominatum. 2. Arthrodia, when a round head is articulated with a superficial cavity, as is the case of the os humeri and scapula. 3. Ginglimus, or hinge-like articulation, as in the 62 Osteology. connection of the thigh-bone with the tibia. The enarthrosis and arthrodia allow of motion to all sides ; the ginglimus only of flection and extension. The syfiarthrosis, or immoveable articula- tion, includes, 1. The suture, when the two bones are indented into each other, as is the case with the parietal bones. 2. Gomphosis, when one bone is fixed into another, in the manner the teeth are placed in their sockets. The term amphiarthrosis is applied to those articulations which partake both of the synar- throsis and diarthrosis, as is the case with the bones of the vertebrae, which are capable of motion in a certain degree, although they are firmly connected together by intermediate car- tilages. What is called symphysis is the union of two bones into one; as in the lower jaw, for instance, which in the foetus consists of two distinct bones, but becomes one in a more ad- vanced age, by the ossification of the uniting cartilage. When bones are thus joined by the means of cartilages, the union is styled synchondro- sis ; when by ligaments, syneurosis. Cartilages are white, solid, smooth, and elastic substances, between the hardness of bones and ligaments, and seemingly of a fi- brous texture. We are not able to trace any vessels into their substance by injection, nor are they ever found tinged in animals that have been fed with madder. Osteology. 63 They may be distinguished into, 1st, Those which are connected with the bones; and, 2dly, Those" which belong to other parts of the body. The first serve either to cover the ends and cavities of bones intended for mo- tion, as in the articulations, where by their smoothness they facilitate motions, which the bones alone could not execute with so much freedom; or they serve to unite bones toge- ther, as in the symphysis pubis, or to length- en them, as in the ribs. Many of them ossifying as we advance in life, their number is less in the adult than in the foetus, and of course there are fewer bones in the old than in the young subject. Of the second class of cartilages, or those belonging to the soft parts, we have instances in the larynx, where we find them useful in the formation of the voice, and for the attach- ment of muscles. The periosteum is a fine membrane of a compact cellular texture, reflected from one joint to another, and serving as a common covering to the bones. It has sanguiferous and lymphatic vessels, and is supplied with nerves from the neighbouring parts. It ad- heres very firmly to their surface, and by its smoothness facilitates the motion of muscles. It likewise supports the vessels that go to be distributed through the substance of the bones, and may serve to strengthen the articulations. At the extremities of bones, where it is found covering a cartilage, it has by some been im- properly considered as a distinct membrane, 64 Osteology. and named perichondrium. This, in its use and structure, resembles the periosteum. Where it covers the bones of the skull, it has gotten the name of pericranium. The periosteum is not a production of the dura mater, as the ancients, and after them Havers, imagined; nor are the bones formed by the ossification of this membrane, at least when it is in a sound state, as some late wri- ters have supposed. The periosteum is deficient in the teeth above the sockets, and in those parts of bones to which ligaments or tendons are attached. The marrow is a fat oily substance, filling the cavities of bones. In the great cavities of long bones it is of a much firmer consistence than in the cells of their spongy part. In the former it inclines somewhat to a yellowish tinge, and is of the consistence of fat; in the latter it is more fluid, and of a red colour. This difference in colour and consistence is owing to accidental causes; both kinds are of the same nature, and may both be described under the common name of marrow, though some writers give the name only to the fat-like substance, and call the other the medullary juice. The marrow is contained in a very fine and transparent membrane, which is supplied with a great number of blood-vessels, chiefly from the periosteum. This membrana medullaris adheres to the inner surface of the bones, and furnishes an infinite number of minute bags or vesicles for inclosing the marrow, Osteology. 65 which is likewise supported in the cavities of the bones by the long filaments of their reti- cular substance. Besides the vessels from the periosteum, the membrana medullaris is furnished with others, which in the long bones may be seen passing in near the extremities of the bone, and sending off numerous branches that rami- fy through all the vesicles of this membrane. The bones, and the cells containing the marrow, are likewise furnished with lympha- tics. By their means, the marrow, like the fat, may be taken up in a greater quantity than it is secreted; and hence it is that so little is found in the bones of those who die of linger- ing diseases. It is still a matter of controversy, Whether the marrow is sensible or not? We are cer- tainly not able to trace any nerves to it; and from this circumstance, and its analogy to fat, Haller has ventured to consider it as insensi- ble. On the other hand, Duverney asserts, that an injury done to this substance in a liv- ing animal was attended with great pain. In this dispute physiologists do not seem to have sufficiently discriminated between the marrow itself and the membranous cells in which it is contained. The former, like the fat, being nothing more than a secreted, and of course an inorganized, matter, may with propriety be ranked among the insensible parts, as much as inspissated mucus or any other secreted mat- ter in the body; whereas the membrana me- dullaris being vascular, though it possesses but I 66 Osteology. an obscure degree of feeling in a sound state, is not perfectly insensible. The marrow was formerly supposed to be intended for the nourishment and renewal of the bones; hut this doctrine is now pretty ge- nerally and deservedly exploded. It seems probable that the marrow is to the bones what fat is to the soft parts. They both serve for some important purposes in the animal cecono- my; but their particular use has never yet been clearly ascertained. The marrow, from the transudation of the oil through the bones of a skeleton, is supposed to diminish their brittleness ; and Havers, who has written pro- fessedly on the bones, describes the canals by which the marrow is conveyed through every part of their substance, and divides them into longitudinal and transverse ones. He speaks of the first as extending through the whole length of the bone; and of the latter, as the passages by which the longitudinal ones com- municate with each other. The similarity of these to the large cancelli in burnt bones, and the transudation of the oil through the bones of the skeleton, seems to prove that some such passages do actually exist. The synovial glands are small bodies,* sup- posed to be of a glandular structure, and ex- ceedingly vascular, secreting a fluid of a clear mucilaginous nature, which serves to lubricate the joints. They are placed in small cavities * It is now much doubted, however, whether the appear- ances in the joints, which are usually called glands, are any thing more than assemblages of fat. Osteology. 67 in the articulations, so as to be capable of be- ing gently compressed by the motion of the joint, which expresses their juice in propor- tion to the degree of friction. When the sy- novia is wanting, or is of too thick a consist- ence, the joint becomes stiff and incapable of flexion or extension. This is what is termed anchilosis. Ligaments are white, glistening, inelastic bands, of a compact substance, more or less broad or thick, and serving to connect the bones together. They are distinguished by different names adapted to their different forms and uses. Those of the joints are called ei- ther round or bursal. The round ligaments are white, tendinous, and inelastic. They are strong and flexible, and are found only in the joint of the knee, and in the articulation of the os femoris with the os innominatum. The bursal, or capsular ligaments, surround the whole joint like a purse, and are to be found in the articulations which allow motion every way, as in the articulation of the arm with the scapula. Of those sacs called Burse mucosa, a few were known to former anatomists, but by much the greater number have been since discover- ed by Dr. Monro,* who observes, that they are to be met with in the extremities of the body only; that many of them are placed en- tirely on the inner sides of the tendons, be- tween these and the bones. Many others co- ver not only the inner, but the outer sides of * See Descripion of the Burs* Mucosa, kc. 68 Osteology. the tendons, or are interposed between the ten- dons and external parts, as well as between those and the bones. Some are situated between the tendons and external parts only or chiefly, some between contiguous tendons, or between the tendons or the ligaments and the joints. A few such sacs are observed where the processes of bones play upon the ligaments, or where one bone plays upon another. Where two or more ten- dons are contiguous, and afterwards separate from each other, we generally find a common bursa divided into branches, with which it communicates ; and a few bursae of contigu- ous tendons communicate with each other.— Some in healthy children, communicate with the cavities of the joints ; and in many old people he has seen such communications form- ed by use or worn by friction, independent of disease. Their proper membrane is thin and trans- parent, but very dense, and capable of con- fining air or any other fluid. It is joined to the neighbouring parts by the common cellular sub- stance. Between the bursa and the hard sub- stance of bone, a thin layer of cartilage or of tough membrane is very generally interpos- ed. To the cellular substance on the outside of the bursa, the adipose substance is con- nected; except where the bursa covers a ten- don, cartilage, or bone, much exposed to pres- sure or friction. In several places a mass of fat, covered with the continuation of the membrane of the Osteology. 69 bursa, projects into its cavity. The edges of this are divided into fringes. The inner side of the membrane is smooth, and is extremely slippery from the liquor se- creted in it. The structure of the bursae bears a strong resemblance to the capsular ligaments of the joints. 1. The inner layer of the ligament, like that of the bursae, is thin and dense. 2. It is connected to the external ligaments by the common cellular substance. 3. Between it and the bones, layers of cartilage, or the articular cartilages, are interposed. 4. At the sides of the joints, where it is not subjected to violent pressure and friction, the adipose substance is connected with the cellular mem- brane. 5. Within the cavities of the joints we observe masses of fat projecting, covered with similar blood-vessels, and with similar fimbriae hanging from their edges. 6. In the knee the upper part of such a mass of fat forms what has been called the mucilaginous gland of the joint, and the under part projects into the bursa behind the ligament which ties the patella to the tibia. 7. The liquor which lubricates the bursae has the same colour, con- sistence, and properties as that of the joints, and both are affected in the same manner by heat, mineral acids, and ardent spirits. 8. In some places the bursae constantly communi- cate with the cavities of the joints, in others they generally do so ; from which we may in- fer a sameness of structure. When we examine the fimbriae common to the fatty bodies of the joints and bursa?, and 70 Osteology. which have been supposed to be the ducts of glands lodged within the masses of fat, we are not able to discover any glandular appear- ance within them. And although we observe many vessels dispersed upon the membranes of the fatty bodies and fimbriae ; and that we cannot doubt that these fimbriae consist of ducts which contain a lubricating liquor, and can even press such a liquor from them; yet their cavities and orifices are so minute, that they are not discoverable even by the assistance of magnifying-glasses. These fimbriae appear, therefore, to be ducts like those of the ure- thra, which prepare a mucilaginous liquor, without the assistance of any knotty or glan- dular organ. Upon the whole, the synovia seems to be furnished by invisible exhalent arteries by the ducts of the fimbriae, and by oil exuding from the adipose follicles by passages not yet disco- vered. The word skeleton, which by its etymology implies simply a dry preparation, is usually applied to an assemblage of all the bones of an animal united together in their natural or- der. It is said to be a natural skeleton, when the bones are connected together by their own proper ligaments; and an artificial one, when they are joined by any other substance, as wire, 8*c. The skeleton is generally divided into the :s head, trunk, and extremities. The first divi- "% sion includes the bones of the cranium and face. The bones of the trunk are the spine, ribs, sternum, and bones of the pelvis. Osteology. 71 The upper extremity on each side consists of the two bones of the shoulder, viz. the sca- pula and clavicle ; the bone of the arm, or os humeri; the bones of the fore-arm, and those of the hand. The lower extremity on each side of the trunk consists of the thigh-bone and the bones of the leg and foot. Sect. II. Of the Bones of the Head. The head is of a roundish figure, and some- what oval.* Its greatest diameter is from the forehead to the occiput; its upper part is call- ed vertex, or crown of the head; its anterior or fore-part the face ; and the upper part of this sinciput, or forehead; its sides the tem- ples ; its posterior, or hind-part, the occiput; and its inferior part the basis. The bones of the head may be divided into those of the cranium and face. * The bones of the foetus being perfectly distinct, and the muscles in young persons not acting much, the shape of the head has been supposed to depend much on the management of children when very young. Vesalius, who has remarked the dif- ference in people of different nations, observes, for instance, that the head of a Turk is conical, from the early use of the turban ; whilst that of an Englishman is flattened by the chin-stay. Some of the latest physiologists suppose, with good reason, that this difference is chiefly owing to certain natural causes with which we are as yet unacquainted. 7 2 Osteology. 1. ito;z£'s of the Cranium and Face* There are eight bones of the cranium, viz. the coronal bone, or os frontis; the two pari- etal bones, or ossa bregmatis; the os occipi- tis; the two temporal bones; the sphenoid bone; and the os ethmoides, or cribriforme. Of these, only the os occipitis and ossa bregmatis are considered as proper to the cra- nium ; the rest being common both to the cra- nium and face. These bones are all harder at their surface than in their middle; and on this account they are divided into two tables, and a middle spon- gy substance called diplb'e. In this, as in all the other bones, we shall consider its figure, structure, processes, de- pressions, and cavities; and the manner in which it is articulated with the other bones. The os frontis has some resemblance in shape to the shell of the cockle. Externally it is convex, its concave side being turned to- wards the brain. This bone, in the places where it is united to the temporal bones, is very thin, and has there no diploe. It is like- wise exceedingly thin in that part of the orbit of the eye which is nearest to the nose. Hence it is, that a wound in the eye, by a sword or any other pointed instrument, is sometimes productive of immediate death. In these cases, the sword passing through the weak part of the bone, penetrates the brain, and divides the nerves at their origin; or perhaps opens Osteology. 73 some blood-vessel, the consequences of which are soon fatal. We observe on the exterior surface of this bone five apophyses or processes, which are easily to be distinguished. One of these is placed at the bottom and narrowest part of the bone, and is called the nasal process, from its supporting the upper end of the bones of the nose. The four others are called angular or orbitar processes. They assist to form the orbits, which are the cavities on which the eyes are placed. In each of these orbits there are two processes, one at the interior or great angle, and the other at the exterior or little angle of the orbit. They are called the angu- lar processes. Between these a ridge is ex- tended in form of an arch, and on this the eye- brows are placed. It is called the orbitar or superciliary ridge, and in some measure covers and defends the globe of the eye. There is a hole in this for the passage of the frontal ves- sels and nerves. This arch is interrupted near the nose by a small pit, in which the ten- don of the musculus obliquus major of the eye is fixed. From the under part of each su- perciliary ridge a thin plate runs a considera- ble way backwards, and has the name of orbi- tar ; the external and fore-part of this plate forms a sinuosity for lodging the lacrymal gland. Between the orbitar plates there is a large discontinuation of the bone, which is fill- ed up by the cribriform part of the os ethmoi- des. K 74 Osteology. On examining the inner surface of this bone at its under and middle part, we observe an elevation in form of a ridge, which has been called the spinous process ; it ascends for some way, dividing the bone into two considerable fossae, in which the anterior lobes of the brain are placed. To a narrow furrow in this ridge is attached the extremity of the falx, as the membrane is called, which divides the brain into two hemispheres. The furrow becoming gradually wider, is continued to the upper and back part of the bone. It has the falx fixed to it, and part of the longitudinal sinus lodged in it. Besides the two fossae, there are many depressions, which appear like digital impres- sions, and owe their formation to the promi- nent circumvolutions of the brain. In the foetus, the forehead is composed of two distinct bones ; so that in them the sagit- tal suture reaches from the os occipitis to the nose. This bone is almost every where com- posed of two tables and a diploe. These two tables separating from each other under the eyes, form two cavities, one on each side of the face, called the frontal sinuses. These sinuses are lined with a soft membrane, called membrana pituitaria. In these sinuses a mu- cus is secreted, which is constantly passing through two small holes into the nostrils, which it serves to moisten. The os frontis is joined by suture to many of the bones of the head, viz. to the parietal, maxillary, and temporal bones ; to the os eth- moides; os sphenoides; os unguis ; and ossa Osteology. 75 nasi. The suture which connects it with the parietal bones is called the coronal suture. The parietal bones are two in number; they are very thin, and even transparent in some places, the particular figure of each of these bones is that of an irregular square, bordered with indentations through its whole circumfer- ence, except at its lower part. It will be ea- sily conceived, that these bones which com- pose the superior and lateral parts of the cra- nium, and cover the greatest part of the brain, form a kind of vault. On their inner surface we observe the marks of the vessels of the du- ra mater; and at their upper edge the groove for the superior longitudinal sinus. The ossa parietalia are joined to each other by the sagittal suture ; to the os sphenoides and ossa temporum by the squamous suture ; to the os occipitis by the lambdoidal suture,* so called from its resemblance to the Greek letter lambda ; and to the os frontis by the co- ronal suture. In the foetus, the parietal bones are sepa- rated from the middle of the divided os frontis by a portion of the cranium then unossified. The occipital bone forms the posterior and inferior parts of the skull; it approaches near- ly to the shape of a lozenge, and is indented throughout three parts of its circumference. * The lambdoidal suture is sometimes very irregular, being composed of many small sutures, which surround so many lit- tle bones called ossa triquetra, though perhaps improperly, as they are not always triangular. 76 Osteology. There is a considerable hole in the inferior portion of this bone, called the foramen mag- num, through which the medulla oblongata passes into the spine.—The nervi accessorii, and vertebral arteries, likewise pass through it. Behind the condyles are two holes for the passage of cervical veins into the lateral si- nuses ; and above them are two others for the passage of the eighth pair and accessory nerves out of the head. At the sides, and a little on the anterior part of the foramen magnum, are two processes, called the condyles, one on each side; they are of an oval figure, and are covered with cartilage. The external surface of this bone has a large transverse arched ridge, under which the bone is very irregular, where it affords attach- ment to several muscles. On examining its inner surface, we may observe two ridges in form of a cross; one ascending from near the foramen magnum to the top of the bone; the upper end of this in which the falx is fixed, is hollow, for lodging the superior longitudinal sinus, and the under end has the third process of the dura mater fixed to it. The other ridge, which runs horizontally, is likewise hollow for containing the lateral sinuses. Four fossae are formed by the cross, two above and two below. In the former are placed the posterior lobes of the brain, and in the latter the lobes of the cerebellum. At the basis of the cranium, we observe the cuneiform process (which is the name given to the great apophysis at the fore part of Osteology. 77 this bone^; it serves for the reception of the medulla oblongata. The os occipitis is of greater strength and thickness, than either of the other bones of the head, though irregularly so; at its inferi- or part, where it is thinnest, it is covered by a great number of muscles. This bone, from its situation, being more liable to be injured by falls, than any other bone of the head, nature has wisely given it the greatest strength at its upper part, where it is most exposed to danger. It is joined to the parietal bones by the lambdoidal suture, and to the ossa temporum, by the additamentum of the temporal suture. It is likewise connected to the os sphenoides by the cuneiform process. It is by means of the os occipitis that the head is united to the trunk, the two condyles of this bone being connected to the superior oblique processes of the first vertebra of the neck. There are two temporal bones, one on each side.—We may distinguish in them two parts; one of which is called the squamous or scaly part, and the other pars petrosa from its hard- ness. This last is shaped like a pyramid. Each of these divisions affords processes and cavities: externally there are three pro- cesses ; one anterior, called the zygomatic process ; one posterior, called the mastoid or mamillary process, from its resemblance to a nipple ; and one inferior, called the styloid pro- cess, because it is shaped like a stiletto, or dag- ger. 78 Osteology. The cavities are, 1. The meatus auditorius externus. 2. A large fossa which serves for the articulation of the lower jaw ; it is before the meatus auditorius, and immediately under the zygomatic process. 3. The stylo-mastoid hole, so called from its situation between the styloid and mastoid processes; it is likewise styled the aquaeduct of Fallopius, and affords a passage to the portio dura of the auditory, or seventh pair of nerves. 4. Below, and on the fore-part of the last foramen, we observe part of the jugular fossa, in which the begin- ning of the internal jugular vein is lodged. An- terior and superior to this fossa is the orifice of a foramen through which passes the carotid artery. This foramen runs first upwards and then forwards, forming a kind of elbow, and ter- minates at the end of the os petrosum.—At this part of each temppral bone, we may observe the opening of the Eustachian tube, a canal which passes from the ear to the back part of the nose. In examining the internal surface of these bones, we may remark the triangular figure of their petrous part which separates two fos- sae ; one superior and anterior ; the other in- ferior and posterior: the latter of these com- poses part of the fossa, in which the cerebel- lum is placed; and the former, a portion of the least fossa for the basis of the brain. On the posterior side of the pars petrosa, we ob- serve the meatus auditorius internus, into which enters the double nerve of the seventh pair. On the under side of this process, part of a hole appears, which is common to the Osteology. 79 temporal and occipital bones; through it the lateral sinus, the eighth pair, and accessory nerves, pass out of the head. The pars petrosa contains several little bones called the bones of the ear; which, as they do not enter into the formation of the cranium, shall be described when we are treat- ing of the organs of hearing. The ossa temporum are joined to the ossa malarum, by the zygomatic sutures ; to the pa- rietal bones, by the squamous sutures ; to the os occipitis, by the lambdoidal suture ; and to the sphenoid bone, by the suture of that name. This bone, from its situation amidst the other bones of the head, has sometimes been called cuneiforme. It is of a very irregular figure, and has been compared to a bat with its wings extended. It is commonly divided into its middle part or body, and its sides or wings. The fore part of the body has a spine or ridge, which makes part of the septum nari- um. The upper part of each wing forms a share of the temple. The fore part of this belongs to the orbit; while the under and back part, termed spinous process, is lodged in the base of the skull at the point of the pars petrosa. But two of the most remarka- ble processes are the ptergoid or aliform, one on each side of the body of the bone, and at no great distance from it. Each of these pro- cesses is divided into two wings, and of these the exterior one is the widest. The other ter- minates in a hook-like process. 80 Osteology. The internal surface of this bone affords three fossae. Two of these are formed by the wings of the bone, and make a part of the lesser fossae of the basis of the cranium. The third, which is smaller, is on the top of the body of the bone ; and is called sella turcica, from its resemblance to a Turkish saddle. This fossa, in which the pituitary gland is placed, has posteriorly and anteriorly process- es called the clinoid processes. There are twelve holes in this bone, viz. six on each side. The first is the passage of th| optic nerve and ocular artery; the second, or large slit, transmits the third, fourth, sixth, and first part of the fifth pair of nerves with the ocular vein; the third hole gives passage to the second branch of the fifth pair; and the fourth hole to the third branch of the fifth pair of nerves. The fifth hole is the passage of the artery of the dura mater. The sixth hole is situated above the ptergoid process of the sphenoid bone; through it a reflected branch of the second part of the fifth pair passes. Within the substance of the os sphenoides there are two sinuses separated by a bony plate. They are lined with the pituitary mem- brane ; and, like the frontal sinuses, separate a mucus which passes into the nostrils. The os sphenoides is joined to all the bones of the cranium; and likewise to the ossa max- illaria, ossa malarum, ossa palati, and vo- mer. Osteology. 81 This bone makes part of the basis of the skull, assists in forming the orbits, and affords attachment to several muscles. The os ethmoides is situated at the fore part of the basis of the cranium, and is of a very irregular figure. From the great number of holes with which it is pierced, it is some- times called os cribriforme or sieve-like bone. It consists of a middle part and two sides. The middle part is formed of a thin bony plate, in which are an infinite number of holes that afford a passage to filaments of the ol- factory nerve. From the middle of this plate, both on the outside and from within, there rises up a process, which may be easily distinguished. The inner one is called cris- ta galli, from its supposed resemblance to a cock's comb. To this process the falx of the dura mater is attached. The exterior pro- cess, which has the same common basis as the crista galli, is a fine lamella which is united to the vomer; and divides the cavity of the nostrils, though unequally, it being generally a little inclined to one side. The lateral parts of this bone are compos- ed of a cellular substance ; and these cells are so very intricate, that their figure or number cannot be described. Many writers have on this account called this part of the bone the labyrinth. These cells are externally covered with a very thin bony lamella. This part of the bone is called the os planum, and forms part of the orbit. 82 Osteology, The different cells of this bone, which are numerous, and which are every where lined with the pituitary membrane, evidently serve to enlarge the cavity of the nose, in which the organ of smelling resides. This bone is joined to the os sphenoides, os frontis, ossa maxillaria, ossa palati, ossa nasi, ossa unguis, and vomer. The ancients, who considered the brain as the seat of all the humours, imagined that this viscus discharged its redundant moisture through the holes of the ethmoid bone. And the vulgar still think, that abscesses of the brain discharge themselves through the mouth and ears, and that snuff is liable to get into the head; but neither snuff nor the matter of an abscess are more capable of passing through the cribriform bone, than the serosi- ty which they supposed was discharged through it in a common cold.—All the holes of the ethmoid bone are filled up with the branches of the olfactory nerve. Its inner part is likewise covered with the dura mater, and its cells are every where lined with the pituitary membrane; so that neither matter nor any other fluid can possibly pass through this bone either externally or internally. Mat- ter is indeed sometimes discharged through the nostrils; but the seat of the disease is in the sinuses of the nose, and not in the brain; and imposthumations are observed to take place in the ear, which suppurate and discharge themselves externally. Osteology. 83 Before we leave the bones of the head, we wish to make some general observations on its structure and figure.—As the cranium might have been composed of a single bone, the ar- ticulation of its several bones being absolute- ly without motion, it may be asked perhaps, Why such a multiplicity of bones, and so great number of sutures? Many advantages may possibly arise from this plurality of bones and sutures, which may not yet have been observed. We are able, however, to point out many useful ends, which could only be ac- complished by this peculiarity of structure.— In this, as in all the other works of nature, the great wisdom of the Creator is evinced, and can- not fail to excite our admiration and gratitude. The cranium, by being divided into sever- al bones, grows much faster and with greater facility, than if it was composed of one piece only. In the foetus, the bones, as we have before observed, are perfectly distinct from each other. The ossification begins in the middle of each bone, and proceeds gradually to the circumference. Hence the ossification, and of course the increase of the head, is car- ried on from an infinite number of points at the same time, and the bones consequently ap- proach each other in the same proportion. To illustrate this doctrine more clearly, if it can want further illustration, suppose it necessa- ry for the parietal bones which compose the upper part of the head, to extend their ossifi- cation, and form the fore part of the head likewise.—Is it not evident, that this process 84 Osteology. would be much more tedious than it is now, when the os frontis and the parietal bones are both growing at the same time ? Hence it hap- pens, that the heads of young people, in which the bones begin to touch each other, increase slowly ; and that the proportionate increase of the volume of the head is greater in three months in the foetus, than it is perhaps in twenty-four months at the age of fourteen or fifteen years. The sutures, exclusive of their advantage in suspending the processes of the dura ma- ter, are evidently of great utility in prevent- ing the too great extent of fractures of the skull.—Suppose, for instance, that by a fall or blow, one of the bones of the cranium be- comes fractured. The fissure, which in a head composed of only one bone, would be liable to extend itself through the whole of it, is checked, and sometimes perhaps stopped by the first suture it meets, and the effects of the injury are confined to the bone on which the blow was received. Ruysch indeed, and some others, will not allow the sutures to be of any such use; but cases have been met with where they seemed to have had this ef- fect, and in young subjects their utility in this respest must be still more obvious. The spherical shape of the head seems like- wise to render it more capable of resisting ex- ternal violence than any other shape would do. In a vault, the parts mutually support and strengthen each other, and this happens in the cranium. Osteology. 85 2. Proper Bones of the Face. The face, which consists of a great number of bones, is commonly divided into the upper and lower jaws. The upper jaw consists of thirteen bones, exclusive of the teeth. Of these, six are placed on each side of the max- illa superior, and one in the middle. The bones, which are in pairs, are the os- sa malarum, ossa maxillaria, ossa nasi, ossa unguis, ossa palati, and ossa spongiosa inferi- ora. The single bone is the vomer. These are the prominent square bones which are placed under the eyes, forming part of the orbits and the upper part of the cheeks. Each of them affords three surfaces ; one ex- terior and a little convex ; a second superior and concave, forming the inferior part and sides of the orbit; and a third posterior, irre- gular, and hollowed for the lodgment of the lower part of the temporal muscle, The angles of each bone form four process- es, two of which may be called orbitar process- es ; of these the upper one is joined by suture to the os frontis, and that below to the maxil- lary bone. The third is connected with the os sphenoides by means of the transverse su- ture ; and the fourth is joined to the zygoma- tic process of the temporal bone, with which it forms the zygoma. These bones, which are of a very irregular, figure, are so called because they form the most considerable portion of the upper jaw. 86 Osteology. They are two in number, and generally re- main distinct through life. Of the many processes which are to be seen on these bones, and which are connect- ed with the bones of the face and skull, we shall describe only the most remarkable. One of these processes is at the upper and fore part of the bone, making part of the side of the nose, and called the nasal process. An- other forms a kind of circular sweep at the in- ferior part of the bone, in which are the alve- oli or sockets for the teeth: this is called the alveolar process. A third process is united to the os malae on each side. Between this and the nasal process there is a thin plate, which forms a share of the orbit, and lies over a passage for the superior maxillary vessels and nerves.—The alveolar process has posterior- ly a considerable tuberosity on its internal surface, called the maxillary tuberosity. Behind the alveolar process we observe two horizontal lamellae, which uniting together, form a part of the roof of the mouth, and di- vide it from the nose. The hollowness of the roof of the mouth is owing to this partition's being seated somewhat higher than the alveo- lar process.—At the fore part of the horizon- tal lamellae there is a hole called foramen inci- sivum, through which small blood-vessels and nerves go between the mouth and nose. In viewing these bones internally, we ob- serve a fossa in the inferior portion of the l^asal process, which with the os unguis and Osteology, 87 os spongiosum inferius, forms a passage for the lachrymal duct. Where these two bones are united to each other, they project somewhat upwards and forwards, leaving between them a furrow, in- to which the lower portion of the septum na- si is admitted^ Each of these bones being hollow, a consi- derable sinufc is formed under its orbitar part. This cavity, which is usually named after Highmore, though it was described by Fallopi- us and others before his time, is lined with the pituitary membrane. It is intended for the same purposes as the other sinuses of the nose, and opens into the nostrils. The ossa maxillaria are connected with the greater part of the bones of the face and crani- um, and assist in forming not only the cheeks, but likewise the palate, nose, and orbits. The ossa nasi form two irregular squares. They are thicker and narrower above than below. Externally they are somewhat convex, and internally slightly concave. These bones constitute the upper part of the nose. At their fore part they are united to each other, above to the os frontis, by their sides to the ossa maxillaria superiora, posteriorly and in- teriorly to the septum narium, and below to the cartilages, that compose the rest of the nostrils. These little transparent bones owe their name to their supposed resemblance to a fin- ger-nail. Sometimes they are called ossa la- chrymalia, from their concurring with the na- 88 Osteology. sal process of each maxillary bone in forming a lodgement for the lachrymal sac and duct. The ossa unguis are of an irregular figure. Their external surface consists of two smooth parts, divided by a middle ridge. One of these parts, which is concave and nearest to the nose, serves to support the lachrymal sac and part of the lachrymal duct. The other, which is flat, forms a small part of the orbit. Each of these bones is connected with the os frontis, os ethmoides, and os maxillare su- perius. These bones which are situated at the back part of the roof of the mouth, between the os sphenoides and the ossa maxillaria superiora, are of a very irregular shape, and serve to form the nasal and maxillary fossa, and a small portion of the orbit. Where they are united to each other, they rise up into a spine on their internal surface. This spine appears to be a continuation of that of the superior max- illary bones, and helps to form the septum narium. These bones are joined to the ossa maxilla- ria superiora, os ethmoides, os sphenoides, and vomer. This bone derives its name from its resem- blance to a ploughshare. It is a long and flat bone, somewhat thicker at its back than at its fore part. At its upper part we observe a furrow extending through its whole length. The posterior and largest part of this furrow receives a process of the sphenoid bone. From this the furrow advances forwards, and be- Osteology. # 89 coming narrower and shallower, receives some part of the nasal lamella ethmoidea; the rest serves to support the middle cartilage of the nose. The inferior portion of this bone is placed on the nasal spine of the maxillary and palate bones, which we mentioned in our description of the ossa palati. The vomer is united to the os sphenoides, os ethmoides, ossa maxillaria superiora, and ossa palati. It forms part of the septum nari- um, by dividing the back part of the nose into two nostrils. The parts which are usually described by this name, do not seem to deserve to be dis- tinguished as distinct bones, except in young subjects. They consist of a spongy lamella in each nostril, which is united to the spongy lamina of the ethmoid bone, of which they are by some considered as a part. Each of these lamellae is longest from be- hind forwards ; with its convex surface turn- ed towards the septum narium, and its con- cave part towards the maxillary bone, covering the opening of the lachrymal duct into the nose. These bones are covered with the pituitary membrane ; and, besides their connection with the ethmoid bone, are joined to the ossa max- illaria superiora, ossa palati, and ossa unguis. The maxilla inferior, or lower jaw, which in its shape resembles a horse-shoe, consists of two distinct bones in the foetus; but these unite together soon after birth, so as to form M 90 Osteology, only one bone. The upper edge of this bone, like the os maxillare superius, has an alveolar process, furnished with sockets for the teeth. On each side the posterior part of the bone rises almost perpendicularly into two process- es. The highest of these, called the coronoid process, is pointed .-and thin, and serves for the insertion of the temporal muscle. The other, or condyloid process, as it is called, is shorter and thicker, and ends in an oblong rounded head, which is received into a fossa of the temporal bone, and is formed for a moveable articulation with the cranium. This joint is furnished with a moveable cartilage. At the bottom of each coronoid process, on its inner part, we observe a foramen extend- ing under the roots of all the teeth, and ter- minating at the outer surface of the bone near the chin. Each of these canals transmits an artery, vein, and nerve, from which branches are sent off to the teeth. The lower jaw is capable of a great variety of motion. By sliding the condyles from the cavity towards the eminences on each side, we bring it horizontally forwards, as in biting; or we may bring the condyles only forward, and tilt the rest of the jaw backward, as in opening the mouth. We are likewise able to slide the condyles alternately backwards and forwards from the cavity to the eminence, and vice versa, as in grinding the teeth. The car- tilages, by adapting themselves to the differ- ent inequalities in these several motions of the Osteology. 91 jaw, serve to secure the articulation, and to prevent any injuries from friction. The alveolar processes are composed of an outer and inner bony plate, united together by thin partitions, which at the fore part of the jaw divide the processes into as many sockets as there are teeth. But at the back part of the jaw, where the teeth have more than one root, we find a distinct cell for each root. In both jaws these processes begin to be formed with the teeth; they likewise accompany them in their growth, and gradually disappear when the teeth are removed. 3. Of the Teeth. The teeth are bones of a particular struc- ture, formed for the purposes of mastication and the articulation of the voice. It will be necessary to consider their composition and fi- gure, their number and arrangement, and the time and order in which they appear. In each tooth we may distinguish a. body, a neck, and a root or fangs. The body of the tooth is that part which appears above the gums. The root is fixed into the socket, and the neck is the middle part between the two. The teeth are composed of two substances, viz. enamel and bone. The enamel, or the vitreous or cortical part of the tooth, is a white and very hard and compact substance peculiar to the teeth, and appears fibrous o*r 92 Osteology. striated when broken. This substance is thick* est on the grinding surface, and becoming gra- dually thinner, terminates insensibly at the neck of the tooth. Ruysch * affirmed, that he could trace the arteries into the hardest part of the teeth; Liewenhoeckf suspected the fibres of the enamel to be so many vessels; and Monro % says, he has frequently injected the vessels of the teeth in children, so as to make the inside of the cortex appear perfectly red. But it is certain, that it is not tinged by a madder diet, and that no injection will ever reach it, so that it has no appearance of be- ing vascular §. The bony part, which composes the inner substance of the body, neck, and root of the tooth, resembles other bones in its structure, but it is much harder than the most compact part of bones in general. As a tooth when once formed receives no tinge from a madder diet, and as the minutest injections do not pe- netrate into its substance, this part of the tooth has, like the enamel, been supposed not to be vascular. But when we consider that the fangs of a tooth are invested by a perios- teum, and that the swellings of these fangs are analogous to the swellings of other bones, we may reasonably conclude, that there is a similarity of structure; and that this bony part has a circulation through its substance, * Thefadr 10. no. 27. f Arcan. Natur continuat. Epiftbl. % Anat. of the Human Bone's. § Hunter on the Teeth. Osteology. 93 although from its hardness we are unable to demonstrate its vessels. In each tooth we find an inner cavity, into which enter an artery, vein, and nerve. This cavity begins by a small opening, and becom- ing larger, terminates in the body of the tooth. In advanced life this hole sometimes closes, and the tooth is of course rendered insensible. The periosteum surrounds the teeth from their fangs to a little beyond their bony sock- ets, where we find it adhering to the gums. This membrane, while it incloses the teeth, serves at the same time to line the sockets, so that it may be considered as common to both. The teeth are likewise secured in their sock- ets by means of the gums ; a red, vascular, firm, and elastic substance, that possesses but little sensibility. In the gums of infants we find a hard ridge extending through their whole length, but no such ridge is to be seen in old people who have lost their teeth. The number of the teeth in both jaws at full maturity, usually varies from twenty-eight to thirty-two. They are commonly divided into three classes, viz. incisores, canini, and grind- ers or molares.* The incisores are the four teeth in the fore part of each jaw. They have each of them two surfaces ; one anterior * Mr. Hunter has thought proper to vary this division. He retains the old name of incisores to the four fore teeth, but he distinguishes the canine teeth by the name of the cuspidati. The two teeth ••which are next to these, and which have been usu- ally ranked with the molares, he calls the bicuspides; and he gives the name ofgrinders only to the three last teeth on each side. 94 Osteology. and convex, the other posterior and slightly concave, both of which terminate in a sharp edge. They are called incisores from their use in dividing the food. They are usually broader and thicker in the upper than in the under jaw; and, by being placed somewhat obliquely, generally fall over the latter. The canini derive their name from their re- semblance to a dog's tusks, being the longest of all the teeth. We find one on each side of the incisores, so that there are two canini in each jaw. Their fang resembles that of the incisores, but is much larger; and in their shape they appear like an incisor with its edge worn off, so as to terminate in a narrow point. These teeth not being calculated for cutting and dividing the food like the incisores, or for grinding it like the molares, seem to be intend- ed for laying hold of substances.* The molares or grinders, of which there are ten in each jaw, are so called, because from their shape and size they are fitted for grinding the food. Each of the incisores and canini is furnished only with one fang; but in the molares of the under jaw we constantly find two fangs, and in those of the upper jaw three fangs. These fangs are sometimes se- parated into two points, and each of these points has sometimes been described as a distinct fang. * Mr. Hunter remarks of these teeth, that we may trace in them a similiarity in shape, situation, and use, from the most imperfectly carnivorous animal, which we believe to be the hu- man species, to the lion, which is the most perfectly carnivo- rous. Osteology. 95 The two first of the molares, or those near- est to the canine teeth on each side, differ from the other three, and are with great pro- priety named bicuspides by Mr. Hunter. They have sometimes only one root, and seem to be of a middle nature between the incisores and the larger molares. The two next are much larger. The fifth or last grinder on each side is smaller and shorter than the rest; and from its not cutting the gum till after the age of twenty, and sometimes not till much later in life, is called dens sapientie. There is in the structure and arrangement of all these teeth an art which cannot be suf- ficiently admired. To understand it properly, it will be necessary to consider the under jaw as a kind of lever, with its fixed points at its articulations with the temporal bones:—it will be right to observe, too, that its powers arise from its different muscles, but in elevation chiefly from the temporalis and masseter; and that the aliment constitutes the object of re- sistance. It will appear, then, that the mola- res, by being placed nearest the centre of mo- tion, are calculated to press with a much greater force than the other teeth, independent of their grinding powers which they possess by means of the pterygoid muscles; and that it is for this reason we put between them any hard body we wish to break. The canini and incisores are placed farther from this point, and of course cannot exert so much force ; but they are made for cutting and tearing the food, and this form seems to make amends for their deficiency in strength. 96 Osteology. There are examples of children who have come into the world with two, three, and even four teeth; but these examples are very rare; and it is seldom before the seventh, eighth, or ninth month after birth, that the incisores, which are the first formed, begin to pass through the gum. The symptoms of denti- tion, however, in consequence of irritation from the teeth, frequently take place in the fourth or fifth month.—About the twentieth or twenty-fourth month, the canini and two molares make their appearance. The dangerous symptoms that sometimes accompany dentition, are owing to the pres- sure of the teeth on the gum, which they ir- ritate so as to excite pain and inflammation. This irritation seems to occasion a gradual wasting of the gum at the part, till at length the tooth makes its appearance. The symptoms are more or less alarming, in proportion to the resistance which the gum affords to the teeth, and according to the num- ber of teeth which may chance to seek a pas- sage at the same time. Were they all to ap- pear at once, children would fall victims to the pain and excessive irritation; but Nature has so very wisely disposed them, that they usually appear one after the other, with some distance of time between each. The first in- cisor that appears is generally in the lower jaw, and is followed by one in the upper jaw. Sometimes the canini, but more commonly one of the molares, begins to pass through the gum first. Osteology. 97 These 20 teeth, viz. eight incisores, four canini, and eight molares, are called tempora- ry or milk teeth, because they are all shed be- tween the age of seven and fourteen, and are succeeded by what are called the permanent or adult teeth. The latter are of a firmer texture, and have larger fangs. These adult teeth being placed in a distinct set of alveoli, the upper sockets gradually dis- appear, as the under ones increase in size, till at length the temporary, or upper teeth, hav- ing no longer any support, consequently fall out. To these 20 teeth, which succeed the tem- porary ones, 12 others are afterwards added, viz. three molares on each side in both jaws: and in order to make room for this addition, we find that the jaws gradually lengthen in proportion to the growth of the teeth; so that with 20 teeth, they seem to be as completely filled as they are afterwards with 32. This is the reason why the face is rounder and flatter in children than in adults. With regard to the formation of the teeth, we may observe, that in a foetus of four months, the alveolar process appears only as a shallow longitudinal groove, divided by minute ridges into a number of intermediate depressions ; in each of which we find a small pulpy substance surrounded by a vascular membrane. This pulp gradually ossifies, and its lower part is lengthened out to form the fang. When the bony part of the tooth is formed, its surface begins to be incrusted with the enamel. How N 98 Osteology. the latter is formed and deposited, we are not yet able to determine. The rudiments of some of the adult teeth begin to be formed at a very early period, for the pulp of one of the incisores may general- ly be perceived in a foetus of eight months, and the ossification begins in it soon after birth. The first bicuspis begins to ossify about the fifth or sixth, and the second about the seventh year. The first adult grinder cuts the gum about the 12th, the second about the 18th, and the third, or dens sapientie, usually between the 20th and 30th year. The teeth, like other bones, are liable to be affected by disease. Their removal is like- wise the natural consequences of old age; for as we advance in life, the alveoli fill up, and the teeth, especially the incisores, fall out. When this happens, the chin projects forward, and the face is much shortened. 4. Of the Os Hyoides.* The os hyoides, which is placed at the root of the tongue, was so called by the ancients on account of its supposed resemblance to the Greek letter v. * This bone is very seldom preserved with the skeleton, and cannot be included among the bones of the head, or any other division of the skeleton Thomas Bartholin has perhaps very properly described it among parts contained in the mouth ; but the generality of anatomical writers have placed it, as it is here, after the bones of the face. Osteology. 99 It will be necessary to distinguish in it, its body, horns, and appendices. The body, which is the middle and broad- est part of the bone, is so placed that it may be easily felt at the fore part of the throat. Anteriorly it is irregularly convex, and its in- ner surface is unequally concave. Its cornua, or horns, which are flat and a little bent, be- ing much longer than the body part, may be described as forming the sides of the «. The appendices, or little horns, as they are called by M. Windslow, and some other writers, are two processes which rise up from the articu- lations of the cornua with the body, and are usually connected with the styloid process on each side by means of a ligament. The uses of this bone are to support the tongue, and afford attachment to a great num- ber of muscles; some of which perform the motions of the tongue, while others act on the larynx and fauces. Sect. III. Of the Bo7ies of the Trunk. The trunk of the skeleton consists of the spine, the thorax, and the pelvis. 1. Of the Spine, The spine is composed of a great number of bones called vertebra, forming a long bo- ny column, in figure not much unlike the let- ter/. This column, which extends from the 100 Osteology. head to the lower part of the body, may be said to consist of two irregular and unequal pyramids, united to each other in that part of the loins where the last lumbar vertebra joins the os sacrum. The vertebrae of the upper and longest py- ramid are called true vertebra, in contradis- tinction to those of the lowermost pyramid, which, from their being immoveable in the adult, are s\y\e& f dse vertebra. It is upon the bones of the spine that the body turns ; and it is to this circumstance they owe their name, which is derived from the Latin verb vertere, to turn jMmsk. 2Mc«li«?ill C«ll«~tv The true vertebrae are divided into three classes of cervical, dorsal and lumbar verte- brae.—The false vertebrae consist of the os sa- crum and os coccygis. In each vertebra, as in other bones, it will be necessary to remark the body of the bone, its processes, and cavities. The body, which is convex before, and concave behind, where it assists in forming the cavity of the spine, may be compared to part of a cylinder cut off' transversely. Each vertebra affords seven processes. The first is at the back part of the vertebra, and from its shape and direction is named the spinous process. On each side of this are two others, which, from their situation with re- spect to the spine, are called transverse pro- cesses. The four others are styled oblique or articular processes. They are much smaller than the spinous or transverse ones. Two of Osteology. 101 them are placed on the upper, and two on the lower part of each vertebra, rising from near the basis of each transverse process. They have gotten the name of oblique processes, from their situation with respect to the processes with which they are articulated ; and they are sometimes styled articular processes, from the manner in which they are articulated with each other; the two superior processes of one vertebra being articulated with the two infe- rior processes of the vertebra above it. Each of these processes is covered with cartilage at its articulation, and their articulations with each other are by a species of ginglimus. In each vertebra, between its body and its processes, we find a hole large enough to ad- mit a finger. These holes or foramina, corre- spond with each other through all the vertebrae, and form the long bony channel in which the spinal marrow is placed. We may likewise observe four notches in each vertebra. Two of these notches are at the upper, and two at the lower part of the bone, between the oblique processes and the body of the ver- tebra. Each of these notches meeting with a similar opening in the vertebra above or be- low it, forms a foramen for the passage of blood-vessels, and of the nerves out of the spine. The bones of the spine are united toge- ther by means of a substance, which in young subjects appears to be of a ligamentous, but in adults more of a cartilaginous nature. This intervertebral substance, which forms a kind 102 Osteology. of partition between the several vertebrae, is thicker and more flexible between the lumbar vertebrae than in the other parts of the spine, the most considerable motions of the trunk being performed on those vertebrae. This substance being very elastic, the extension and flexion of the body, and its motion backwards and forwards, or to either side, are performed with great facility. This elasticity seems to be the reason why people who have been long stand- ing, or have carried a considerable weight, are found to be shorter than when they have been long in bed. In the two first instances the intervertebral cartilages (as they are usu- ally called) are evidently more exposed to compression than when we are in bed in an horizontal posture. In advanced life these cartilages become shrivelled, and of course lose much of their elasticity. This may serve to account for the decrease in stature and the stooping forward which are usually to be observed in old peo- ple. Besides the connection of the several verte- brae by means of this intervertebral substance, there are likewise many strong ligaments, both external and internal, which unite the bones of the spine to each other. Their union is al- so strengthened by a variety of strong muscles that cover and surround the spine. The bones of the spine are found to dimi- nish in density, and to be less firm in their texture in proportion as they increase in bujk; so that the lowermost vertebrae, though the Osteology. 103 largest, are not so heavy in proportion as the upper ones. By this means the size of these bones is increased without adding to their weight: a circumstance of no little importance in a part like the spine, which, besides flexi- bility and suppleness, seems to require light- ness as one of its essential properties. In very young children, each vertebra con- sists of three bony pieces united by cartilages which afterwards ossify. There are seven vertebrae of the neck— they are of a firmer texture than the other bones of the spine. Their transverse process- es are forked for the lodgment of muscles, and at the bottom of each we observe a fora- men, through which pass the cervical artery and vein. The first and second of these ver- tebrae must be described more particularly. The first approaches almost to an oval shape —On its superior surface it has two cavities which admit the condyles of the occipital bone with which it is articulated. This verte- bra, which is called atlas from its supporting the head, cannot well be described as having either body or spinous process, being a kind of bony ring. Anteriorly, where it is articu- lated to the odontoid process of the second vertebra, it is very thin. On its upper surface it has two cavities which admit the condyles of the occipital bone. By this connection the head is allowed to move forwards and back- wards, but has very little motion in any other direction. 104 Osteology. The second vertebra has gotten the name of dentata, from its having, at its upper and an- terior part, a process called the odontoid or tooth-like process, which is articulated with the atlas, to which this second vertebra may be said to serve as an axis. This odontoid pro- cess is of a cylindrical shape, somewhat flat- tened, however, anteriorly and posteriorly. At its fore-part where it is received by the atlas, we may observe a smooth, convex, articulat- ing surface. It is by means of this articula- tion that the head performs its rotatory motion, the atlas in that case moving upon this odon- toid process as upon a pivot. But when this motion is in any considerable degree, or, in other words, when the head moves much ei- ther to the right or left, all the cervical verte- brae seem to assist, otherwise the spinal mar- row would be in danger of being divided trans- versely by the first vertebra. The spinous process of each of the cervi- cal vertebrae is shorter, and their articular pro- cesses more oblique, than in the other bones of the spine. These 12 vertebrae are of a middle size be- tween those of the neck and loins. At their sides we may observe two depressions, one at the upper and the other at the lower part of the body of each vertebrae; which uniting with similar depressions in the vertebrae above and below, form articulating surfaces, covered with cartilages, for receiving the heads of the ribs ; and at the fore-part of their transverse process Osteology. 105 (excepting the two last) we find an articulating surface for receiving the tuberosity of the ribs. These five vertebrae differ only from those of the back in their being larger, and in hav- ing their spinous processes at a greater dis- tance from each other. The most consider- able motions of the trunk are made on these vertebras; and these motions could not be per- formed with so much ease, were the process- es placed nearer to each other. The os sacrum, which is composed of five or six pieces in young subjects, becomes one bone in more advanced age. It is nearly of a triangular figure, its infe- rior portion being bent a little forwards. Its superior part has two oblique processes which are articulated with the last of the lumbar ver- tebrae ; and it has likewise commonly three small spinous processes, which gradually be- come shorter, so that the lowermost is not so long as the second, nor the second as the up- permost. Its transverse processes are formed into one oblong process, which becomes gra- dually smaller as it descends. Its concave or anterior side is usually smooth, but its poste- rior convex side has many prominences (the most remarkable of which are the spinous pro- cesses just now mentioned,) which are filled up and covered with the muscular and tendi- nous parts behind. This bone has five pair of holes, which af- ford a passage to blood-vessels, and likewise to the nerves that are derived from the spinal marrow, which is continued even here, being O 106 Osteology. lodged in a triangular cavity, that becomes smaller as it descends, and at length termi- nates obliquely at the lower part of this bone. Below the third division of the os sacrum, this canal is not completely bony as in the rest of the spine, being secured at its back part only by a very strong membrane, so that a wound at this part must be extremely dangerous. The os sacrum is united laterally to the os- sa innominata or hip-bones, and below to the coccyx. The coccyx, which, like the os sacrum, is in young people made up of three or four dis- tinct parts, usually becomes one bone in the adult state. It serves to support the intestinum rectum; and, by its being capable of some degree of motion at its articulation with the sacrum, and being like that bone bent forwards, we are en- abled to sit with ease. This bone is nearly of a triangular shape, , being broadest at its upper part, and from thence growing narrower to its apex,'where it is not bigger than the little finger. It has got its name from its supposed re- semblance to a cuckow's beak. It differs greatly from the vertebrae, being commonly without any processes, and having no cavity for the spinal marrow, or foramina for the transmission of nerves. The spine, of which we have now finished the anatomical description, is destined for ma- ny great and important uses. The medulla spinalis is lodged in its bony canal secure from Osteology. 107 external injury. It serves as a defence to the abdominal and thoracic viscera, and at the same time supports the head, and gives a ge- neral firmness to the whole trunk. We have before compared it to the letter^ and its different turns will be found to render it not very unlike the figure of that letter.— In the neck we see it projecting somewhat for- ward to support the head, which without this assistance would require a great number of muscles.—Lower down, in the thorax, we find it taking a curved direction backwards, and of course increasing the cavity of the chest. Af- ter this, in the loins, it again projects forwards in a direction with the centre of gravity, by which means we are easily enabled to keep the body in an erect posture, for otherwise we should be liable to fall forward. Towards its inferior extremity, however, it again recedes backward, and thus assists in forming the pel- vis, the name given to the cavity in which the urinary bladder, intestinum rectum, and other viscera are placed. If this bony column had been formed only of one piece, it would have been much more easily fractured than it is now: and by confin- ing the trunk to a stiff situation, a variety of motions would have been altogether prevent- ed, which are now performed with ease by the great number of bones of which it is compos- ed. It is firm, and yet to this firmness there is added a perfect flexibility. If it be required to carry a load upon the head, the neck be- 108 Osteology. comes stiff with the assistance of its muscles, and accommodates itself to the load, as if it was composed only of one bone— In stooping likewise, or in turning to either side, the spine turns itself in every direction, as if all its bones were separated from each other. In a part of the body, like the spine, that is made up of so great a number of bones, and intended for such a variety of motion, there must be a greater danger of dislocation than fracture ; but we shall find, that this is very wisely guarded against in every direction by the processes belonging to each vertebra, and by the ligaments, cartilages, he. by which these bones are connected with each other. 2. Of the Bones of the Thorax. The thorax, or chest, is composed of ma- ny bones, viz. the sternum which is placed at its anterior part, twelve ribs on each side which make up its lateral parts, and the dor- sal vertebrae which constitute its posterior part. These last have been already described. The sternum is the long bone which ex- tends itself from the upper to the lower part of the breast anteriorly, and to which the ribs and the clavicles are articulated. In children it is composed of several bones united by cartilages ; but as we advance in life, most of these cartilages ossify, and the ster- num in the adult state is found to consist on- Osteology. 109 ly of three pieces, and sometimes becomes one bone. It is however generally described as being composed of three parts—one superior, which is broad, thick, and short; and one in the middle, which is thinner, narrower, and longer than the other. It terminates at its lower part by a third piece, which is called the xyphoid, or sword- like cartilage, from its supposed resemblance to the blade of a sword, and because in young subjects it is commonly in a cartilaginous state. We have already observed, that this bone is articulated with the clavicle on each side. It is likewise joined to the fourteen true ribs, viz. seven on its right and seven on its left side. The ribs are bones shaped like a bow, form- ing the sides of the chest. There are twelve on each side. They are distinguished into true and false ribs : The seven upper ribs which are articulated to the sternum are call- ed true ribs, and the five lower ones that are not immediately attached to that bone are call- ed false ribs. On the inferior and interior surface of each rib, we observe a sinuosity for the lodgment of an artery, vein, and nerve. The ribs are not bony through their whole length, their anterior part being cartilaginous. They are articulated with the vertebrae and sternum. Every rib (or at least the greater number of them) has at its posterior part two processes ; one at its extremity called the head of the rib, by means of which it is articulated 110 Osteology. with the body of two vertebrae; and another, called its tuberosity, by which it is articulated with the transverse process of the lowest of these two vertebrae. The first rib is not arti- culated by its extremity to two vertebrae, be- ing simply attached to the upper part of the first vertebra of the back. The seven superi- or or true ribs are articulated anteriorly with the sternum by their cartilages ; but the false ribs are supported in a different manner—the eighth, which is the first of these ribs, being attached by its cartilage to the seventh ; the ninth to the eighth, &c. The two lowermost ribs differ likewise from all the rest in the following particulars: they are articulated only with the body of the ver- tebra, and not with a transverse process ; and anteriorly, their cartilage is loose, not being attached to the cartilages of the other ribs; and this seems to be, because the most consi- derable motions of the trunk are not perform- ed on the lumbar vertebrae alone, but likewise on the two last vertebrae of the back; so that if these two ribs had been confined at the fore part like the other ribs, and had been likewise articulated with the bodies of two vertebrae, and with the transverse processes, the moti- on of the two last vertebrae, and consequently of the whole trunk, would have been impeded. The ribs help to form the cavity of the tho- rax ; they afford attachment to different mus- cles ; they are useful in respiration; and they serve as a security to the heart and lungs. Osteology. ill 3. Of the Bones of the Pelvis. The pelvis is composed of the os sacrum, os coccygis, and two ossa innominata. The two first of these bones were included in the account of the spine, to which they more pro- perly belong. In children, each os innominatum is com- posed of three distinct bones; but as we ad- vance in life the intermediate cartilages gradu- ally ossify, and the marks of the original se- paration disappear, so that they become one irregular bone ; still however continuing to re- tain the names of ilium, ischium, and pubis, by which their divisions were originally dis- tinguished, and to be described as three dif- ferent bones by the generality of anatomists. The os ilium forms the upper and most consi- derable part of the bone, the os ischium its lower and posterior portion, and the os pubis its fore part. The os ilium or haunch bone, is articulated posteriorly to the os sacrum by a firm cartila- ginous substance, and is united to the os pu- bis before and to the os ischium below. Its superior portion is thin, and terminates in a ridge called the crista or spine of the ilium, and more commonly known by the name of the haunch. This crista rises up like an arch; being turned somewhat outwards, so as to re- semble the wings of a phaeton. Externally this bone is unequally prominent and hollowed for the lodgment of muscles; in- ternally we find it smooth and concave. At 112 Osteology. its lower part there is a considerable ridge on its inner surface. This ridge extends from the os sacrum, and corresponds with a similar prominence both on that bone and the ischi- um ; forms with the inner part of the ossa pu- bis what in midwifery is termed the brim of the pelvis. The crista or spine, which at first is an epi- physis, has two considerable tuberosities ; one anteriorly, and the other posteriorly, which is the largest of the two: these, from their pro- jecting more than the parts of the bone below them, have gotten the name of spinal process- es. From the anterior spinous process, the sartorius and tensor vaginae femoris muscles have their origin; and below the posterior process we observe a considerable niche in the bone, which, in the recent subject, is formed into a large foramen, by means of a strong ligament that is stretched over its low- er part from the os sacrum to the sharp-point- ed process of the ischium. This hole affords a passage to the great sciatic nerve, and to the posterior crural vessels under the pyriform muscle, part of which likewise passes out here. The os ischium, or hip-bone, which is of a very irregular figure, constitutes the lower lateral parts of the pelvis, and is commonly divided into its body, tuberosity, and ramus. The body forms the lower and most consider- able portion of the acetabulum, and sends a sharp-pointed process backwards, called the spine of the ischium. To this process the li- Osteology. 113 gament adheres, which was just now spoken of, as forming a foramen for the passage of the sciatic nerve.—The tuberosity, which is the lowest part of the trunk, and supports us when we sit, is large and irregular, affording origin to several muscles. From this tubero- sity we find the bone becoming thinner and narrower. This part, which has the name of ramus or branch, passes forwards and up- wards, and concurs with the ramus of the os pubis, to form a large hole called the foramen magnum ischii, or thyroideum, as it is some- times named, from its resemblance to a^door or shield. This hole, which in the recent subject is closed by a strong membrane called the obturator ligament, affords through its whole circumference attachment to muscles. At its upper part where we observe a niche in the bone, it gives passage to the obturator vessels and nerves, which go to the inner part of the thigh. Nature seems every where to avoid an unnecessary weight of bone, and this foramen, no doubt, serves to lighten the bones of the pelvis. The os pubis or share-bone, which with its fellow forms the fore-part of the pelvis, is the smallest division of the os innominatum. It is united to its fellow by means of a strong cartilage, which forms what is called the sym- physis pubis. In each os pubis we may distinguish the body of the bone, its angle, and ramus. The body or outer part is united to the os ilium. The angle comes forward to form the sym- P 114 Osteology. physis, and the ramus is a thin process which unites with the ramus of the ischium, to form the foramen thyroideum. The three bones we have described as com- posing each os innominatum, all assist in form- ing the acetabulum, in which the head of the os femoris is received. This cavity is every where lined with a smooth cartilage, excepting at its inner part, where we may observe a little fossa, in which are lodged the mucilaginous glands of the joint. We may likewise notice the pit or depression made by the round ligament, as it is improper- ly called, which, by adhering to this cavity and to the head of the thigh-bone, helps to secure the latter in the socket. These bones, which are united to each other and to the spine by many very strong ligaments, serve to support the trunk, and to connect it with the lower extremities ; and at the same time to form the pelvis or bason, in which are lodged the intestines and urinary bladder, and in women the uterus ; so that the study of this part of osteology is of the utmost importance in midwifery. It is worthy of observation, that in women the os sacrum is usually shorter, broader, and more hollowed, the ossa ilia more expanded, and the inferior opening of the pelvis larger than in men. Sect. IV. Of the Extremities. These parts of the skeleton consist of the upper extremity and the lower. Osteology. 115 1. Of the Upper Extremity. This consists of the shoulder, the arm, and the hand. 1. Of the shoulder. The shoulder consists of two bones, the clavicula and the scapula. The former, which is so named from its re- semblance to the key in use amongst the an- cients, is a little curved at both its extremities like an italic f. It is likewise called jugulum, or collar-bone, from its situation. It is about the size of the little finger, but longer, and be- ing of a very spongy substance is very liable to be fractured. In this, as in other long bones, we may distinguish a body and two extremi- ties. The body is rather flattened than round- ed. The anterior extremity is formed into a slightly convex head, which is nearly of a tri- angular shape. The inferior surface of the head is articulated with the sternum. The posterior extremity, which is flatter and broad- er than the other, is connected to a process of the scapula, called acromion. Both these articulations are secured by ligaments, and in that with the sternum we meet with a move- able cartilage, to prevent any injury from fric- tion. The clavicle serves to regulate the motions of the scapula, by preventing it from being 116 Osteology. brought too much forwards, or carried too far backwards. It affords origin to several mus- cles, and helps to cover and protect the sub- clavian vessels, which derive their name from their situation under this bone. The scapula, or shoulder-blade, which is nearly of a triangular shape, is fixed to the posterior part of the true ribs, somewhat in the manner of a buckler. It is of a very un- equal thickness, and, like all other broad, flat bones, is somewhat cellular. Exteriorly it is convex, and interiorly concave, to accommo- date itself to the convexity of the ribs. We observe in this bone three unequal sides, which are thicker and stronger than the body of the bone, and are therefore termed its costa. The largest of the three, called also the basis, is turned towards the vertebrae. Another, which is less than the former, is below this ; and the third, which is the least of the three, is at the upper part of the bone. Externally the bone is elevated into a considerable spine, which rising small at the basis of the scapula, becomes gradually higher and broader, and divides the outer surface of the bone into two fossae. The superior of these, which is the smallest, serves to lodge the supra spinatus muscle ; and the inferior fossa, which is much larger than the other, gives origin to the infra spinatus. This spine terminates in a broad and flat process at the top of the shoulder, called the processus acromion, to which the clavicle is articulated. This process is hollowed at its lower part to allow a passage to the supra and Osteology. 117 infra spinati muscles. The scapula has like- wise another considerable process at its upper part, which, from its resemblance to the beak of a bird, is called the coracoid process. From the outer side of this coracoid process, a strong ligament passes to the processus acromion, which prevents a luxation of the os humeri up- wards. A third process begins by a narrow neck, and ends in a cavity called glenoid, for the connection of the os humeri. The scapula is articulated with the clavicle and os humeri, to which last it serves as a ful- crum ; and by varying its position it affords a greater scope to the bones of the arm in their different motions. It likewise gives origin to several muscles, and posteriorly serves as a de- fence to the trunk. 2. Bones of the Arm. The arm is commonly divided into two parts, which are articulated to each other at the el- bow. The upper part retains the name of arm, properly so called, and the lower part is usu- ally called the fore-arm. The arm is composed of a single bone call- ed os humeri. This bone, which is almost of a cylindrical shape, may be divided into its bo- dy and its extremities. The upper extremity begins by a large, round smooth head, which is admitted into the glenoid cavity of the scapula. On the upper and fore part of,the bone there is a groove for lodging the long head of the biceps mus- 118 Osteology. cle of the arm ; and on each side of the groove, at the upper end of the bone, there is a tuber- cle to which the spinata muscles are fixed. The lower extremity has several processes and cavities. The principal processes are its two condyles, one exterior and the other inte- rior, and of these the last is the largest. Be- tween these two we observe two lateral protu- berances, which, together with a middle cavi- ty, form as it were a kind of pully upon which the motions of the fore-arm are chiefly per- formed. At each side of the condyles, as well exteriorly as interiorly, there is another emi- nence which gives origin to several muscles of the hand and fingers. Posteriorly and superi- orly, speaking with respect to the condyles, we observe a deep fossa which receives a consi- derable process of the ulna ; and anteriorly and opposite to this fossa, we observe another, which is much less and receives another pro- cess of the same bone. The body of the bone has at its upper and anterior part a furrow which begins from be- hind the head of the bone, and serves to lodge the tendon of a muscle. The body of the os humeri is hollow through its whole length, and, like all other long bones, has its marrow. This bone is articulated at its upper part to the scapula. This articulation, which allows motion every way, is surrounded by a capsu- lar ligament; that is sometimes torn in luxa- tion, and becomes an obstacle to the easy re- duction of the bone. Its lower extremity is articulated with the bones of the fore-arm. Osteology. 119 The fore-arm is composed of two bones, the ulna and radius. The ulna or elbow-bone is much less than the os humeri, and becomes gradually small- er as it descends to the wrist. At its upper part it has two processes and two cavities. Of the two processes, the largest, which is situ- ated posteriorly, and called the olecranon, is admitted into the posterior fossa of the os hu- meri. The other process is placed anteriorly, and is called the coronoid process. In bending the arm it enters into the anterior fossa of the os humeri. This process being much smaller than the other, permits the fore-arm to bend inwards; whereas the olecranon, which is shaped like a hook, reaches the bottom of its fossa in the os humeri as soon as the arm be- comes straight, and will not permit the fore- arm to be bent backwards. The ligaments likewise oppose this motion. Between the two processes we have describ- ed, there is a considerable cavity called the sygmoid cavity, divided into two fossae by a small eminence, which passes from one pro- cess to the other; it is by means of this cavity and the two processes, that the ulna is articu- lated with the os humeri by ginglimus. At the bottom of the coronoid process interi- orly, there is a small sygmoid cavity, which serves for the articulation of the ulna with the radius. The body of the ulna is of a triangular shape: Its lower extremity terminates by a small'head and a little styloid process. The 12(5 Osteology. ulna is articulated above to the os humeri— both above and below to the radius, and to the wrist at its lower extremity. All these ar- ticulations are secured by means of ligaments. The chief use of this bone seems to be to sup- port and regulate the motions of the radius. The radius, which is so named from its sup- posed resemblance to the spoke of a wheel, is placed at the inside of the fore-arm. It is somewhat larger than the ulna, but not quite so long as that bone. Its upper part is cylin- drical, hollowed superiorly to receive the out- er condyle of the os humeri. Laterally it is admitted into the little sygmoid cavity of the ulna, and the cylindrical part of the bone turns in this cavity in the motions of pronation and supination.* This bone follows the ulna in flexion and extension, and may likewise be moved round its axis in any direction. The lower extremity of the radius is much larger and stronger than its upper part; the ulna, on the contrary, is smaller and weaker below than above ; so that they serve to supply each other's deficiencies in both those parts. On the external side of this bone, we ob- serve a small cavity which is destined to re- ceive the lower end of the ulna; and its low- er extremity is formed into a large cavity, by means of which it is articulated with the bones * The motions of pronation and supination may be easily described. If the palm of the hand, for instance, is placed on the surface of a table, the hand may be said to be in a state of pronation; but if the back part of the hand is turned towards the table, the hand will be then in a state of supination. Osteology. 121 of the wrist, and on this account it is some- times called manubrium manus. It supports the two first bones of the wrist on the side of the thumb, whereas the ulna is articulated with that bone of the wrist which corresponds with the little finger. Through the whole length both of this bone and the ulna, a ridge is observed, which affords attachment to an interosseous ligament. This ligament fills up the space between the two bones. 3. Bones of the Hand. The carpus or wrist consists of eight small bones of an irregular shape, and disposed in two unequal rows. Those of the upper row are articulated with the bones of the fore-arm, and those of the lower one with the metacarpus. The ancient anatomists described these bones numerically; Lyserus seems to have been the first who gave to each of them a par- ticular name. The names he adopted are founded on the figure of the bones, and are now pretty generally received, except the first, which instead of ».ti,x.( divides to form the lateral sinuses, ggg, The lambdoid suture, h, The left squamous su- ture, i, The parietal bone- Fig. 4. The anterior surface of the Ossa Nasi. A, The upper part, which joins the os fron- tis. B, The under end, which joins the car- tilage of the nose. C, The inner edge, where they join each other. Fig. 5. The posterior surface of the Ossa Nasi. A A, Their cavity, which forms part of the arch of the nose. B B, Their ridge or spine, which projects a little to be fixed to the fore- part of the septum narium. Fig. 6. The external surface of the Os Max- illare Superius of the left side. A, The nasal process. B, The orbitar plate. C, The unequal surface which joins the os malae. D, The external orbitar hole. E, The opening into the nostril. F, The pa- Osteology, 149 late-plate. G, The maxillary tuberosity. H, Part of the os palati. I, The two dentes inci- sores. K, The dens caninus. L, The two small dentes molares. M, The three large dentes molares. Fig. 7. The internal surface of the Os Max- illare Superius and Os Palati. A, The nasal process. B B, Eminences for the connection of the os spongiosum inferius. D, The under end of the lachrymal groove. E, The antrum maxillare. F, The nasal spine, between which and B is the cavity of the nos- tril. G, The palate-plate. H, The orbitar part of the os palati. I, The nasal plate. K, The suture which unites the maxillary and pa- late bones. L, The pterygoid process of the palate bones. Fig. 8. The external surface of the right Os Unguis. A, The orbitar part. B, The lachrymal part. C, The ridge between them. Fig. 9. The internal surface of the right Os Unguis. This side of the bone has a furrow opposite to the external ridge; all behind that is irre- gular, where it covers part of the ethmoidal cells. 150 Osteology. Fig. 10. The external surface of the left Os Malje. A, The superior orbitar process. B, The inferior orbitar process. C, The malar pro- cess. D, The zygomatic process. E, The orbitar plate. F, A passage for small vessels into or out of the orbit. Fig. 11. The internal surface of the left Os Mal^. A, The superior orbitar process. B, The inferior orbitar process. C, The malar pro- cess. D, The zygomatic process. E, The internal orbitar plate or process. Fig. 12. The external surface of the right Os Spongiosum Inferius. A, The anterior part. B, The hook-like process for covering part of the antrum maxil- lare. C, A small process which covers part of the under end of the lachrymal groove. D, The inferior edge turned a little outwards. Fig. 13. The internal surface of the Os Spon- giosum Inferius. A, The anterior extremity. B, The upper edge which joins the superior maxillary and palate bones. Fig. 14. The posterior and external surface of the right Os Palati. A, The orbitar process. B, The nasal la- mella. C, The pterygoid process. D, The palate process. Osteology. 151 Fig. 15. The anterior and external surface of the right Os Palati. A, The orbitar process. B, An opening through which the lateral nasal vessels and nerves pass. C, The nasal lamella. D, The pterygoid process- E, The posterior edge of the palate process for the connection of the velum palati. F, The inner edge by which the two ossa palati are connected. Fig. 16- The right side of the Vomer, A, The upper edge which joins the nasal lamella of the ethmoid bone and the middle cartilage of the nose. B, The inferior edge, which is connected to the superior maxillary and palate bones. C, The superior and pos- terior part which receives the processus azy- gos of the sphenoid bone. Fig. 17- The Maxilla Inferior. A, The chin- B, The base and left side. C, The angle. D, The coronoid process. E? The condyloid process. F, The beginning of the inferior maxillary canal of the right side, for the entry of the nerve and blood-vessels. G, The termination of the left canal- H, The two dentes incisores. I, The dens caninus. K, The two small molares. L, The three large molares. Fig. 18. The different classes of the Teeth. 1, 2, Afore and back view of the two ante- rior dentes incisores of the lower jaw. 3, 4. 152 Osteology. Similar teeth of4 the upper jaw. 5, 6, A fore and back view of the dentes canini. 7, 8, The anterior dentes molares. 9, 10, 11, The posterior dentes molares. 12, 13, 14, 15, 16, Unusual appearances in the shape and size of the teeth. Fig, 19. The external surface of the Os Hy oides. A, The body. B B, The cornua. C C, The appendices. Plate XXII. Fig. 1. A Posterior View of the Sternum and Clavicles, with the ligament connect- ing the clavicles to each other. a, The posterior surface of the sternum. b b, The broken ends of the clavicles, c c c c, The tubercles near the extremity of each cla- vicle, d, The ligament connecting the cla- vicles. Fig. 2. A Fore-view of the Left Scapula, and of a half of the Clavicle, with their , Ligaments- a, The spine of the scapula, b, the acro- mion, c, The inferior angle- d, Inferior cos- i ta. e, Cervix, f, Glenoid cavity, covered with cartilage for the arm-bone, g g, The capsular ligament of the joint- h, Coracoid Osteology. 153 process, i, The broken end of the clavicle. k, Its extremity joined to the acromion. 1, A ligament coming out single from the acromion to the coracoid process, m, A ligament com- ing out single from the acromion, and dividing into two, which are fixed to the coracoid pro- cess- Fig. 3. The Joint of the Elbow of the Left Arm, with the Ligaments. a, The os humeri, b, Its internal condyle. c c, The two prominent parts of its trochlea appearing through the capsular ligament, d, The ulna, e, The radius, f, The part of the ligament including the head of the radius. Fig. 4. The Bones of the Right-Hand, with the Palm in view. a, The radius, b, The ulna, c, The sca- phoid bone of the carpus, d, The os lunare. e, The os cuneiforme. f, The os pisiforme. g, Trapezium, h, Trapezoides. i, Capita- turn, k, Unciforme. 1, The four metacar- pal bones of the fingers, m, The first pha- lanx, n, The second phalanx, o, The third phalanx, p, The metacarpal bone of the thumb, q, The first joint, r, The second joint. Fig. 5. The Posterior View of the Bones of the Left Hand- The explication of Fig. 4. serves for this figure ; the same letters pointing out the same bones, though in a different view. U 154 Osteology, Fig. 6. The Upper Extremity of the Tibia, with the Semilunar Cartilages of the Joint of the Knee, and some Ligaments. a, The strong ligament which connects the rotula to the tubercle of the tibia, b b, The parts of the extremity of the tibia, co- vered with cartilage, which appear within the semilunar cartilages, c c, The semilunar car- tilages, d, The two parts of what is called the cross ligament. Fig. 7- The Posterior View of the Joint of the Right Knee. a, The os femoris cut- b, Its internal con- dyle, c, Its external condyle, d, The back part of the tibia, e, The superior extremity of the fibula, f, The edge of the internal se*- milunar cartilage, g, An oblique ligament. h, A larger perpendicular ligament, i, A li- gament connecting the femur and fibula. Fig. 8- The Anterior View of the Joint of the Right Knee. b, The internal condyle- c, Its external condyle, d, The part of the os femoris, on which the patella moves- e, A perpendicu- lar ligament, f f, The two parts of the cru- cial ligaments, g g, The edges of the two moveable semilunar cartilages, h, The tibia. i, The strong ligament of the patella, k, The back part of it where the fat has been dis- sected away. 1, The external depression, m, The internal one. n, The cut tibia. Osteology. 155 Fig. 9. A View of the inferior part of the Bones of the Right Foot- ,, a, The great knob of the os calcis. b, A prominence on its outside, c, The hollow for the tendons, nerves, and blood-vessels, d, The anterior extremity of the os calcis. e, Part of the astragalus, f, Its head covered with cartilage, g, The internal prominence of the os naviculare. h, The os cuboides. i, The os cuneiforme internum ; k,—Medium ; 1,—Externum, m, The metatarsal bones of the four lesser toes, n, The first—o, The second—p, The third phalanx of the four les- ser toes, q, The metatarsal bones of the great toe. r, Its first—s, Its second joint. Fig. 10- The Inferior Surface of the two large Sesamoid Bones, at the first Joint of the Great Toe. Fig. 11. The Superior View of the Bones of the Right Foot. a, b, as in Fig- 9. c, The superior head of the astragalus, d, &c. as in Fig. 9. Fig. 12. The View of the Sole of the Foot, with its Ligaments. a, The great knob of the os calcis. b, The hollow for the tendons, nerves, and blood- vessels, c, The sheaths of the flexores pol- licis and digitorum longi opened, d, The strong cartilaginous ligament supporting the head of the astragalus, e, h, Two ligaments which unite into one, and are fixed to the 156 Osteology. metatarsal bone of the great toe. f, A liga- ment from the knob of the os calcis to the metatarsal bone of the little toe. g, A strong triangular ligament, which supports the bones of the tarsus, i, The ligaments of the joints of the five metatarsal bones. Fig. 13. a, The head of the thigh bone of a child, b, The figamentum rotundum con- necting it to the acetabulum, c, The capsu- lar ligament of the joint with its arteries in- jected, d, The numerous vessels of the mu- cilaginous gland injected. Fig. 14. The Back-view of the Cartilages of the Larynx, with the Os Hyoides. a, The posterior part of the base of the os hyoides. b b, Its cornua. c, The appendix of the right side, d, A ligament sent out from the appendix of the left side, to the sty- loid process of the temporal bone, e, The union of the base with the left cornu. f f, The posterior sides of (g) the thyroid carti- lage, h h, Its superior cornua. i i, Its in- ferior cornua. k, The cricoid cartilage. 11, The arytenoid cartilages, m, The entry into the lungs, named glottis, n, The epiglottis. o o, The superior cartilages of the trachea. p, Its ligamentous back part. Fig. 15. The Superior Concave surface of the Sesamoid Bones at the first joint of the Great Toe, with their Ligaments. a, Three sesamoid bones. b, The liga- mentous substance in which they are formed. Of the Integuments, &c. 157 Part II. OF THE SOFT PARTS IN GE- NERAL ; Of the Common Integuments, with their appen- dages ; and of the Muscles. ANATOMICAL writers usually proceed to a description of the muscles after having finished the osteology; but we shall deviate a little from the common method, with a view to describe every thing clearly and dis- tinctly, and to avoid a tautology which would otherwise be unavoidable. All the parts of the body are so intimately connected with each other, that it seems impossible to convey a just idea of any one of them, without being in some measure obliged to say something of others; and on this account we wish to mention in this place the names and situation of the principal viscera of the body, that when mention is hereafter made of any one of them in the course of the work, the reader may at least know where they are placed. After this little digression, the common in- teguments, and after them the muscles will be described; we then -propose to enter into an examination of the several viscera and their different functions. In describing the brain. 158 Of the Integuments, &c. occasion will be taken to speak of the nerves and animal spirits. The circulation of the blood will follow the anatomy of the heart, and the secretions and other matters will be intro- duced in their proper places. The body is divided into three great cavities. Of these the uppermost is formed by the bones of the cranium, and incloses the brain and cere- bellum. The second is composed of the vertebrae of the back, the sternum, and true ribs, with the additional assistance of muscles, membranes, and common integuments, and is called the thorax—It contains the heart and lungs. The third, and inferior cavity, is the abdo- men. It is separated from the thorax by means of the diaphragm, and is formed by the lumbar vertebrae, the os sacrum, the os- sa innominata, and the false ribs, to which we may add the peritonaeum, and a variety of muscles. This cavity incloses the stomach, intestines, omentum or cawl, liver, pancreas, spleen, kidneys, urinary bladder, and parts of generation. Under the division of common integuments are usually included the epidermis, or scarf- skin, the reticulum mucosum of Malpighi, the cutis or true skin, and the membrana adi- posa—The hair and nails, as well as the se- baceous glands, may be considered as appen- dages to the skin. Of the Integuments, csft. 159 Sect. I. Of the Skin. 1. Of the ScART-skin. The epidermis, cuticula, or scarf-skin, is a fine, transparent, and insensible pellicle, des- titute of nerves and blood-vessels, which in- vests the body, and every where covers the true skin. This scarf-skin, Which seems to be very simple, appears, when examined with a microscope, to be composed of several la- minae or scales which are increased by pres- sure, as we may observe in the hands and feet, where it is frequently much thickened, and becomes perfectly callous. It seems to adhere to the cutis by a number of very mi- nute filaments, but may easily be separated from it by heat, or by maceration in water- Some anatomical writers have supposed that it is formed by a moisture exhaled from the whole surface of the body, which gradually hardens when it comes into contact with the air. They were perhaps induced to adopt this opinion, by observing the speedy regeneration of this part of the body when it has been by any means destroyed, it appearing to be renewed on all parts of the surface at the same time; where- as other parts which have been injured, are found to direct their growth from their circum- ference only towards their centre. But a demon- strative proof that the epidermis is not a fluid 160 Of the Integuments, &fc. hardened by means of the external air, is that the foetus in utero is found to have this cover- ing. Lieuwenhocck supposed its formation to be owing to the expansion of the extremities of the excretory vessels which are found eve- ry where upon the surface of the true skin. Ruysch attributed its origin to the nervous papillae of the skin; and Heister thinks it pro- bable, that it may be owing both to the pa- pillae and the excretory vessels. The cele- brated Morgagni, on the other hand, contends,* that it is nothing more than the surface of the cutis, hardened and rendered insensible by the liquor amnii in vitero, and by the pressure of the air. This is a subject, however, on which we can advance nothing with certainty. The cuticle is pierced with an infinite num- ber of pores or little holes, which afford a passage to the hairs, sweat, and insensible per- spiration, and likewise to warm water, mer- cury, and whatever else is capable of being taken in by the absorbents of the skin. The lines which we observe on the epidermis be- long to the true skin. The cuticle adjusts it- self to them, but does not form them. 2. Of the Rete Mucosum. Between the epidermis and cutis we meet with an appearance to which Malpighi, who first described it, gave the name of rete muco- sum, supposing it to be of a membranous structure, and pierced with an infinite number of pores; but the fact is, that it seems to be * Adversar. Anat. u. Animadver. 2. Of the Integuments, &c. 161 nothing more than a mucous substance which may be dissolved by macerating, it in water, while the cuticle and cutis preserve their tex- ture. The colour of the body is found to depend on the colour of this rete mucosum ; for in ne- groes it is observed to be perfectly black, whilst the true skin is of the ordinary colour. The blisters which raise the skin when burnt or scalded, have been supposed by some to be owing to a rarefaction of this mucus ; but they are more probably occasioned by an increased action of the vessels of the part, together with an afflux and effusion of the thinner parts of the blood- 3. Of the Cutis, or True Skin. The cutis is composed of fibres closely compacted together, as we may observe in lea- ther, which is the prepared skin of animals. These fibres form a thick net-work, which every where admits the filaments of nerves, and an infinite number of blood-vessels and lymphatics. The cutis, when the epidermis is taken off, is found to have, throughout its whole sur- face, innumerable papillae, which appear like very minute granulations, and seem to be cal- culated to receive the impressions of the touch, being the most easily observed where the sense of feeling is the most delicate, as in the palms of the hands and on the fingers. X 162 Of the Integuments, &c. These papillae are supposed by many ana- tomical writers to be continuations of the pul- py substance of nerves, whose coats have ter- minated in the cellular texture of the skin. The great sensibility of these papillae evi- dently proves them to be exceedingly nervous; but surely the nervous fibrellae of the skin are of themselves scarcely equal to the formation of these papillae, and it seems to be more pro- bable that they are formed like the rest of the cutis. These papillae being described, the uses of the epidermis and the reticulum mucosum will be more easily understood; the latter serving to keep them constantly moist, while the for- mer protects them from the external air, and modifies their too great sensibility. 4. Of the Glands of the Skin. In different parts of the body we meet, within the substance of the skin, with certain glands or follicles, which discharge a fat and oily humour that serves to lubricate and soft- en the skin. When the fluid they secrete has acquired a certain degree of thickness, it approaches to the colour and consistence of suet; and from this appearance they have de- rived their name of sebaceous glands. They are found in the greatest number in the nose, ear, nipple, axilla, groin, scrotum, vagina, and prepuce. Besides these sebaceous glands, we read, in anatomical books, of others that are de- Of the Integuments, &c. 163 scribed as small spherical bodies placed in all parts of the skin, in much greater abundance than those just now mentioned, and named mi- liary, from their supposed resemblance to mil- let-seed. Steno, who first described these glands, and Malpighi, Ruysch, Verheyen, Windslow, and others, who have adopted his opinions on this subject, speak of them as hav- ing excretory ducts, that open on the surface of the cuticle, and distil the sweat and mat- ter of insensible perspiration ; and yet, not- withstanding the positive manner in which these pretended glands have been spoken of, we are now sufficiently convinced that their existence is altogether imaginary. 5. Of the Insensible Perspiration and Sweat. The matter of insensible perspiration, or in other words, the subtile vapour that is continu- ally exhaling from the surface of the body, is not secreted by any particular glands, but seems to be derived wholly from the extremities of the minute arteries that are everywhere dis- persed through the skin. These exhaling ves- sels are easily demonstrated in the dead sub- ject, by throwing water into the arteries ; for then small drops exude from all parts of the skin, and raise up the cuticle, the pores of which are closed by death ; and in the living subject, a looking-glass placed against the skin, is soon obscured by the vapour. Bidloo fancied he had discovered ducts leading from the cutis to 164 Of the Integuments, &c. the cuticle, and transmitting this fluid ; but in this he was mistaken. When the perspiration is by any means in- creased, and several drops that were insensi- ble when separate, are united together and con- densed by the external air, they form upon the skin small, but visible, drops called sweat.* This particularly happens after much exercise, or whatever occasions an increased determin- ation of fluids to the surface of the body; a greater quantity of perspirable matter being in such cases carried through the passages that are destined to convey it off. It has been disputed, indeed, whether the insensible perspiration and sweat are to be considered as one and the same excretion, dif- fering only in degree; or whether they are two distinct excretions derived from different sources. In support of the latter opinion, it has been alleged, that the insensible perspi- ration is agreeable to nature, and essential to health, whereas sweat may be considered as a species of disease. But this argument proves nothing ; and it seems probable, that both the insensible vapour and the sweat are exhaled in a similar manner, though they differ in quantity, and probably in their qualities ; the former being more limpid, and seemingly less impregnated with salts than the latter: at any rate we may consider the skin as an emuncto- ry through which the redundant water, and * Lieuwenhoeck asserts that one drop of sweat is formed by the conflux of fifteen drops of perspirable vapour. Of the Integuments, &c. 165 sometimes the other more saline parts of the blood, are carried off. But the insensible per- spiration is not confined to the skin only—a great part of what we are constantly throwing off in this way is from the lungs. The quantity of fluid exhaled from the human body by this insensible perspiration is very considerable. Sanctorius* an Italian physician, who indefati- gably passed a great many years in a series of sta- tical experiments, demonstrated long ago what has been confirmed by later observations, that the quantity of vapour exhaled from the skin and from the surface of the lungs, amounts nearly to 5-8ths of the aliment we take in. So that if in the warm climate of Italy a person eats and drinks the quantity of eight pounds in the course of a day, five pounds of it will pass off by insensible perspiration, while three pounds only will be evacuated by stool, urine, saliva, &c. But in countries where the degree of cold is greater than in Italy, the quantity of perspired matter is less; in some of the more northern climates, it being found not to equal the discharge by urine. It is likewise observ- * The insensible perspiration is sometimes distinguished by the name of this physician, who was born in the territories of Venice, and was afterwards a professor in the university of Pa- dua. After estimating the aliment he took in, and the sensible secretions and discharges, he was enabled to ascertain with great accuracy the weight or quantity of insensible perspiration by means of a statical chair which he contrived for this purpose; and from his experiments, which were conducted with great industry and patience, he was led to determine what kinds of solid or li- quid aliment increased or diminished it. From these experi- ments he formed a system, which he published at Venice in 1614, in the form of aphorisms, under the title of " Ars dc Medicina Statica." 166 Of the Integuments, £jfc. ed to vary according to the season of the year, and according to the constitution, age, sex, diseases, diet, exercise, passions, &C. of different people. From what has been said on this subject, it will be easily conceived, that this evacuation cannot be either much increased or diminish- ed in quantity without affecting the health. The perspirable matter and the sweat are in some measure analogous to the urine, as ap- pears from their taste and saline nature.* And it is worthy of observation, that when either of these secretions is increased in quantity, the other is diminished ; so that they who per- spire the least, usually pass the greatest quan- tity of urine, and vice versa. 6. Of the Nails. The nails are of a compact texture, hard and transparent like horn. Their origin is still a subject of dispute. Malpighi supposed them to be formed by a continuation of the papillae of the skin: Ludwig, on the other hand, maintained, that they were composed of the extremities of blood-vessels and nerves; both these opinions are now deservedly rejected. They seem to possess many properties in common with the cuticle; like it they are nei- ther vascular nor sensible, and when the cuti- * Minute chrystals have been observed to shoot upon the clothes of men who work in glass-houses. Haller Elem. Phys. Of the Integuments, &c, 167 cle is separated from the true skin by macera- tion or other means, the nails come aw< y with it. They appear to be composed of different layers, of unequal size, applied one over the other. Each layer seems to be formed of lon- gitudinal fibres. In each nail we may distinguish three parts, viz. the root, the body or middle, and the ex- tremity. The root is of a soft, thm, and white substance, terminating in the form of a cres- cent ; the epidermis adheres very strongly to this part; the body of the nail is broader, red- der, and thicker, and the extremity is of still greater firmness. The nails increase from their roots, and not from their upper extremity. Their principal use is to cover and defend the ends of the fingers and toes from external injury. 7. Of the Hair. The hairs, which from their being general- ly known do not seem to require any definition, arise from distinct capsules or bulbs seated in the cellular membrane under the skin.* Some * Malpighi, and after him the celebrated Ruysch, supposed the hairs to be continuations of nerves, being of opinion that they originated from the papillae of the skin, which they consi- dered as nervous; and as a corroborating proof of what they advanced, they argued the pain we feel in plucking them out; but later anatomists seem to have rejected this doctrine, and con- sider the hairs as particular bodie-, not arising from the papillae (for in the parts where the papilla: abound most there are n0 hairs,) but from bulbs or capsules, which are peculiar to them. 168 Of the Integuments, &c. of these bulbs inclose several hairs. They may be observed at the roots of the hairs which form the beard or whiskers of a cat. The hairs, like the nails, grow only from below by a regular propulsion from their root, where they receive their nourishment. Their bulbs, when viewed with a microscope, are found to be of various shapes. In the head and scrotum they are roundish; in the eye- brows they are oval; in the other parts of the body they are nearly of a cylindrical shape. Each bulb seems to consist of two membranes, between which there is a certain quantity of moisture. Within the bulb the hair separates into three or four fibrillae; the bodies of the hairs, which are the parts without the skin, vary in softness and colour according to the difference of climate, age, or temperament of body.* Their general use in the body does not seem to be absolutely determined; but hairs in par- ticular parts, as on the eye-brows and eye-lids, are destined for particular uses, which will be mentioned when those parts are described. 8. Of the Cellular Membrane and Fat. The cellular membrane is found to invest the most minute fibres we are able to trace; so that by modern physiologists, it is very pro- * The hairs differ likewise from each other, and may not be improperly divided into two classes ; one of which may include the hair of the head, chin, pubes, and axillse; and the other, the softer hairs, which are to be observed almost every where on the surface of die body. Of the Integuments, &c. 169 perly considered as the universal connecting medium of every part of the body. It is composed of an infinite number of mi- nute cells united together, and communicating with each other. The two diseases peculiar to this membrane are proofs of such a commu- nication ; for in the emphysema all its cells are filled with air, and in the anasarca they are universally distended with water. Besides these proofs of communication from disease, a fa- miliar instance of it may be observed among butchers, who usually puncture this membrane, and by inflating it with air add to the good ap- pearance of their meat. The cells of this membrane serve as re- servoirs to the oily part of the blood or Fat, which seems to be deposited in them, either by transudation through the coats of the ar- teries, that ramify through these cells, or by particular vessels, continued from the ends of arteries. These cells are not of a glandular structure, as Malpighi and others after him have supposed. The fat is absorbed and car- ried back into the system by the lymphatics. The great waste of it in many diseases, par- ticularly in the consumption, is a sufficient proof that such an absorption takes place. The fulness and size of the body are in a great measure proportioned to the quantity of fat contained in the cells of this membrane. In the living body it seems to be a fluid oil, which concretes after death. In graminivo- rous animals, it is found to be of a firmer con- sistence than in man. Y 170 Of the Muscles. The fat is not confined to the skin alone, being met with every where in the interstices of muscles, in the omentum, about the kid- neys, at the basis of the heart, in the orbits, The chief uses of the fat. seem to be to afford moisture to all the parts with which it is connected; to facilitate the action of the muscles ; and to add to the beauty of the body, by making it every where smooth and equal. Sect. II. Of the Muscles. The muscles are the organs of motion. The parts that are usually included under this name consist of distinct portions of flesh, suscepti- ble of contraction and relaxation ; the motions of which in a natural and healthy state, are subject to the will, and for this reason they are called voluntary muscles. But besides these, there are other parts of the body that owe their power of contraction to their mus- cular fibres ; thus the heart is of a muscular texture, forming what is called a hollow mus- cle ; and the urinary bladder, stomach, intes- tines, &c. are enabled to act upon their con- tents, merely because they are provided with muscular fibres. These are called involuntary muscles, because their motions are not depen- dent on the will. The muscles of respiration, being in some measure influenced by the wiu\ are said to have a mixed motion. Of the Muscles. 171 The names by which the voluntary muscles are distinguished, are founded on their size, figure, situation, use, or the arrangement of their fibres, or their origin and insertion. But besides these particular distinctions, there are certain general ones that require to be noticed. Thus, if the fibres of a muscle are placed parallel to each other in a straight direction, they form what are styled a rectilinear muscle ; if the fibres cross and intersect each other, they constitute a compound muscle ; a radiated one, if the fibres are disposed in the manner of rays; or a penniform muscle, if, like the plume of a pen, they are placed obliquely with re- spect to the tendon. Muscles that act in opposition to each other, are called antagonista; thus every extensor muscle has a flexor for its antagonist, and vice versa. Muscles that concur in the same ac- tion are styled congeheres. The muscles being attached to the bones, the latter may be considered as levers that are moved in different directions by the contrac- tion of those organs. The end of a muscle which adheres to the most fixed part is usually called the origin, and that which adheres to the more moveable part, the insertion, of the muscle. In every muscle we may distinguish two kinds of fibres ; the one soft, of a red colour, sensible and irritable, edAXed fleshy fibres; the other of a firmer texture, of a white glisten- ing colour, insensible, without irritability or the power of contracting, and named tendinous 172 Of the Muscles. fibres. They are occasionally intermixed ; but the fleshy fibres generally prevail in the belly or middle part of a muscle, and the tendi- nous ones in the extremities. If these tendi- nous fibres are formed into a round slender chord, they form what is called the tendon of the muscle; on the other hand, if they are spread into a broad flat surface, the extremity of the muscle is styled aponeurosis. The tendons of many muscles, especially when they are long and exposed to pressure or friction in the grooves formed for them in the bones, are surrounded by a tendinous sheath or fascia, in which we sometimes find a small mucous sac or bursa mucosa, which obviates any inconvenience from friction. Sometimes we find whole muscles, and even several mus- cles, covered by a fascia of the same kind, that affords origin to many of their fibres, dip- ping down between them, adhering to the ridges of bones, and thus preventing them from swelling too much when in action. The most remarkable instance of such a covering is the fascia lata of the thigh. Each muscle is inclosed by a thin covering of cellular membrane, which has been some- times improperly considered as peculiar to the muscles, and described under the name of pro- pria membrana musculosa. This cellular co- vering dips down into the substance of the muscle, connecting and surrounding the most minute fibres we are able to demonstrate, and affording a support to their vessels and nerves. Lieuwenhoeck fancied he had discovered, by Of the Muscles, 173 means of his microscope, the ultimate division of a muscle, and that he could point out the sim- ple fibre, which appeared to him to be an hun- dred times less than a hair; but he was after-- wards convinced how much he was mistaken on this subject, and candidly acknowledged, that what he had taken for a simple fibre was in fact a bundle of fibres. It is easy to observe several of these fasci- culi or bundles in a piece of beef, in which, from the coarseness of its texture, they are very evident. The red colour which so particularly distin- guishes the muscular or fleshy parts of ani- mals, is owing to an infinite number of blood- vessels that are dispersed through their sub- stance. When we macerate the fibres of a muscle in water, it becomes of a white colour like all other parts of the body divested of their blood. The blood-vessels are accompa- nied by nerves, and they are both distributed in such abundance to these parts, that in endea- vouring to trace the course of the blood-ves- sels in a muscle, it would appear to be formed altogether by their ramifications ; and in an attempt to follow the branches of its nerves, they would be found to be equal in propor- tion. If a muscle is pricked or irritated, it im- mediately contracts. This is called its irrita- ble principle ; and this irritability is to be con- sidered as the characteristic of muscular fibres, and may serve to prove their existence in parts that are too minute to be examined by the eye. 174 Of the Muscles. This power, which disposes the muscles to contract when stimulated, independent of the will, is supposed to- be inherent in them ; and is therefore named vis insita. This property is not to be confounded with elasticity, which the membranes and other parts of the body possess in a greater or less degree in common with the muscles; nor with sensibility, for the heart, though the most irritable, seems to be the least sensible of any of the muscular parts of the body. After a muscular fibre has contracted, it soon returns to a state of relaxation, till it is excited afresh, and then it contracts and re- laxes again. We may likewise produce such a contraction, by irritating the nerve leading to a muscle, although the nerve itself is not affected. This principle is found to be greater in small than in large, and in young than in old, animals. In the voluntary muscles these effects of con- traction and relaxation of the fleshy fibres are produced in obedience to the will, by what may be called the vis nervosa, a property that is not to be confounded with the vis insita. As the existence of a vis insita different from a vis nervea, was the doctrine taught by Doctor Haller in his Elem. Phys. but is at present called in question by several, particularly Doc- tor Monro, we think it necessary to give a few objections, as stated in his Observations on the Nervous System: Of the Muscles. 175 " The chief experiment (says the Doctor) which seems to have led Dr. Haller to this opinion, is the well known one, that the heart and other muscles, alter being detached from the brain, continue to act spontaneously, or by stimuli may be roused into action for a con- siderable length of time ; and when it cannot be alleged, says Dr. Haller, that the nervous fluid is by the mind, or otherwise, impelled into the muscle. " That in this instance, we cannot compre- hend by what power the nervous fluid or ener- gy can be put in motion, must perhaps be granted: But has Dr. Haller given a better explanation of the manner in which his sup- posed vis insita becomes active ? " If it be as difficult to point out the cause of the action of the vis insita as that of the action of the vis nervea, the admission of that hew power, instead of relieving, would add to our perplexity. " We should then have admitted, that two causes of a different nature were capable of producing exactly the same effect; which is not in general agreeable to the laws of nature. " We should find other consequences arise from such an hypothesis, which tend to weak- en the credibility of it. For instance, if in a sound animal the vis nervea alone produces the contraction of the muscles, we will ask what purpose the vis insita serves ? If both operate, are we to suppose that the vis nervea, impelled by the mind or living principle, gives the order, which the vis insita executes, and 176 Of the Muscles. that the nerves are the internuntii; and so ad- mit two wise agents employed in every the most simple action ? But instead of speculating far- ther, let us learn the effect of experiments, and endeavour from these to draw plain conclu- sions. " 1. When I poured a solution of opium in water under the skin of the leg of a frog, the muscles, to the surface of which it was appli- ed, were very soon deprived of the power of contraction. In like manner, when I poured this solution into the cavity of the heart, by opening the vena cava, the heart was almost instantly deprived of its power of motion, whe- ther the experiment was performed on it fixed in its place, or cut out of the body. u 2. I opened the thorax of a living frog; and then tied or cut its aorta, so as to put a stop to the circulation of its blood. " I then opened the vena cava, and poured the solution of opium into the heart; and found, not only that this organ was instantly deprived of its powers of action, but that in a few minutes the most distant muscles of the limbs were extremely weakened. Yet this weakness was not owing to the want of circu- lation, for the frog could jump about for more than an hour after the heart was cut out. " In the first of these two experiments, we observed the supposed vis insita destroyed by the opium; in the latter, the vis nervea; for it is evident that the limbs were affected by the sympathy of the brain, and of the nervous Of the Muscles. 177 system in general, with the nerves of the heart. 3. When the nerve of any muscle is first divided by a transverse section, and then burnt with a hot iron, or punctured with a nee- dle, the muscle in which it terminates con- tracts violently, exactly in the same manner as when the irritation is applied to the fibres of the muscle. But when the hot iron, or nee- dle, is confined to the nerve, Dr. Haller him- self must have admitted, that the vis nervea, and not the vis insita, was excited. But here I would ask two questions. " First, Whether we do not as well under- stand how the vis nervea is excited when irri- tation is applied to the muscle as when it is ap- plied to the trunk of the nerve, the impelling power of the mind seeming to be equally wanting in both cases ? " Secondly, If it appears that irritation ap- plied to the trunk of a nerve excites the vis nervea, why should we doubt that it can equal- ly well excite it when applied to the small and very sensible branches and terminations of the nerve in the muscle ? " As, therefore, it appears that the suppos- ed vis insita is destroyed or excited by the same means as the vis nervea; nay, that when, by the application of opium to the heart of a frog, after the aorta is cut and the circulation interrupted, we have destroyed the vis insita, the vis nervea is so much extinguished, that the animal cannot act with the distant muscles Z 178 Of the Muscles. of the limb; and that these afterward grow very torpid, or lose mucii of their supposed vis insita ; it seems clearly to follow, that there is no just ground for supposing that any other principle produces the contraction of a mus- cle." The vis nervosa, or operation of the mind, if we may so call it, by which a muscle is brought into contraction, is not inherent in the muscle like the vis insita ; neither is it perpe- tual, like this latter property. After long con- tinued or violent exercise, for example, the vo- luntary muscles become painful, and at length incapable of further actipn ; whereas the heart and other involuntary muscles, the motions of which depend solely on the vis insita, continue through life in a constant state of action, with- out any inconvenience or waste of this inherent principle. The action of the vis nervosa on the volun- tary muscles, constitutes what is called mus- cular motion ; a subject that has given rise to a variety of hypotheses, many of them inge- nious, but none of them satisfactory. Borelli and some others have undertaken to explain the cause of contraction, by suppos- ing that every muscular fibre forms as it were ' a chain of very minute bladders, while the nerves which are distributed through the mus- cle, bring with them a supply of animal spi- rits, which at our will fill these bladders, and by increasing their diameter in width, shorten them, and of course the whole fibre. Of the Muscles. 179 Borelli supposes these bladders to be of a rhomboidal shape; Bernoulli on the other hand contends that they are oval. Our coun- tryman, Cowper, fancied he had filled them with mercury ; the cause of this mistake was probably owing to the mercury's insinuating itself into some of the lymphatic vessels. The late ingenious Mr. Elliot undertook to account for the phenomena of muscular motion on prin- ciples very different from those just now men- tioned. He supposed that a dephlogisticated state of the blood is requisite for muscular ac- tion, and that a communication of phlogiston to the blood is a necessary effect of such action. We know that the muscular fibre is shorten- ed, and that the muscle itself swells when in action ; but how these phenomena are produc- ed, we are unable to determine. We likewise know that the nerves are essential to muscu- lar motion; for upon dividing or making a li- gature round the nerve leading to a muscle, the latter becomes incapable of motion. A li- gature made on the artery of a muscle pro- duces a similar effect; a proof this, that a re- gular supply of blood is also equally necessa- ry to muscular motion. The cause of palsy is usually not to be sought for in the muscle affected, but in the nerve leading to that mus- cle, or in that part of the brain or spinal mar- row from which the nerve derives its origin. 180 Of the Muscles. Of the particular Muscles. As the enumeration and description of the particular muscles must be dry and unenter- taining to the generality of readers, yet can- not be altogether omitted in a work of this nature, it appeared eligible to throw this part of the subject into the form of a table ; in which the name, origin, insertion, and princi- pal use of each muscle, will be found describ- ed in few words, and occasionally its etymolo, gy when it is of Greek derivation or difficult to be understood. A TABLE of the MUSCLES, arranged according to their Situation. [N. B. This table does not include all the muscles of the body; thofe belonging to the eyes, internal car, intestinum rectum, and the male and female organs of generation, being described in other parts of the work. The reader will be pleased to observe likewise, that although all the muscles (a few only excepted) are in pairs, mention is here made only of the muscles of one side.] Muscles situated unJtr the integu- m n s of the cra- miun • - - i Name. Occipito frontalis. 2. Corrugator super- cihi. Origin. From the transverse ridge of the os oc- cipitis. From above the join- ing of the os fron- tis, os nasi, and os maxillare. Insertion. Into the skin of the eye-brows. Into the inner part of the occipito- frontalis. Use. To pull the skin of the head back- wards, and to raise the eye-brows and skin of the fore- head. To draw the eye- brows towards each other, and to wrin- kle the forehead. Name. ------------of the eye-lids I. Orbicularis palpe- brarum. 2. Levator palpebral superioris. -------—-of the external ear i. Attollens auricu- 1a.m. r: 2. Anterior auriculx. 3. Retrahentes ( *) au- riculx. Musclbs of the car- tilages of the ear i. Tragicus. Origin. From around the edge of the orbit. From the bottom of the orbit, near the optic foramen. From the tendon of the occipito fron- talis near the os temporis. From near the back part of the zygoma. From the outer and back part of the root of the mastoid process. From the outer and middle part of the concha, near the tragus. Insertion, Use, Into the nasal pre- To shut the eye. cess of the os maxillare. Into the cartilage of To open the eye, the upper eye-lid. Into the upper part To raise the ear. of the ear. 09 to Into an eminence be- hind the helix. Into the convex part of the concha. To raise this emi- nence, and to pull it forwards. To stretch the con- cha, and pull the ear backwards. Into the upper part To depress the con- of the tragus'. cha, and pull the point of the tragus a little outwardi, to * These *re three small ilcndcr muscles. The inferior one i» sometime* wanting -ofthenose, Name. i. Antitragicus. 3. Transversus-auri- culae. 4. Helicis major. Heli icis minor, 1. Compressor naris. (*) Origin. From the root of the inner part of the helix. From the upper part of the concha. From theupper, ante- rior, and acute part of the he'ix. Frcm the lower and fore part of the he- lix. From the outer part of the root of the ala nasi. Insertion. Use. Into the upper part To dilate the mouth of the anti-tragus. of the concha. Into the inner part of the helix. Into the cartilage of the helix, a little above the tragus. Into the helix, near the fissure in its cartilage. Into the naval pro- cess of the os max- illare, and anterior extremity of the os nasi. To stretchthe conclia and scapha, and likewise to pull the parts it is connect- ed with towards each other. To depress the upper part of the helix. To ccntract the fis- sure. To straighten the nos- trils, and likewise to conugate the skin of the nose. Co * The nose is affected by fibres of the occipito frontalis, and by several muscles of the face \ but this pair, the compressores, is the qq only one that is proper to it. C3 Name. .----------of the mouth and lips, i. Levator labii supe- riors, alasque nasi. 2. Levator anguli oris. 3. Zygomaticus ma- jor. 4. Zygomaticus mi- nor. c. Buccinator. Origin. From the outer part of the orbitar pro- cess of the os max- illare and from the nasal process of that b me, where it joins the os frontis. From the os maxil- lare superius, be- tween the orbitar foramen and the first dens molaris. From the os malas near the zygoma- tic suture. Immediately above the origin of the zyg. major. From the alveoli of the dentes molares in the upper and lower jaws. Insertion. Into the upper lip and alaof theno'-.e. Into the orbicularis oris at the angle of the mouth. Into the angle of the mouth. Into the angle of the mouth. Into the angle of the mouth. Use. To draw the upper lip and skin of the nose upwards and outwards. To raise the corner of the mouth. To raise the angle of the mouth, and make the cheek prominent, as in laughing. To raise the angle of the mouth oblique- ly outwards. To contract the mouth and draw the angle of it outwards and backwards. & Name. Depressor labii su- per i oris, alacque nasi. Depressor oris. anguli 8. Depressor labii in- ferioris. Levator labii infe- rioris. Origin. From the os maxill. super, immediately above the gums of the dentes incisores. At the side of the chin from the low- er edge of the max- illa inferior. From the lower and anterior part of the maxilla inferi- or. From near the gums of the incisores and caninusof the max- illa inferior. Insertion. Into the root of the ala nasi and upper lip. Into the angle of the mouth. Into the under lip. Into the under lip and skin of the chin. Use. To draw the ala nasi and upperlip down- wards. To draw the corner of the mouth down- wards. To draw the under lip downwards and somewhat outwards To raise the under lip and skin of the chin. 5t Co <"> ST" Co jo. Orbicularis ris. (*) O- To shut the mouth by constraining the lips. (*) This muscle is in a great measure, if not wholly, formed by the buccinator, tygomatici, depressores, Mid other muscles that move the hps. Its fibres surround the mouth like a ring. CO fyfuscLEs of the low- er jaw, - - - Name. i. Temporalis, 2. Masseter.(f) Origin. From part of the os bregmatis and os frontis; squamous part of the os tem- poris ; back part of the os make, and the temporal pro- cess of the os sphe- noides. (*) From the malar pro- cess of the os max- illare, and the low- er edges of the os malae. and of the zygomatic process of the os temporis. Insertion. Use. Intothe coronoid pro-, To move the lower cess of the lower jaw upwards. jaw. coronoid process, and that part of the j aw which supports that and the con- dyloid process. to move the jaw a little forwards and backwards. c* 3 Into the basis of the To raise and likewise ^ (♦) Some of its fibres likewise have their origin from a strong fascia that covers the muscle and adheres to the bone round the whole circumference of its origin. When we remove this covering, we find the muscle of a semicircular shape with its fibres, con- verging and forming a strong middle tendon. (t) So called from its use in chewing, its derivation being from n*TtT*tt**: manduc», " H eat." Name. 3. Pterygoideus in- ternus. , Pterygoideus ex- ternus. --------situated at the fore part of the neck. 1 .Latissimuscolli. (f) Origin. From the inner sur- face of the outer wing of the ptery- goid process of the os sphenoides, and from the process of the os palati that helps to form the pterygoid fossa. From the external ala of the pterygoid process, a small part ofthe adjacent os maxillare, and a ridge in the ten po- ral process of the os sphenoides. From the cellular membrane covering Insertion. Into the lower jaw on its inner side and near its angle. Use. To raise the lower jdv- and *mw it a little to one side. Into the fore part of To move the jaw for- ££ the condyl id pro cess « f the lower jaw, and likewise of the capsular li- gament. Into the side of the chin and intejm- wards and to the opposite side ; , * ) and at the same time to prevent the ligament of the joint fr"m being pinched. To draw the cheeks and skin of the face (*) This happens when the muscle acts singly. When both act, the jaw is brought horizontally forwards. (+) 'Ihisbioad and thin muscular expansion, which i* situated immediately under the common unc; ur.ents, 15 by Winslow named mucuhu cutaneus. Galen gave it the name of ^«r„^» ^t.s (Flatysma-myoides J ; the etymology of wh.ch is from •fc-T..^ dUatati,, and mvs, musculus, and n&*s> forma. IS"" Co Co. CO Name. Z. Mastoideus (*). Muscles fituated be- tween the trunk and the os hyoides. ,Omo-hyoideus(f). Origin. the pectoral, del- toid, and trapezius muscles. From the upper part of the sternum and from the upper and fore part of the clavicle. From the upper costa of the scapula near its niche; from part of a ligament that extends across this niche, and some- Insertion. ments of the cheek. Into the mastoid pro- cess, and as far back as the lamb- doidal suture. Into the basis of the os hyoides. Use. downwards; and when the mouth is shut, to draw ;ill that part of the skin to which it is con- nected below the lowerjawupwards. To move the head to one side, or when both muscles ;ct,to bend it forwards. To draw the os hyoi- des in an oblique direction down- wards. CO 00 Co <"> Co (•) This, on account of its two origins, is by Albinus described as two distinct muscles, which he names stemo-mastoideus and deido- mastoideus. (■p This muscle does not always arise from the coracoid process, it seems to have been improperly named coraco-byoiJes by .Louglaa and Albinus. Winslow calls it omo-hyoideus, on account of its general origin from the scapula. Name. t. Sterno-hyoideus. 3. Hyo-thyroideus. 4. Sterno-thyroideus. 5. Crico-thyroideus. Origin. Insertion. Use. times by a few fi- bres, from the co- racoid process. From the cartilage of the first rib, the in- ner and upper part of the sternum, and a small part of the clavicle. From part of the ba- sis and horn of the os hyoides. From between the cartilages of the 1st and 2d ribs at the upper and in- ner part of the sternum. From the anterior part and side of the cricoid cartilage. Into the basis of the os hyoides. Into a rough oblique line at the side of the thyroid carti- lage. Immediately under the hyo-thyroi- deus. Into the lower part and inferior horn of the thyroid car- tilage. To draw the os hyoi- des downwards. To raise the thyroid cartilage, or de- press the os hyoi- des. To pull the thyroid cartilage down- wards. To pull the cricoid cartilage upwards and backwards, or the thyroid for- wards and down- wards. situated be- tween the os hyoi- des and lower jaw. i. Diagiasticus (*.) 3.Stylo-hyoideus(f). Mylo-hyoideus (§) Genohyoide- us. Origin, From a fossa at the root of the mastoid process, and like- wise from the os hyoides. From the basis of the styloid process. From the inside of the lower jaw, be- tween the last dens molaris and the chin. From the inside of the chin. Insertion, Into the lower and anterior part of the chin. Into the side and fore part of the os hy- oides near its base. Into the basis of the os hyoides. Into the base of the os hyoides. Use, To draw the lower jaw downwards. To draw the os hyoi- des obliquely up wards. To move the os hyoi- des to either side, forwards or up- wards. To move the os hy- oides forwards or upwards. o 3 Co 6* (*\ From S.c v»™ fUveHterJ because it has two fleshy bellies with a*middle tendon. This tendon passes through the stylo-hyoideus. (t' In some subjects we meet with another muscle, which from its having nearly the same origin, insertion, and use as this, has been Da7*eds?name°d from iu arising near the dentes molares 0«m.J and its being inserted into the os hyoides. (4 i From y*»"3"> mtntum, the " chin-' Name. 5. Genio glossus. 6. Hyo-glosbUs(*). 7. Lingualis. 8. Stylo-glossus. 6 ■tgt Insertion. From •}■■-: inside of Into the tongue and th-.- cnin. basis of the os hy- oides. From the horn, ba- Into the tongue late- sis, and appendix rally. of the os hyoides Laterally from the Into the extremity of root of the tongue. the tongue. 9. Stylo-pharyngxus. From the styl- 'id pro- cess, and some- times also from a ligament that ex- tends from'thence to the angle of the lowr jaw. From the basis of the styloid process. to. Circumflexus-pa- From near the bony lati. part of the Eusta- chian tube, and Into the side of the tongue from the rout to near its tip. Into the side of the pharynx and pos- terior part of the thyroid cartilage. Into the semilunar edge ->f the os pa- lati and the velum Use. To move the t^r-cn;* in various direc- tions To draw the tongue downward and in- v ards. To shorten the tongue and draw it back- wards. To move the tongue backwards and to one side. To raise the thyroid cartilage and pha- r\nx, and likewise to dilate the latter. To dilate and draw the velum oblique- ly downwards. (*) From *'t*it torhu, and >x«.e«», lingua, " the tongue." Name. Qrigin. Insertion. Us, pendulum palati n. Levator palati. Muscles situated about the fauces, i. Pil-ito-pharyn- gseus. Into the velum pen- To pull the velum dulum palati. backwards. Into the upper and posterior part of the thyroid cartilage. from the spinous process of the os sphenoides. From the membra- nous part of the Eustachian tube, and the extremity of the os petrosum. From the lower and anterior part of the cartilaginous extre- mity of the Eusta- chian tube f);the tendinous expan- fion of the circum- flexus palati; and the velum pendu- lum palati near the ba is and back part of the uvula. (*) This muscle in its course forms a round tendon, which, after crossing ovei a kind of hook formed by the inner plate of the pterygoid process of the sphenoid bone, expands into a tendinous membrane. i ) Tht two fibres that arise from the Eustachian tube are described as a distinct muscle by Albinus, under the name of salpingo- .fbaryngeus. They serve to dilate the mouth »f the tube. To raise the pharynx and thyroid carti- lage, or to pull the velum and uvula backwards and downwards. Name. Origin. Insertion. Use. Muscles at the back part of the pharynx. to 2. Constrictor isthmi faucium. 3. Azygos uvulae. Constrictor pha-i ryngis superior. 2. Constrictor pharynx gis medius (J). From near the basis of the tongue late- rally. From the end of the suture that unites the ossa palati. From the cuneiform process- of the oc- cipital bone; the pterygoid process 6f the os sphenoi- des, and from each jaw near the last dens molaris (f). From the horn arid appendix of the os To raise the tongue anddrawthe velum towards it (* ) Into the velum pen- dulum palati, near the basis and fore part of the uvula. Into the extremity of To shorten the uvu- the uvula. la, and bring it forwards and up- wards. Into the middle of To move the pharynx the pharynx. upwards and for- wards, and to com- press its upperpart. Into the middle of To draw the os hyoi- the processus cunei- des and pharynx (*) This muscle, and the palato-pharyngasus, likewise serve to close the passage into the fauces, and to carry the food into the pha- rynx. t The three orders of fibres here mentioned, with a few others derived from the tongue, have given occasion to Douglas to de- scribe them as four distinct muscles, under the names of uphalo-pharyngaus, mylo-pbaryngeus, ptery-pbaryngaus, and glosso-pbaryngaus. f\) Douglas makes two muscles of this the hyo-pharyngaus and syndesmo-pharyngaus. 3 3. Name. j .Constrictor pharyfi- gis inferior (*). about the glottis - - - I. Crico arytacnoide- us lateralis. 2. Crico-arytaenoide- us posticus. 3. Arytaenoideus ob- liquis. 4. Arytaenoideus transversus. 5. Thyreoarytaenoi- deus. {•) Th« cri( Origin. hyoides, and from the ligament that unites it with the thyroid cartilage. From the cricoid and thyroid cartilages. From the side of the cricoid cartilage. From the cricoid car- tilage posteriorly. From the basis of one of the arytaenoid cartilages From one of the ary- taenoid cartilages laterally. From the posterior and under part of the thyroid carti- lage. iryngxus and thyro-pharyn, Insertion. formis of the occi- pital bone, about its middle and be- fore the great fo- ramen. Into the middle of the pharynx. Into the basis of the arytaenoid carti- lage laterally. lnt the basis of he arytaenoidcartilage posteriorly Near the extremity of the other arytae- noid cartilage. Into the other arytae- n. id cartilage late- rally. Into the arytaenoid cartilage. jseui of Douglas. Use. upwards, and to compress the latter. To compress part of the pharynx. To open the glottis. To open the glottis. To draw the parts it is connected w ith towards each other. To shut the glotts. To draw the arytae- noid cartilage ror- wards. Muscles at the fore part of the neck, close to the verte- brae Name. (5. Arytacno-epiglot- tideus. 7. Thyreo-epiglotti- deus. Rectus capitis in- ternus major. 2. Rectus capitis in- ternus minor. 3. Rectus capitis la teralis-. Origin. From the upper part of the ;rytaenoid cart'lage laterally. Fn-m the thyroid car- tilage. Insertion. Into the side of the epiglottis. Into the side of the epiglottis. From the anterior ex- tremities of the transverse proces- ses of the five 1 w- ermost cervical ver- tebrae. From the anterior and upper p^rt of the first cervical vertebra From the anterior and Upper p rt of Into the fore part of the cuneiform pro- cess of the os occi- pitis. Near the basis of the condyloid process of tire os occipitis. Into the os occipi- tis, opposite to the Use. To move the epiglot- tis outwards. To pull the epiglot- tis obliquely down- wards (*.) To bend the head forwards. To assist the last de- scribed muscle. To move the head to one side. SP- SS Co (•) When either this ©r the preceding muicle acts with its fellow, the epiglottis is drawn directly downwards up»n the glottu Name. Origin. Insertion. Use. £ the transverse pro- stylo mastoid fora- cess of the first cer- men. vical vertebra. 4, Longus colli. Within the thorax, In'o the second cer- To pull the neck to laterally from the^- yicalvertebra ante- one side (*). bodies of the three riorly. uppermost dorsal ^O vertebrae; from the ^ basis and fore part *£, of the transverse ^ processesoffhefirst ^ and second dorsal ^ vertebrae, and of co the last cervical ^« vertebra; and last- '? ly, from the ante- rior extremities of the transverse pro- cesses of the 6thj 5th, 4th, and 3d cervical vertebrae. (») When both muscles act, the neck is drawn directly forwards. -------at the fore part of the abdo- men - Name. i. Obliquus externus. 2. Obliquus internus. Qri° m. From theloweredges of the eight infe- rior ribs, near their cartilages. From the spinous pro- cess of the three lowermost lumbar vertebrae, the back part of the os sa- Inscrtion. Use. Into the linea alba (* ),cssapubis(f), and spine cf the ilium \x). To compress and fup- port the viscera, assistin evacuating the faeces and urine, draw down the ribs, and bend the trunk forwards, or ob- liquely to one bide. Into the cartilages cf To assist the obliquus all the false ribs, externus. linea alba (§), and fore part of the pubis. (*) The linea alba is that tendinous expansion which reaches from the cartilago ensiformis to the os pubis. It is formed by the inter- lacement of the tendinous fibres of the oblique and transverse muscles, and on this account some anatomists have considered these as three digastric muscles. (t) A little above the pubis the tendinous fibres of this muscle separate from each other, so as to form an opening called the ring of the obliquus externus, and commonly, though improperly, the ring of the abdominal muscles, there being no such aperture either in the transversalis or obliquus internus. This ring in the male subject affords a passage to the spermatic vessels, and in the female to the round ligament of the uterus. (\) From the anterior and upper spinous process of the ilium, this muscle is stretched tendinous to the os pubis, and thus forms what is called l>y some Fallopius's, and by others Faupart's ligament. The blood-vessels pass under it to the thigh. ' ($) The tendon formed by the upper part of this muscle in its way to the linea alba is divided into two layers. The posterior layer runs ondcr, ;uid the anterior one over, the rectus muscle1. Name; 3. Transversalis. Origin. Insertion. Use. Into the linea alba To compress the ab- and cartilago ensi- dominaLviscera. formis. 4. Rectus abdominis. crum, the spine of the ilium, and back part of F-illopius's ligament (*). From the cartilages of the seven inferi- or ribs; the trans- verse processes of the last dorsal, and four upper lumbar vertebra?; the inner part of Fallopius's ligament and the spine of the ilium. From the upper edge Into the cartilages of To compress the fore of the pubis and the symphysis pu- bis. part of the abdo- men, and to bend the trunk forwards. the 5 th, fith. and 7 th ri s, and the ed./.e of the carti- lago ensiformis(f) (*) From this part it detaches some fibres which extend downwards upon the spermatic chord, and form what is described as the cremaster muscle. t) The fibres of the rectus are generally divided by three tendinou. intersections. The two upper thirds of this muscle passing be- tween the tendinous layers of the obliquus internus, are inclosed as it were in a sheath ; but at its lower part we find it immediately contiguous to the peritonasum, the inferior portion of the tendon of the transversalis passing orcr the rectus, and adhering to the anus- rior layer of the obliquus interna*. OB $ Name. 5. Pyramidalis (*). Must lis at the fore part of the thorax. 1. Pectoralis Major. 2. Subclaviust. Origin. From the anterior and upper part of the pubis. From the cartilagi- nous ends of the 5th and 6th ribs ; the sternum, and anterior part of the clavicle. From the cartilage of the first rib. Insertion. Into the linea alba and inner edge of the rectus, com- monly about two inches above die pubis. Into the upper and in- ner part of the os humeri (f). Into the under face of the vicle. sur- cla- Use. To afflft the lower portion of the rec- tus. To draw the arm for- wards, or oblique- ly forwards. To move the clavi- cle forwards arid downwards, and to as isr in raising the first rib. 3 S- Co Co (•) This muscle is sometimes wanting. , (t) The fibres of this muscle pass towards the axilla in a folding manner, and with those of the latissunus dorsi form the arm-pit. Muscles that con- cur in forming the thorax, Name. Pectoralis minor (*). 4. Serratus Magnus. 1. Diaphragma (f). 2. LeVatores consta* rum. . Intercostales exter- ni, . Intercostales inter- terni (J). Origin. From the upper edges of the 3 d, 4th, and 5 th ribs. From the eight supe- rior ribs. From the transverse processes of the last cervical and the eleven upper dor- sal vertebrae. From the lower edge of each upper rib. Insertion. Intothe coracoid pro- cess of the sca- pula. Into the basis of the scapula. Use. To move the scapu- la forwards and downwards, or to elevate the ribs. To bring the scarula forwards. O o Into the upper side of To move the ribs up- each rib, near its wards and out- tuberosity. wards. Into the superioredge To elevate the ribs. of each lower rib. (*) This and some other muscles derive their name of serratus, from their arising by a number of tendinous or fleshy digitations, re- sembling the teeth of a saw (serra.J (t) For a description of the diaphragm, see Part IV. Sect. IV. (i) The origin, insertion, and us* of the internal intercostales, are similar to those of the external. The reader, however, will b« pleased to observe, that the ibtercostales externi occupy the spaces between the ribs only from the spine to their cartilages ; from thence to the sternum, there being only a thin membrane, which is spread over the intercostales interni j and the latter, on the contrary, ess- tend only from the sternum to the angles of each rifc. 3 Name. part and trunk, —at the back of the neck 5.Sterno-costales(*). Trapezius (f), or cucullaris. n Origin. From the cartilagoen- tiformis, and low- er and middle part of the sternum. From the middle of the os occipitis, and the spinous pro cesses cf the two inferior cervical, and of all the dor- sal, vertebrae (%). Insertion. Into the cartilages of the 2d, 3d, 4th, 5th, and 6th ribs. Into the posterior half of the clavicle, part of the acro- mion, and the spine of the scapula. Use. To depress the cari^ lages of the ribs. To move the scapula. The fibres of the external mufrtes run obliquely forwards; thofe of the internal obliquely backwards. This difference in the direc- tion of thdr ^ Galen to fuppofe that they were intended for different ufes ; that the external intercoftals, for inftance Tveo elevate and tL internal ones todepress the ribs. Fallopius seems to have been the first who ^^^^%^ of7his doctrine, which has since been revived by Boyle, and more lately still by Hamberger, whose theoretical arguments on this sub- ^^""ot^^ each side. Vesalius, and after him Douglasand Albiiuis,<:on- sider them as forming a single muscle, which, on account of its shape, they name triangularu. Verheyen, Winslow, and Haller, more ^Lr;^. bTthem as so many separate muscles, which, on account of their origin and insertion, they name firno-coftale,. P (Pt Inamedi oyRioLnurfron, ?,«*, on account of its quadrilateral fliape. Columbus and others gave it the name of Allans, ^^^^Si^^W, united with thofe of its fellow in the aapc of the neck, form what is called the ligament Name. z. Rhomboideus (*). 3. Latissimus dorsi. 4. Serratus posticus. inferior Origin. From the fpinous pro- ceffes of the three lowermoftcervical, and of all the dor- fal vertebras. From part of the spine of the os ili- um, the spinous processes of the os sacrum and lumbar vertebrae, and of six or eight of the dor- sal vertebrae; also from the four infe* rior false ribs neap their cartilages. From the spinous pro- cesses of the two lowermost dorsal, and of three of the lumbar vertebras. Insertion. Into the basis of the scapula.. Into the os humeri, at the inner edge of the groove for lodging the long head of the biceps muscle. Into the lower edges of the three or four lowermostribsnear their cartilages. Use. To move the scapula upwards and back- wards. To draw the os hu- meri downwards and backwar ds,and to roll it upon its axis. to O to To draw the ribs out- wards,downwards, and backwards. e* Co (*) This muscle consists of two distinct portions, which are described as separate muscksby Albinus, under the names of rbomboMcus minor and rhomboideus my or. Name, 5. Levator scapulae. 6. Serratus superior posticus. 7. Splenius (*); Origin. From the transverse processes of the four uppermost vertebrae colli. From the lower part of the ligamentum colli, the spinous process of the low- ermost cervical vertebra, and of the two superior dorfal vertebras. From the spinous processes of the four or five upper- most vertebrae of the back, and of the lowermost cer- vical vertebral Insertion. Use. Into the upper angle To move the scapula of the scapula. forwards and up- wards. Into the 2d, 3d, and To expand the tho- 4th ribs. rax. Into the transverse processes of the two first cervical vertebrx, the up- per and back part of the mastoid pro- cess, and a ridge on the os occipitis. To move the backwards. head a- Co Co (*) According to some writers, this muscle has gotten its name from itsmemblanee to the spleen; others derive it hcmjpimum •jplint. o C/J Name. Origin. Insertion, Use. 8. Complexus (*). 9. Trachelo-mastoi- deus (f). 10. Rectus capitis po- fticus major. 11 • Rectus capitis po- sticus minor. 12. Obliquus superior capitis. From the transverse processes of thefour or five uppermost dorsal, and of the six lowermost cer- vical vertebrae. From the transverse processes of the first dorsal vertehra, and four or five of the lowermost, cer- vical vertebrae. From thespinouspro- cess of the second cervical vertebra. From the first verte- bra of the neck. From the transverse process Of the first cervical vertebra. Into the os occipitis Into the mastoid pro- cess. Into the os occipitis. Into the os occipitis. Into the os occipitis. To draw the head backwards. To draw the head ba<-kwa;ds. To extend the head and draw it back- wards. To assist the rectus major. To draw the head. backwards. (•> So named on account of its complicated stricture. (f) So named from its origin from the neck (Tp»yer of fleshy fibres is found crowng, and intimately adhering to it. This portion, which is descrih:'■' by Albinus, under the name of transversalis .ervtcis, m. ) \ ) pr« : ' .y be considered as an appendage to the long.'ssi mus dorsi. It arises from the transverse processes of the five or six superior dorsal verteur^, and is inserted into the transverse processes of O Name, 16. Spinalis dorsi. 17. Semi-fpinalis dor- si. 18. Multifidus Spi- nae (*). Origin. From the spinous processes of the up- permost lumbar and lowermost dorsal vertebrae. From the transverse processes of the 7th, 8th, 9th, and ioth vertebrae of the back. From the os sacrum, ilium, oblique and transverse proces- ses of the lumbar vertebrae, trans- verse processes of Insertion. Use. to o Into the spinous pro- To extend the verte- cesses of the nine bra;. superiordorsal ver- tebrae. Into the spinous pro- To extend the fpine cesses of the four obliquely back- uppermost dorsal, wards. «§ and lowermost of the cervical verte- <*». &- brae. rt> Into the spinous pro- To extend the back b* cesses of the lum- and draw it back- gt bar, dorsal, and six wards, or to one 5 of the cervical ver- side. tebrae. Co the six inferior cervical vertebrae. By means of this appendage the longissimus dorsi may serve to move the neck to one side, or oblique- ly backwards. *) Anatomists in general have unnecessarily multiplied the muscles of the spine. Albinus has the merit of having introduced greater simplicity into this part of myology. Under the name of multifidus spina, he has very properly included those portions of muscular flesh intermixed with tendinous fibres, situated close to the back part of th* spine, and which are described by Douglas under the names of transversales colli, dorsi, \Sf lumborum. Name. Origin. Insertion. Use. lo. Semi-spinalis col- li. 20. Scalenus (*). 21. Inter-spinalis (f ). the dorsal, and four of the cervical vertebrae. From the transverse processes of the five or six uppermost dorsal vertebrae. From the transverse processes of the five inferior cervi- cal vertebrae. From the upper part of each of the fpi- nous processes of the six inferior cer- vical vertebrae. Into the spinous pro- cesses of the 2d, 3d, 4th, 5th, and 6th cervical ver- tebras. Into the upper and outer part of the first and second ribs. Into the under part of each of the spinous processes of the vertebrae above. To stretch the neck obliquely back- wards. To move the neck forwards, or to one side. To draw the spinous processes towards each other. 3 «*•. »- (*) The ancients gave it this name from its resemblance to an irregular triangle ( These muscles are to be found only in the neck and loins ; which have been described, as the inter-trans-vir sales dorsi being rather ^VThtTnd^herilowing pair of muscle, derive their name of psoa, from*-, lumbus, on account of their situation at the anterior part- of the loins. Name. &rigin. u 3. Iliacus internus. From the inner lip, hollow part, and edge of the os ilium. 4. Quadratus lumbo- From the posterior rum (*). part of the spine of the ilium. 5. Coccygseus. From the posterior and inner edge of the spine of the ischium. Insertion. In common with the psoas magnus. Into the transverse processes of the four uppermost lumbar vertebrae, the inferior edge of the last rib, and the side of the low- ermost dorsal ver- tebra. Into the lower part of the os sacrum, and almost the whole length of the os coccygis late- rally. Use, To assist the psoas magnus. To support the spine, or to draw it to one side. To draw the os coc- cygis forwards and inwards (f). (*) So called from its shape, which is that of an irregular square. .... ,1 _._» Ar .t,. (V Some of the fibres of this muscle ace united with those of the levator ani, fo that it assists in closing the lower part ot we pelvis. to o n ..on the scapu- la and tipper part of the osJiumeri, Name. i. Deltoidcs (*). 2, Supra-spinatus.. 3. lnfra-spinatus. 4. Teres minor (f). 5, Teres major. 6. Subscapularis. Origin. From the clavicle, processus acromi- on, and spine of the scapula. From the basis, spine, and upper costa of the scapula. From the base and spine of the scapu- la. From the inferior co- sta of the scapula. From the inferior angle, and inferior costa of the scapu- la. From the basis, supe* rior and inferior co- sta of the scapula* Insertion. Into the anterior and middle part of the os humeri. Into a large tubero- sity at the head of the os humeri. Into the upper and middle part of the tuberosity. Into the lower part of the tuberosity. Into the ridge at the inner side of the groove formed for the long head of the biceps. Into the upper part of a small tuberosity at the head of the os humeri. Use. To raise the arm- To raise the arm. To roll the os humeri outwards. To assist the infra spinatus. To assist in the rota- tory motion of the arm. To roll the arm in- wards. -(*) Sto named from its supposed resemblance to the Greek a reversed. l.\ ) This and the following pair are called tores, from their being; of a long and round shape. Muscles on the os humeri, - - - Name, Coraco-brachia- lis (*). r. Biceps flexor cubi- ti. Insertion. Into the middle and inner side of the os humeri. Into the tuberosity at the upper end of the radius. Origin. From the coracoid process of the sca- pula. By two heads, one from the coracoid process, and the other, or long head, from the upper and outer edge of the glenoid cavity of the scapula. From the os humerij below, and at each side of the tendon of the deltoides. By three heads; the first, from the infe- rior costa of the scapula; the se- cond, from the up*- per and outer part (*) This mustle affords a passage to the mttsculo-cutaneiBui nerve. Use. To roll the arm for- wards and upwards. Tobend the fore-arm. * Brachialis internus. Triceps extensor cubiti. Into a small tubero- sity at the fore part of the coronoid pro- cess of the ulna. Into the upper and outer part of the olecranon. To assist in bending U\e fore-arm. To extend the fore- arm. fc> Name. on the fore- arm, - - i. Supinator longus. 2. Extensor carpi ra- dialis longus. 3. Extensor carpi ra- dialis brevis. 4. Extensor digito rum communis. Origin. of the os humeri; and the third, from the back part of that bone. From the outer ridge and anterior sur- face of the os hu- meri, a little above its outer condyle. Immediately below the origin of the supinator longus. From the outer and lower part of the outer condyle of the os humeri, and the upper part of the radius. From the outer con- dyle of the os hu- meri. Insertion. Use, to 13 Into the radius near its styloid process. Into the upper part of the metacarpal bone of the fore- finger. Into the upper part of the metacarpal bone of the middle finger. To assist in turning the palm <^f the hand upwards. To extend the wrist. To assist the extensor longus. Into the back part of To extend the fingers all the '^ones of the fore finger. •3 Name. Origin. Insertion. Use. nans. 7. Anconaeus (*). 5. Extensor minimi From the outer con. digiti. dyle of the os hu- meri. 6. Extensor carpi ul- From the outer con- dyle of the os hu- meri. From the outer con- dyle of the os hu- meri. 8. Flexor carpi ulna- From the inner con- ris. dyle of the os hu- meri, and a .terior edge of the olecra- non (f). From the inner con- dyle of the os hu- meri. 9. Palmaris longus. Into the b^nes of the To ext nd the little little finger. finger. Into the metacarpal To assist in extending bone of the little the wrist. finger. Into the outer edge To extend the fore of the ulna. arm. Into the os pisiforme. To assist in bending the hand. Into the internall an- To bend the hand. nular ligament, and aponeurosis p ilmaris \%). 3 a* (*) So called from »y«»v, cubitus. (+) Betwe n t te two origins of this muscle we find the ulrtor-nerve going to the fore arm. , i The a oneurosis palm .ris is a tendinous membrane that , xtoids over the palm of the hand. Some anatomists have supposed it to be a production of the tendo,, of this iruscle, but without sufii --M grounds; for in seme subjects we find the palmaris longus inserted whoiiy -.to the annular lij/animt, sr is ^o be perfectly distinct from this aponeurosis; and it now and then happens, that no palmaris longus u to be found, whertas this expansion is never deficient. 03 Nam*. yOt Flexor £arpi ra, dialis. 11. Pronator radii teres. 12. Flexor sublimis perforatus (*). 13. Supinator radii brevis. Origin. From the inner con- dyle of the os hu- meri. From the outer con- dyle of the os hu- meri, and coronoid process of the ulna. From the inner con- dyle of the os hu- meri, inner edge of the coronoid pro- cess of the ulna, and upper and an- terior part of the radius. From the outer con- dyle of the os hu- meri, and posterior surface and outer edge of the ulna. Insertion. Into the metacarpal bone of the fore finger. Into the anterior and convex edge of the radius near its middle. Into the second bone pf each finger. Into the anterior, in- ner, and upper part of the radius. Use. & To bend the hand. To roll the hand in- wards. To bend the second Qr joint of the finger. To roll thp radius outwards. I (•) This muscle is named perforator, en account of the four tendons in which it terminates, being perforated bf those of another muscle, the perferans. Name. 14. Abductor cis longus. 15. Extenfor pollicis. 16. Extensor pollicis. 17. Indicator. 0i tgt polli- From the middle and back part of the ulna, interofleous ligament, and ra- dius. minor From the back part of the ulna, and in- terofleous li gament and radius. major From the back of the ulna and interofTe- ous ligament. From the middle of the ulna. 18. Flexor profundus perforans. 19. Fleicor pollicis* longus From the upper and fore part of the ulna, and interos- seous ligament. From the upper and fore part of the radius* Insertion. By two tendons into the os trapezium, and first bone of the thumb. Into the convex part of the second bone of the thumb. Into the third and last bone of the thumb. Into the metacarpal bone of the fore- finger. Into the fore part of the last bone of each of the fingers. Use. To stretch the first bone of the thumb outwards. To extend the second bone of the thumb obliquely out- wards. To stretch the thumb obliquely back- wards. To extend the fore- finger. To bend the last joint of the fingers. Into the last joint of To bend the last joint the thumb. of the.thumb.. a- c-r 60 to 09 Name, 20. Pronator quadratus. Origin, radii From the inner and lower part of the ulna. Insertion. Into the radius, op- posite to its origin. MuscLEsonthehand, 1. Lumbricales (*). From the tendons of Into the tendons of the perforans. 2. Abductor bravis From the fore part pollicis. of the intern d an- nular ligament, os scaphoides, and one of the tendons of the abductor lon- gus pollicis. 3. Opponens pollicis. From the inner and antei ior part of the internal annular li- gament, and from the os scaphoides. the extensor digi- torum communis. Into the outer side of the 2d bone of the thumb, near its root. Use. To roll the radius in- wards, nd of course to assist in the pronation of the hand. To bend the first, and to exiend the two last joints of the fingers (f). To move the thumb from the lingers. Into the first bone of To move the thumb the thumb. inwards, and to turn it upon its axis. (") So named from their being shaped somewhat like the lumbricus or earth-worm. (■|- Fallopius whs the first who remarked the two opposite uses of this muscle. Their extending power is owing to their connection with the extensor communis. to «S Name. Origin. Insertion. Use. rt 4. Flexor brevis pol* licis. 5. Abductor pollicis. 6. Abductor indices. 7. Palmaris brevis. 8. Abductor minimi digiti. 9. Flexor parvus mi- nimi digiti. From the os trapezdi- des, internal annu- lar ligament, os magnum, and os unciforme From the metacarpal bone of the middle finger. From the inner side of the first bone of the thumb, and from the os trape- zium. From the internal an- nular ligament, and aponeurosis pal- maris. From the internal an- nular ligament and os pisiforme. From the os uncifor- me and internal an- nular ligament. Into the ossa sesa- moidea and second bone of the thumb. Into the basis of the second bone of the thumb. Into the first bone of the fore finger pos- teriorly. Into the os pisiforme, and the skin cover- ing the abductor minimi digiti. Into the side of the first bone of the little finger. Into the first bone of the little finger. To bend the second joint of the thumb. To move the thumb towards the fin- gers. To move the fore fin- ger towards the thumb. To contract the palm of the hand. To draw the little finger from the rest. To bend the little fin- ger. $ fcO Name. Origin. Insertion. Use. Muscles at the back part of the pelvis, and upper part of the thigh, - 10. Abductor meta- carpi minimi digiti. n. Interossei interni. i2.Interossei externi. i. Glutaeus (f) max- imus. From the os uncifor- me and internal an- nular ligament. Situated, between the metacarpal bones. Situated between the metacarpal bones on the back of the hand. From the spine of the ilium, posterior sa- cro-ischiatic liga- Into the metacarpal bone of the little finger. Into the roots of the fingers. Into the roots of the fingers. To move that bone towards the rest. To extend the fingers and move them to- wards the thumb To extend the fin- gers ; but the first draws the middle finger inwards, the second draws it out- wards, and the third draws thering finger inwards. Into the upper part To extend the thigh of the linea as per a and draw it out- of the os femoxis. wards. (*) The third interosseus internus (for there are four of the externi and three cf the interni) differs from the rest in drawing the middle finger from the thumb. (f) From >-».»">, nates. Name. Origin. Insertion. Use. 2. Glutaeus medius. 3. Glutaeus minimus. 4. Pyriformis (*). 5. Gemini (f). ments, os sacrum, and os coccygis. From the spine and superior surface of the ilium. From the outer sur- face of the ilium and the border of its great niche. From the anterior part of the os sa- crum. By two portions, one from the outer sur- face of the spine of the ischium; the other from the Into the outer and back part of the great trochanter of the os femoris. Into the upper and anterior part of the great trochanter. Into a cavity at the root of the tro- chanter major. Into the same cavity as the pyriformis. To draw the thigh outwards and a lit- tle backwards, and when it is bended, to roll it. To assist the former. To roll the thigh out- wards. To roll the thigh out- wards, and likewise to confine the ten- don of the obtura- tor internus, when (*) So named from its pear-like shape. (t) The two portions of this muscle having been described as two distinct muscles by some anatomists, have occasioned it to be named gimini. The tendon of the obturator internus runs between these two portions. r-* Name. on the thigh (t). 6. Obturator internus. 7. Quadratus (*) fe- moris. 1. Biceps flexor cm- Origin, tuberosity of the ischium and poste- rior sacro-ischiatic ligament. From the superior half of the inner border of the fo- ramen thyroideum. From the tuberosity of the ischium. By two heads; one from the tuberosity of the ischium, Insertion. Use. the latter is in ac-tion. to O Into the same cavity with the former. To roll the thigh out-wards. Into a ridge between the trochanter ma-jor and trochanter To move the thigh outwards. Co <■> e\* 06 minor. Into the upper, and To bend the leg. back part of the fi- bula (t). f*\ This muscle is not of the square shape its name would seem to indicate. ..,.,. 1 i. • u r+ The muscles of the lee and thigh are covered by a broad tendinous membrane called fasaa lata that surrounds them in the man- ner of a sheath. It is sent off from the tendons of the glut*, and other muscles, and dipping down between the muscles it covers ad- here, to the linea aspera, and spreading over the joint of the knee, gradually ^appears on the leg. It M thickest on the inside of the thigh. . {\) The tendon »f this muscle forms the outer hamstring. Name. Origin. Insertion, Use. 2. Semitendinosus. 3. Semi-membrano- sus (*). 4. Tensor vaginae fe- moris. 5. Sartorius. 6. Rectus. the other from the linea aspera near the insertion of the glutaeus maximus. From the tuberosity of the ischium. From the tuberosity of the ischium. From the superior and anterior spi- nous process of the ilium From the superior and anterior spi- nous process of the ilium. By two tendons ; one from the anterior and inferior spi- Into the upper and inner part of the tibia. Into the upper and back part of the •head of the tibia. Into the inner side of the fascia lata, which covers the outside of thethigh. Into the upper and in- To bend the leg in- ner part of the ti- wards (f). bia. To bend and draw the leg inwards. To bend the leg. To stretch the fascia. Into the upper and fore-part of the pa- tella. To extend the leg. s. CO ?5 (•) So named on account of its origin, which is by a broad flat tendon three inches long. (t) Spigelius was the firft who gave this the name of sartorius, or the taylor'e muscle, from, its use in crossing the legs, N3 b3 Name. Origin. Insertion. Use. 7. Gracilis. 8. Vastus externus (*)• 9. Vastus internus. nous process of the ilium ; the other from the posterior edge of the coty- loid cavity. From the fore-part of the ischium and pubis. From the anterior and lower part of the great trochanter, and the outer edge of the linea aspera. From the inner edge of the linea aspera, beginning between the fore-part of the os femoris and the root of the lesser trochanter. Into the upper and in- ner part of the ti- bia. To the upper and out- er part of the pa- tella. Into the upper and inner part of the patella. To bend the leg. To extend \h.z leg. To extend the leg. (*) The vastus externus, vastus internus, and crurseus, are so intimately connected with each other, that some anatomists have been in iuced to consider them as a triceps, or single muscle with three heads. Name. 10. Cruraeus (*). 11. Pectinalis. 12. Abductor longus femoris (f). 13. Abductor brevis femoris. 14- Abductor mag- nus femoris. Origin. From the outer .and anterior part of the lesser trochanter. From the anterior edge of the os pu- bis, or pectinis, as it is sometimes called. From the upper and fore part of the os pubis. From the fore part of the ramus of the os pubis. From the lower and fore part of the ra- mus of the os pu- bis. Insertion. Use. Into the upper part To extend the leg. of the patella. Into the upper and To draw the thigh fore part of the li- inwards, upwards, nea aspera. and to roll it a lit- tle outwards. Near the middle and" back part of the li- nea aspera. Into the inner and upper part of the linea aspera. Into the whole length of the linea aspera. To draw the thigh inwards, upwards, I" and to roll it a lit- tle outwards. (*) Under the crursus we sometimes meet with two small muscles, to which Albinus has given the name of sub-erartti. They ter- minate on each side of the patella, and prevent the capsular ligament from being pinched. When they are wanting, which is very often the case, some of the fibres of the crnrsus are found adhering to the capsula, (f) This and the two following muscles have been usually, but improperly, considered as forming a single muscle with three heads, and on that account named triceps femoris. Musclbs on the leg, Nam*, ij. Obturator exter- nus, i.Gastrocnemius (*) externus. 2. Gastrocnemius (f) internus. Origin. From part of the ob- turator ligament, and the inner half of the circumfe- rence of the fora- men thyroideum. By two heads ; one from the inner con- dyle, the other from the outer con- dyle of the os fe- moris. By two heads; one from the back part of the head of the fibula, the other from the upper and back part of the tibia. Insertion. Into the os femoris near the root of the great trochan- ter. By a great round •■en- don, common to this and the follow- ing muscle. By a large tendon (the tendo achillis) common to this and the former muscle, into the lower and back part of the os calcis. Use, To move the thigh outwards in an ob- lique direction,and likewise to bend and draw it in- wards. To extend the foot. To extend the foot. to 5s- Co (*) r*rpo*»>tjui'*, sura, " the calf of the leg." (|) This muscle is by some anatomists named soltus, on account of its being shaped like the sole-fish. m ** Name. 3. Plantaris (*)■ 4. Popliteus (\), 5. Flexor longus digi- torum pedis {%). Origin. From the upper and posterior part of the outer condyle of the os femoris. From the outer con- dyle of the thigh* From the upper and inner part of the tibia. Insertion. Into the inside of the back part of the os calcis; the upper and part of the Into inner tibia. By four tendons, which, after pas- sing through the perforations in those of the flexor digitorum brevis, are inserted into the last bone of all the toes except the great toet Use. To assist in extend- ing the foot. To assist in bending the leg and rolling it inwards. To bend the last joint of the toe. R Co <-> (») this muscle has gotten the dame of plantaris, from its feeing supposed to furnish the aponeurosis that coVers the sole of the foot; "but it does not in the least contribute to the formation of that tendinous expansion. (+) So called on account of its situation at the ham (poples). t . (|) This muscle, about the middle of the foot, Unites with a fleshy mass, which, from its having first been described by Sylvius, is fsually called massa ta'rnea JacoBX Sylvii. to to Name. 6. Flexor longus pol- licis pedis. 7. Tibialis posticus. 8. Peroneus longus. 9. Peroneus brevis. Origin. Insertion. Use. From the back part, and a little below the head of the fi- bula. From the back part and outer edge of the tihia, and like- wise from the in- terosseous liga- ment and adjacent part of the fibula. From the outer side of the head of the tibia, and also from the upper, anterior, and outer part of the perone or fibu- la, to which it ad- heres for a consi- derable way down. From the outer and fore part of the fi- bula. Into the last bone of To bend tire great the great toe. toe. Into the inner and upper part of the os naviculare and side of the os cu- neiforme medium. To move the foot in- wards. Into the metatarsal To move the foot out- bone of the great wards. toe. Into the metatarsal bone of the little toe. To assist the last de- scribed muScle.. Name. 10. Extensor longus- digitorum pedis. n. Peroneus tertius. 12. Tibialis antrcas. 13. Extensorproprius pollicis pedis. Muscles onthefootr 1. Extensor brevis di- gitorum pedis. Origin. Insertion. Use. From the upper, out- er, and fore part of the tibia, inter- osseous ligament, and inner edge of the fibula. From the fore-part of the lower half of the fibula, and from the interos- seous ligament. From the upper and fore part of the ti- bia. From the upper and fore part of the ti- bia. From the upper and anterior part of the os calcis. By four tendons into the first joint of the smaller toes. Into the metatarsal bone of the little toe. Into the os cunei- forme internum. Into the convex sur- face of the bones of the great toe. By four tendons ; one of which joins the tendon of the ex- ternus longus polli- cis, and the other three the tendons of the extensor di- gitorum longus. To extend the toes. To ihe bend the foot. To bend the foot. To extend the great toe. To extend the toes. Name. Origin, Insertion. Use. to to 00 t. Flexor brevis d»gi- torum pedis. 3. Abductor pollicis pedis. 4. Abductor minimi digiti, 5, Lumbricales pedis. From the lower part of the os calcis. From the inner and lower part of the os calcis. From the outer tuber- cle of the os calcis, the root of the me- tatarsal bone of the little toe, and also from the aponeuro- sis plantaris. From the tendons of the flexor longus digitorum pedis. By four tendons, which, after af- fording a passage to those of the flex- or longus, are in- serted into the se- cond phalanx of each of the small toes. Into the first joint of the great toe. Into the outer side of the first joint of the little toe. Into the tendinous ex- pansion at the up- per part of the toes. To bend the second joint of the toes. To move the great toe from the other toes. To draw the little toe outwards. To draw the toes iar wards. 09 Name. Origin. Insertion. 6. Flexor brevis pol- licis pedis. 7. Abductor pollicis pedis. 3. Transversalis pe- dis. 9. Flexor brevis mi- nimi digiti pedis. 10. Interossei pedis interni (*). .--------■------exter- ni (f). From the inferior and anterior part of the os calcis, and also from the inferior part of the os ca- neiformeexternum. From near the roots of the metatarsal bones of the 2d, 3d, and 4th toes. From the outer and under part of the anterior end of the metatarsal bone of the little toe. From the basis of the metatarsal bone of the little toe. Situated between the metatarsal bones. By two tendons into the first joint of the great toe. Into the outer os se- samoideum, orfirst joint of the great toe. Into the inner os se- samoideum, and anterior end of the metatarsal bone of the great toe. Into the first joint of the little toe. To bend the first joint of the great toe. To draw the great toe nearer to the rest, and a1 so tobend it. To contract the foot. To bend the little toe. (*) The intercssei'interni are three in number; their use is to draw the smaller ft) The interossei externi are four in number j the first serves to move the fore wards. All the interossei assist in extending the toes. toes towards the great toe. toe towards the great toe; the rest move the toes out- 230 Of the Muscles. EXPLANATION of PLATES XXIII. and XXIV. Plate XXIII. Fig. l. The Muscles immediately under the common teguments on the anterior part of the body are represented on the right side; and on the left side the Muscles are seen which come in view when the exterior ones are taken away. A, The frontal muscle. B, The tendinous aponeurosis which joins it to the occipital; hence both named occipito-frontalis. C, Attol- lens aurem. D, The ear. E, Anterior au- ris. F F, Orbicularis palpebrarum. G, Le- vator labii superioris alaeque nasi. H, Leva- tor anguli oris. I, Zygomaticus minor. K, Zygomaticus major. L, Masseter. M, Or- bicularis oris. N, Depressor labii inferioris. O, Depressor anguli oris. P, Buccinator. QQ,, Platysma myoides. RR, Sterno-cleido- mastoidaeus. S, Part of the trapezius. T, Part of the scaleni. Superior Extremity—U, Deltoides. V, Pectoralis'major- W, Part of the latissimus dorsi. X X, Biceps flexor cubiti. Y Y, Part of the brachialis externus. Z Z, The begin- ning of the tendinous aponeurosis (from the biceps), which is spread over the muscles of r* if- Of the Muscles. 231 the fore-arm. a a, Its strong tendon inserted into the tubercle of the radius, b b, Part of the brachialis internus- c, Pronator radii te- res, d, Flexor carpi radialis- e, Part of the flexor carpi ulnaris- f, Palmaris longus. g, Aponeurosis palmaris. 3, Palmaris brevis. 1, Ligamentum carpi annulare. 2 2, Abductor minimi digiti- h, Supinator radii longus. i, The tendons of the thumb- k, Abductor pol- licis. 1, Flexor pollicis longus. m m, The tendons of the flexor sublimis perforatus, profundus perforans, and lumbricales—The sheaths are entire in the right hand,—in the left cut open to show the tendons of the flex- or profundus perforating the sublimis. Muscles not referred to—in the left supe- rior extremity—n, Pectoralis minor, seu ser- ratus anticus minor, o, The two heads of (x x) the biceps, p, Coraco-brachialis. q q, The long head'of the triceps extensor cubiti. rr, Teres major. ((, Subscapulars. 11, Ex- tensores radiales. u, Supinator brevis- v, The cut extremity of the pronator teres, w, Flexor sublimis perforatus. x, Part of the flexor profundus, y, Flexor pollicis longus. z, Part of the flexor pollicis brevis- 4, Ab- ductor minimi digiti. 5, The four lumbri- cales- Trunk-—6, Serrated extremities of the ser- ratus anticus major* 7 1, Obliquus externus abdominis. 8 8, The linea alba- 9, The um- bilicus. 10, Pyramidalis. 11 11, The sper- matic cord. On the left side it is covered by 232 Of the Muscles. the cremaster. 12 12, Rectus abdominis. 13, Obliquus internus- 14 14, he. Intercostal muscles. Inferior Extremities—a a, The graci. lis. b b, Parts of the triceps, c c, Pectialis. dd, Psoas magnus- e e, Iliacus internus. yj Part of the glutaeus medius- g, Part of the glutaeus minimus, h, Cut extremity of the rectus cruris, ii, Vastus externus, k, Ten- don of the rectus cruris. / /, Vastus internus. * Sartorius muscle. ** Fleshy origin of the tensor vaginae faemoris or membranosus. Its tendinous aponeurosis covers (i) the vastus externus in the right side, m m, Patella, n n, Ligament or tendon from it to the tibia, o, Rectus cruris, p, Cruraeus. q q, The tibia. r r, Part of the Gemellus or gastrocnemius ex- ternus.* fff, Part of the soleus or gastroc- nemius internus. t, Tibialis anticus. u, Ti- bialis posticus, v v, Peronaei muscles, ww, Extensor longus digitorum pedis, x x, Ex- tensor longus pollicis pedis- y, Abductor pol- licis pedis. Fig- 2- The Muscles, Glands, he. of the Left Side of the face and neck, after the common Teguments and Platysma myoides have been taken off. a, The frontal muscle, b, Temporalis and temporal artery, c, Orbicularis palpebrarum. d, Levator labii superioris alaeque nasi, e, Levator anguli oris, f, Zygomaticus. g, De- pressor labii inferioris. h, Depressor anguli Of the Muscles. 233 oris, i, Birccinator. k, Masseter. 11, Paro- tid gland- m, Its duct- n, Sterno-cleido-mas- toidseus. o, Part of the trapezius, p, Sterno- hyoidaeus. q, Sterno-thyroidaeus. r, Omo- hyoidaeus. f, Levator scapulas- 11, Scaleni. u, Part of the splenius. Fig. 3. The Muscles of the Face and Neck in view after the exterior ones are taken away. a a, Corrugator supercilii. b, Temporalis- c, Tendon of the levator palpebral superioris- d, Tendon of the orbicularis palpebrarum, e, Masseter- f, Buccinator, g, Levator anguli oris, h, Depressor labii superioris alaeque na- si, i, Orbicularis oris- k, Depressor anguli oris. 1, Muscles of the os hyoides. m, Ster- no-cleido-mastoidseus. Fig. 4- Some of the Muscles of the Os Hy* oides and Submaxillary Gland. a, Part of the masseter muscle- b, Poste- rior head of the digrastic. c, Its anterior head, dd, Sterno-hyoidaeus- e, Omo-hyoi- daeus. f, Stylo-hyoidaeus. g, Submaxillary gland in situ. Fig. 5. The Submaxillary Gland and Duct. a, Musculus mylo-nyoidaeus. b, Hyo-glos- sus. c, Submaxillary gland extra situ, d, Its duct. Plate XXIV. Fig. 1. The Muscles immediately under the common teguments on the posterior part G g 234 Of the Muscles. of the body, are represented in the right side; and on the left side the Muscles are seen which come in view when the exterior ones are taken away. Head.—A A, Occipito-frontalis. BT Attol- lens a-urem. C, Part of the orbicularis palpe- brarum. D, Masseter. E, Pterygoidaeus in- ternus. Trunk.—Right side. FFF, Trapezius sen cucullaris. G G G G, Latissimus dorsi. H, Part of the obliquus externus abdominis. Trunk,—Left side.. I, Splenius. Kr Part pf the complexus. L, Levator scapulae. M, Rhomboides. N N, Serratus posticus inferi- or. O, Part of the longissimus dorsi. P, Part of the sacro-lumbalis. Q, Part of the semi-spinalis dorsi. R, Part of the serratus anticus major. Sy Part of the obliquus in- ternus abdominis. Superior Extremity.—Right side. T, Deltoides. U, Triceps extensor cubiti. V, Supinator longus. W W, Extensores carpi radialis lon-gior and brevior. X X, Extensor carpi ulnaris. Y Y, Extensor digitorum com- munis. Z, Abductor indicia 12 3, Exten- sores pollicis. Superior Extremity.—Left side, a, Su- pra spinatus. b, Infra-spinatus. c, Teres minor, d, Teres major, e, Triceps extensor cubiti. f f, Extensores carpi radiales. g, Su- pinator brevis. h, Indicator. 12 3, Exten- sores pollicis. i, Abductor minimi digiti. k, Interossei. Inferior Extremity.—Right side. 1, Glu- taeus maximus. m, Part of the Glutaeus me- Of the Muscles. 235 dius. n, Tensor vaginae femoris. o, Graci- lis, p p, Abductor femoris magnus. q, Part of the vastus internus. r, Semimembrano- sus, s, Semitendinosus- t, Long head of the biceps flexor cruris, u u, Gastrocnemius ex- ternus seu gemellus, v, Tendo Achillis. w, Soleus seu gastrocnemius internus. x x, Pe- ronaeus longus and brevis. y, Tendons of the flexor longus digitorum pedis;—and under them * flexor brevis digitorum pedis- z, Ab ductor minimi digiti pedis. Inferior Extremity.—Left side, m, n, o, p, q, r, s, t, v, w w, x x, y, z, Point the same parts as in the right side, a, Pyrifor- mis. b b, Gemini, c c, Obturator internus. d, Quadratus femoris. e, Coccygaeus. f The short head of the biceps flexor cruris- gg, Plantaris. h, Poplitaeus. i, Flexor longus pollicis pedis. Fig. 2. The Palm of the Eeft Hand after the .common Teguments are removed, to show the Muscles of the Fingers. a, Tendon of the flexor carpi radialis. b, Tendon of the flexor carpi ulnaris. c, Ten- dons of the flexor sublimis perforatus, pro- fundus perforans and lumbricales. d, Ab- ductor pollicis. e e, Flexor pollicis longus- f, Flexor pollicis brevis. g, Palmaris brevis- h, Abductor minimi digiti. i, Ligamentum carpiannulare. k, A probe put under the ten- dons of the flexor digitorum sublimis; which are perforated by 1, the flexor digitorum pro- fundus, m m m m, Lumbricales. n, Abduc- tor pollicis. 236 Of the Abdomen. Fig 3. A Fore-view of the foot and Tendons of the Flexores Digitorum. a, Cut extremity of the tendo Achillis. b Upper part of the astragalus, c, Os calcis. d, Tendon of the tibialis anticus. e, Tendon of the extensor pollicis longus. f, Tendon of the peronaeus brevis. g, Tendons of the flex- or digitorum longus, with the nonus Vesalii. h h, The whole of the flexor digitorum bre- vis. Fig. 4. Muscles of the Anus. a a, An out line of the buttocks, and upper part of the thighs, b, The testes contained in the scrotum, c c, Sphincter ani. d, Anus. e, Levator ani. f f, Erector penis, g g, Ac- celerator urinae- h, Corpus cavernosura ure- thra?. Fig. 5. Muscles of the Penis. a a, b, d, e e, f f, h, point the same as in fig. 4. c, Sphincter ani. gg, Transversalis penis. Part III. OF THE ABDOMEN, OR LOWER BELLY. THE abdomen or lower belly, extends from the lower extremity of the sternum, or the hollow, usually called the pit of the sto- mach, and more properly scrobiculus cordis, to the lower part of the trunk- Of the Abdomen. 237 It is distinguished into three divisions call- ed regions; of these the upper one, which is called the epigastric region, begins immediate- ly under the sternum, and extends to within two fingers breadth of the navel, where the middle or umbilical region begins, and reaches to the same distance below the navel. The third, which is called the hypogastric, includes the rest of the abdomen, as far as the os pu- bis. Each of these regions is subdivided into three others ; two of which compose the sides, and the other the middle part of each region. The middle part of the upper region is call- ed epigastrium, and its two sides hypochondria. The middle part of the next region is the um- bilical region, properly so called, and its two sides are the flanks, or iliac regions. Lastly, the middle part of the lower region retains the name of hypogastrium, and its sides are call- ed inguina or groins. The back part of the abdomen bears the name of lumbar region- These are the divisions of the lower belly, which are necessary to be held in remem- brance, as they frequently occur in surgical and anatomical writing. We will now proceed to examine the contents of the abdomen; and after having pointed out the names and ar- rangement of the several viscera contained in it, describe each of them separately- After having removed the skin, adipose membrane, and abdominal muscles, we disco- ver the peritonaeum or membrane that enve- lopes all the viscera of the lower belly- This being opened, the first part that presents itself 238 Of the Abdomen. is the omentum or cawi, floating on the sur- face of the intestines, which are likewise seen every where loose and moist, and making a great number of circumvolutions through the whole cavity of the abdomen. The stomach is placed in the epigastrium, and under the stomach is the pancreas. The liver fills the right hypochondrium, and the spleen is situ- ated in the left. The kidneys are seen about the middle of the lumbar region, and the uri- nary bladder and parts of generation are seat- ed in the lower division of the belly- Sect. I. Of the Peritonaeum. The peritonaeum is a strong simple mem- brane, by which all the viscera of the abdo- men are surrounded, and in some measure supported. Many anatomical writers, particu- larly Winslow, have described it as being com- posed of two distinct membranous laminae; but their description seems to be erroneous. What perhaps appeared to be a second lamina, being found to be simply a cellular coat, which sends off productions to the blood-vessels passing out of the abdominal cavity. The aorta and vena cava likewise derive a covering from the same membrane, which seems to be a part of the cellular membrane we have already de- scribed. The peritonaeum, by its productions and re- duplications, envelopes the greatest part of the abdominal viscera. It is soft, and capable of considerable extension; and is kept smooth and moist by a vapour which is constantly ex- Of the Abdomen. 239 haling from its inner surface, and is returned again into the circulation by the absorbents. This moisture not only contributes to the softness of the peritonaeum, but prevents the attrition, and other ill effects which would otherwise probably be occasioned, by the mo- tion of the viscera upon each other. When this fluid is supplied in too great a quantity, or the absorbents become incapable of carrying it off, it accumulates, and consti- tutes an ascites or dropsy of the belly; and when by any means the exhalation is discon- tinued, the peritonaeum thickens, becomes dis- eased, and the viscera are sometimes found adhering to each other. The peritonaeum is not a very vascular membrane. In a sound state it seems to be endued with little or no feeling, and the nerves that pass through it appear to belong to the abdominal muscles. Sect. II. Of the Omentum. The omentum, epiploon, or cawl, is a dou- ble membrane, produced from the peritonaeum. It is interlarded with fat, and adheres to the stomach, spleen, duodenum, and colon; from thence hanging down loose and floating on the surface of the intestines. Its size is different in different subjects. In some it descends as low as the pelvis, and it is commonly longer at the left side than the right. This part, the situation of which we have just now described, was the only one known to the ancients under the name of epiploon; 246 Of the Abdomen. but at present we distinguish three omenta, viz. omentum magnum colico gastricnm, omen- tum parvum hepatico gastricum, and omentum colicum. They all agree in being formed of two very delicate laminae, separated by a thin layer of cellular membrane. The omentum magnum colico gastricum, of which we have already spoken, derives its ar- teries from the splenic and hepatic. Its veins terminate in the vena portae. Its nerves, which are very few, come from the splenic and he- patic plexus. The omentum parvum hepatico gastricum, abounds less with fat than the great epiploon. It begins at the upper part of the duodenum, extends along the lesser curvature of the sto- mach as far as the oesophagus, and terminates about the neck of the gall-bladder, and behind the left ligament of the liver, so that it covers the lesser lobe; near the beginning of which we may observe a small opening, first describ- ed by Winslow, through which the whole pouch may easily be distended with air.* The vessels of the omentum parvum are derived chiefly from the coronary stomachic arteries and veins. The omentum colicum begins at the fore part of the ccecum and right side of the colon. It appears as a hollow conical appendage to these intestines, and usually terminates at the back of the omentum magnum. It seems to * This membranous bag, though exceedingly thin and trans- parent, is found capable of supporting mercury, thrown into ;t by the same channel. Of the Abdomen. 241 be nothing more than a membranous coat of the cdecum and colon, assuming a conical shape when distended with air. The uses of the omentum are not yet satis- factorily determined. Perhaps by its softness and looseness it may serve to prevent those adhesions of the abdominal viscera, which have been found to take place when the fat of the omentum has been much wasted. Some authors have supposed, that it assists in the preparation of bile; but this idea is founded merely on conjecture. Sect. III. Of the Stomach. The stomach is a membranous and muscu- lar bag, in shape not unlike a bag-pipe, lying across the upper part of the abdomen, and in- clining rather more to the left than the right side. It has two orifices, one of which receives the end of the oesophagus, and is called the cardia, and sometimes the left and upper ori- fice of the stomach ; though its situation is not much higher than the other, which is styled the right and inferior orifice, and more com- monly the pylorus; both these openings are more elevated than the body of the stomach. The aliment passes down the oesophagus in- to the stomach through the cardia, and after having undergone the necessary digestion, passes out at the pylorus where the intesti- nal canal commences. The stomach is composed of four tunics or coats, which are so intimately connected toge- H h 242 Of the Abdomen. ther that it requires no little dexterity in the anatomist to demonstrate them. The exterior one is membranous, being derived from the peritonaeum.—The second is a muscular tu- nic, composed of fleshy fibres which are in the greatest number about the two orifices.—The third is called the nervous coat, and within this is the villous or velvet-like coat which composes the inside of the stomach. The two last coats being more extensive than the two first, form the folds, which are observed every where in the cavity of this viscus, and more particularly about the pylo- rus ; where they seem to impede the too has- ty exclusion of the aliment, making a conside- rable plait, called valvula pylori. The inner coat is constantly moistened by a mucus, which approaches to the nature of the saliva, and is called the gastric juice; this li- quor has been supposed to be secreted by cer- tain minute glands* seated in the nervous tu- nic, whose excretory ducts open on the sur- face of the villous coat. The arteries of the stomach called the gas- tric arteries are principally derived from the cseliac; some of its veins pass to the splenic, and others to the vena portae; and its nerves are chiefly from the eighth pair or par vagum. * Heister, speaking of these glands, very properly says, " in porcis facile, in homine raro observantur ;" for although many anatomical writers have described their appearance and figure, yet they do not seem to have been hitherto satisfactorily demon- strated in the human stomach ; and the gastric juice is now more generally believed to be derived from the exhalent arteries of the stomach. Of the Abdomen. 243 The account given of the tunics of the sto- mach may be applied to the whole alimentary canal; for both the oesophagus and intestines are, like this viscus, composed of four coats. Before we describe the course of the ali- ment and the uses of the stomach, it will be necessary to speak of other parts which assist in the process of digestion. Sect. IV. Of the Oesophagus. The oesophagus or gullet is a membra- nous and muscular canal, extending from the bottom of the mouth to the upper orifice of the stomach.—Its upper part where the ali- ment is received is shaped somewhat like a funnel, and is called the pharynx. From hence it runs down close to the bo- dies of the vertebrae as far as the diaphragm, in which there is an opening through which ix passes, and then terminates in the stomach about the eleventh or twelfth vertebra of the back. The oesophagus is plentifully supplied with arteries from the external carotid, bronchial, and superior intercostal arteries.; its veins empty themselves into the vena azygos, inter- nal jugular, and mammary veins, he. Its nerves are derived chiefly from the eighth pair- We likewise meet with a mucus in the oeso- phagus, which every where lubricates its in- ner surface, and tends to assist in deglutition. —This mucus seems to be secreted by very minute glands, like the mucus in other parts of the alimentary canal. 244 Of the Abdomen. Sect. V. Of the Intestines. The intestines form a canal, which is usu- ally six times longer than the body to which it belongs. This canal extends from the py- lorus, or inferior orifice of the stomach, to the anus. It will be easily understood, that a part of such great length must necessarily make many circumvolutions, to be confined with so many other viscera within the cavity of the lower belly. Although the intestines are in fact, as we have observed, only one long and extensive canal, yet different parts have been distin- guished by different names. The intestines are first distinguished into two parts, one of which begins at the sto- mach, and is called the thin or small intestines^ from the small size of the canal, when com- pared with the other part, which is called the large intestines, and includes the lower portion of the canal down to the anus. Each of these parts has its subdivisions.— The small intestines being distinguished into duodenum, jejunum, and ilium, and the larger portion into caecum, colon, and rectum. The small intestines fill the middle and fore parts of the belly, while the large intestines fill the sides and both the upper and lower parts of the cavity. The duodenum, which is the first of the small intestines, is so called, because it is about 12 inches long. It begins at the pylo- Of the Abdomen. 245 rus and terminates in the jejunum, which is a part of the canal observed to be usually more empty than t;.e otiier intestines.—This appear- ance gives it its name, and likewise serves to point out where it begins. The next division is the ilium, which of it- self exceeds the united length of the duode- num and jejunum, and has received its name from its numerous circumvolutions. The large circumvolution of the ilium covers the first of the large intestines called the cajcum,* which seems properly to belong to the colon, being a kind of pouch ofabovufour fingers in width, and nearly of the same length, having exteriorly a little appendix, called appendix cceci. The caecum is placed in the cavity of the os ilium on the right side, and terminates in the colon, which is the largest of all the in- testines. This intestine ascends by the right kidney to which it is attached, passes under the hol- low part of the liver, and the bottom of the stomach, to the spleen, to which it is likewise secured, as it is also to the left kidney; and from thence passes down towards the os sa- crum, where, from its straight course, the ca- nal begins to take the name of rectum. There are three ligamentous bands extend- ing through the whole length of the colon, * Anatomists have differed with respect to this division of the intestines.—The method here followed is now generally adopted ; but there are authors who allow the name of cacum only to the little appendix, which has likewise been called the venniform ap- pendix, from its resemblance to a worm in size and length. 246 Of the Abdomen. which, by being shorter than its two inner coats, serve to increase the plaits on the in- ner surface of this gut. The anus which terminates the intestinum rectum, is furnished with three muscles; one of these is composed of circular fibres, and from its use in shutting the passage of the anus is called sphincter ani. The other two are the levatores ani, so call- ed, because they elevate the anus after dejec- tion. When these by palsy, or any other dis- ease lose the power of contracting, the anus prolapses; and when the sphincter is affected by similar causes, the faeces are voided invo- luntarily. It has been already observed, that the in- testinal canal is composed of four tunics; but it remains to be remarked, that here, as in the stomach, the two inner tunics being more extensive than the other two, form the plaits which are to be seen in the inner surface of the intestines, and are called valvules conniven- ies. Some authors have considered these plaits as tending to retard the motion of the faeces, in order to afford more time for the separa- tion of the chyle; but there are others who attribute to them a different use: they con- tend, that these valves, by being naturally in- clined downwards, cannot impede the descent of the faeces, but that they are intended to pre- vent their return upwards. They are probably destined for both these uses; for although these folds incline to their lower side, yet the inequalities they occasion Of the Abdomen. 247 in the canal are sufficient to retard, in some measure, the progressive motion of the faeces, and to afford a greater surface for the absorp- tion of chyle, and their natural position seems to oppose itself to the return of the aliment. Besides these valvule conniventes, there is one more considerable than the rest, called the valve of the colon ; which is found at that part of the canal where the intestinum ilium is join- ed to the colon. This valve permits the ali- mentary pulp to pass downwards, but serves to prevent its return upwards ; and it is by this valve, that glysters are prevented from passing into the small intestines.* Of the little vermiform appendix of the cae- cum, it will be sufficient to say, that its uses have never yet been ascertained. In birds we meet with two of these appendices. The intestines are lubricated by a constant supply of mucus, which is probably secreted by very minute follicles.f This mucus pro- motes the descent of the alimentary pulp, and in some measure defends the inner surface of * This is not invariably the case, for the contents of a gly- ster have been found not only to reach the small intestines, but to be voided at the mouth. Such instances, however, are not common. f Some writers have distinguished these glands into miliary, lenticular, &c.—Brunner and Peyer were the first anatomists who described the glands of the intestines, and their descrip- tions were chiefly taken from animals, these glandular appear- ances not seeming to have been hitherto satisfactorily pointed out in the human subject.—It is now pretty generally believed, that the mucus which every where lubricates tire alimentary canal, is exhaled from the minute ends of arteries; and that these ex- tremities first open into a hollow vesicle, from whence the depo- sited juice of several branches flows out through one commo.t orifice. 248 Of the Abdomen. the intestines from the irritation to which it would, perhaps, otherwise be continually ex- posed from tiie aliment; and which, when in a certain degree, excites a painful disorder cahed colic, a name given to the disease, be- cause us most usual seat is in the intestinum colon. The intestines are likewise frequently dis- tended with air, and this distention sometimes occasions pain, and constitutes the flatulent colic. The arteries of the intestines are continu- ations of the mesenteric arteries, which are derived in two considerable branches from the aorta.—The redundant blood is carried back into the vena portarum. In the rectum the veins are called hemor- rhoidal, and are there distinguished into inter- nal and external: the first are branches of the inferior mesenteric vein, but the latter pass into other veins. Sometimes these veins are distended with blood from obstructions, from weakness of their coats, or from other causes, and what we call the hemorrhoids takes place. In this disease they are sometimes ruptured; and the discharge of blood which consequent- ly follows, has probably occasioned them to be called hemorrhoidal veins. The nerves of the intestines are derived from the eighth pair. Sect. VI. Of the Mesentery. The name of the mesentery implies its sim- ulation amidst the intestines. It is in fact a Of the Abdomen, 249 part of the peritonaeum, being a reduplica- tion * of that membrane from each side of the lumbar vertebrae, to which it is firmly attach- ed, so that it is formed of two laminae, con- nected to each other by cellular membrane. The intestines, in their different circumvo- lutions, form a great number of arches, and the mesentery accompanies them through all these turns; but by being attached only to the hollow part of each arch, it is found to have only a third of the extent of the intestines. That part of this membrane which accom- panies the small intestines is the mesentery, properly so called ; but those parts of it which are attached to the colon and rectum are dis- tinguished by the names of meso-colon and me- so-rectum. There are many conglobate glands dispers- ed through this double membrane, through which the lacteals and lymphatics pass in their way to the thoracic duct. The blood-vessels of the mesentery were described in speaking of the intestines. I i * He who only reads of the reduplication of membranes, will perhaps not easily understand how the peritf nxum and pleura are reflected over the viscera in their several cavities ; for one of these serves the same purposes in the thorax that the other does in the abdomen. This disposition, for the dis- covery of which we are indebted to modern anatomists, con- stitutes a curious part of anatomical knowledge : but the stu- dent, unaided by experience, and assisted only by what the li- mits of this work would permit us to say on the occasion, would probably imbibe only confused ideas of the matter ; and it v-ill perfectly answer the present purpose, if he considers the me- sentery as a membrane attached bv one of its sides to the lum- bar vertebrae, and by the other to the intestines. 250 Of the Abdomen. This membrane, by its attachment to the vertebrae, serves to keep the intestines in their natural situation. The idea usually formed of the colic called miserere, is perfectly errone- ous ; it being impossible that the intestines can be twisted, as many suppose they are, in that disease, their attachment to the mesente- ry effectually preventing such an accident— but a disarrangement sometimes takes place in the intestinal canal itself, which is produc- tive of disagreeable and sometimes fatal con- sequences.—This is by an introsusception of the intestine, an idea of which may be easily formed, by taking the finger of a glove, and involving one part of it within the other. If inflammation takes place, the stricture in this case is increased, and the peristaltic mo- tion of the intestines (by which is meant the progressive motion of the faeces downwards) is inverted, and what is called the iliac passion takes place. The same effects may be occa- sioned: by a descent of the intestine,, or of the omentum either with it or by itself, and thus constituting what is called an hernial rupture; a term by which in general is meant the falling down or protrusion of any part of the intes- tine or omentum, which ought naturally to be contained within the cavity of the belly. To convey an idea of the manner in which such a descent takes place, it will be necessa- ry to observe, that the lower edge of the ten- don of the musculus obliquus externus, is stretched from the fore-part of the os ilium or haunch-bone of the os pubis, and consti- tutes what is called PouparVs or Fallopius's li- Of the Abdomen. 251 gament, forming an opening, through which pass the great crural artery and vein. Near the os pubis the same tendinous fibres are se- parated from each other, and form an opening on each side, called the abdominal ring, through which the spermatic vessels pass in men, and the ligamenta uteri in women. In consequence of violent efforts, or perhaps of natural causes, the intestines are found sometimes to pass through these openings; but the peritonaeum which incloses them when in their natural ca- vity, still continues to surround them even in their descent. This membrane does not be- come torn or lacerated by the violence, as might be easily imagined; but its dilatibility enables it to pass out with the viscus, which it incloses as it were in a bag, and thus forms what is called the hernial sac. If the hernia be under Poupart's ligament, it is called femoral; if in the groin, inguinal ;* and scrotal, if in the scrotum. Different names are likewise given to the hernia as the con- tents of the sac differ, whether of omentum only or intestine, or both:—but these defini- tions more properly belong to the province of surgery. Sect. VII. Of the Pancreas. The pancreas is a conglomerate gland placed behind the bottom of the stomach, towards the first vertebra of the loins; shaped * The hernia congenita will be considered with the male or- gans of generation, with which it is intimately connected. 252 Of the Abdomen. like a dog's tongue, with its point stretched out towards the spleen, and its other end ex- tending towards the duodenum. It is about eight fingers breadth in length, two or three in width, and one in thickness. This viscus, which is of a yellowish colour, somewhat inclined to red, is covered with a membrane which it derives from the peritonae- um. Its arteries, which are rather numerous than large, are derived chiefly from the sple- nic and hepatic, and its veins pass into the veins of the same name.—Its nerves are de- rived from the intercostal. The many little glands of which it has been observed the pancreas is composed, all serve to secrete a liquor called the pancreatic juice, which in its colour, consistence, and other pro- perties, does not seem to differ from the sali- va. Each of these glands sends out a little excretory duct, which, uniting with others, help to form larger ducts; and all these at last terminate in one common excretory duct (first discovered by Virtsungus in 1642), which runs through the middle of the gland, and is now usually called ductus pancreaticus Virt- sungi. This canal opens into the intestinum duodenum, sometimes by the same orifice with the biliary duct, and sometimes by a distinct opening. The liquor it discharges being of a mild and insipid nature, serves to dilute the alimentary pulp, and to incorporate it more easily with the bile. Of the Abdomen. 253 Sect. VIII. Of the Liver. The liver is a viscus of considerable size, and of a reddish colour; convex superiorly and anteriorly where it is placed under the ribs and diaphragm, and of an unequal sur- face posteriorly. It is chiefly situated in the right hypochondrium, and under the false ribs; but it likewise extends into the epigastric re- gion, where it borders upon the stomach. It is covered by a production of the peritonaeum, which serves to attach it by three of its redu- plications to the false ribs. These reduplica- tions are called ligaments, though very differ- ent in their texture from what are called by the same name in other parts of the body. The umbilical cord, too, which in the foetus is per- vious, gradually becomes a simple ligament after birth ; and, by passing to the liver, serves likewise to secure it in its situation. At the posterior part of this organ where the umbilical vessels enter, it is found divid- ed into two lobes. Of these, the largest is placed in the right hypochondrium ; the other, which covers part of the stomach, is called the little lobe. All the vessels winch go to the liver pass in at the fissure we have mention- ed ; and the production of the peritoneum, which invests the liver, was described by Glis- son, an English anatomist, as accompanying them in their passage, and surrounding them like a glove ; hence this production has been commonly known by the name of capsula of Glisson: but it appears to be chiefly a continu- 254 Of the Abdomen. ation of the cellular membrane which covers the vena portae ventralis. The liver was considered by the ancients as an organ destined to prepare and perfect the blood; but later discoveries have proved, that this opinion was wrong, and that the liver is a glandular substance formed for the secretion of the bile. The blood is conveyed to the liver by the hepatic artery and the vena portae. This is contrary to the mode of circulation in other parts, where veins only serve to carry off the redundant blood : but in this viscus the hepa- tic artery, which is derived from the caeliac, is principally destined for its nourishment; and tfee vena portae, which is formed by the union of the veins from most of the abdominal vis- cera, furnishes the blood from which the bile is chiefly to be separated; so that these two series of vessels serve very distinct purposes. The vena portae, as it is ramified through the liver, performs the office both of a vein and an artery; for like the former it returns the blood from the extremities of arteries, while as the latter it prepares it for secretion. The nerves of the liver are branches of the intercostal and par vagum. The bile, after be- ing separated from the mass of blood, in a manner of which mention will be made in another place, is conveyed out of this organ by very minute excretory ducts, called poribi- liarii; these uniting together like the excreto- ry ducts in the pancreas, gradually form larger ones, which at length terminate in a considera- ble canal called ductus hepaticus. Of the Abdomen. 25 5s. Sect. IX. Of the Gall-bladder. The gall-bladder is a little membranous bag, shaped like a pear, and attached to the posterior and almost inferior part of the great lobe of the liver. It has two tunics; of which the exterior one is a production of the peritonaeum. The interior, or villous coat, is supplied with a mu- cus that defends it from the acrimony of the bile. These two coverings are intimately con- nected by means of cellular membrane, which from its firm glistening appearance has gene- rally been spoken of as a muscular tunic. The gall-bladder is supplied with bloods vessels from the hepatic arteries. These branches are called the cystic arteries, and the cystic veins carry back the blood. Its nerves are derived from the same origin as those of the liver. The neck of the gall-bladder is continued in the form of a canal called ductus cysticus* which soon unites with the ductus hepaticus we described as the excretory duct of the li- ver; and forming one common canal, takes the name of ductus coledochus communis, through which both the cystic and hepatic bile are dis^ charged into the duodenum. This canal opens into the intestine in an oblique direction, first passing through the exterior tunic, and then piercing the other coats after running between each of them a very little way. This cecono- my serves two useful purposes;—to promote the discharge of bile and to prevent its return. 256 Of the Abdomen. The bile may be defined to be a natural li- quid soap, somewhat unctuous and bitter, and of a yellowish colour, which easily mixes with water, oil, and vinous spirits, and is capable of dissolving resinous substances. From some late experiments made by M. Cadet,* it ap- pears to be formed of an animal oil, combined with the alkaline base of sea-salt, a salt of the nature of milk, and a calcareous earth which is slightly ferruginous. Its definition seems sufficiently to point out the uses for which it is intended^ It blends the alimentary mass, by dividing and attenu- ating it; corrects the too great disposition to acescency, which the aliment acquires in the stomach; and finally, by its acrimony, tends to excite the peristaltic motion of the intes- tines. After what has been said, it will be conceiv- ed that there are two sorts of bile; one of which is derived immediately from the liver through the hepatic duct, and the other from the gall-bladder. These two biles, however, do not essentially differ from each other. The hepatic bile indeed is milder, and more liquid than the cystic, which is constantly thicker and yellower; and by being bitterer, seems to possess greater activity than the other. Every body knows the source of the hepatic bile, that it is secreted from the mass of blood by the liver; but the origin of the cystic bile * Mem. de 1* Acad, des Sciences, 1767. f The ancients, who were not acquainted with the real use of the liver, considered the bile as an excrementilious and use- less fluid. Of the Abdomen. 257 has occasioned no little controversy ampngst anatomical writers. There are some who con- tend, that it is separated in the substance of the liver, from whence it passes into the gall- bladder through particular vessels. In deer, and in some other quadrupeds, as well as in several birds and fishes, there is an evident communication, by means of particular ves- sels, between the liver and the gall-bladder. Bianchi, Winslow, and others, have asserted the existence of such vessels in the human subject, and named them hepaticystic ducts; but it is certain that no such ducts exist.—In obstructions of the cystic duct, the gall-blad- der has been found shrivelled and empty: so that we may consider the gall-bladder as a reser- voir of hepatic bile ; and that it is an establish- ed fact that the whole of the bile contained in the gall-bladder is derived from the liver; that it passes from the hepatic to the cystic duct, and from that to the gall-bladder. The differ- ence in the colour, consistence, and taste of the bile, is merely the consequence of stagna- tion and absorption. When the stomach is distended with aliment, this reservoir under- goes a certain degree of compression, and the bile passes out into the intestinal canal ; and in the efforts to vomit, the gall-bladder seems to be constantly affected, and at such times dis- charges itself of its contents. Sometimes the bile concretes in the gall- bladder, so as to form what are called gall- Kk 258 Of the Abdomen. stones.* When these concretions pass into the cystic duct, they sometimes occasion ex- quisite pain, by distending the canal in their way to the duodenum; and by lodging in the ductus choledochus communis, and obstruct- ing the course of the bile, this fluid will be absorbed, and by being carried back into the circulation occasion a temporary jaundice. Sect. X. Of the Spleen. The spleen is a soft and spongy viscus, of a bluish colour, and about five or six fingers breadth in length, and three in width, situated in the left hypochondrium, between the sto- mach and the false ribs. That side of it which is placed on the side of the ribs is convex ; and the other, which is turned toward the stomach, is concave. The splenic artery, which is a branch from the caeliac, supplies this viscus with blood, and a vein of the same name carries it back into the vena portae. Its nerves are derived from a particular plexus called the splenic, which is formed by branches of the intercostal nerve, and by the eighth pair, or par vagum. The ancients, who supposed two sorts of bile, considered the spleen as the receptacle of what they called atra bilis. Havers, who * These concretions sometimes remain in the gall-bladder without causing any uneasiness. Dr. Heberden relates, that a gall-stone weighing two drams was found in the gall-bladder of the late Lord Bath, though he had never complained of the jaundice, nor of any disorder which he could attribute to that cause. Med. Trans. Vol. ii. Of the Abdomen. 259 wrote professedly on the bones, determined its use to be that of secreting the synovia; and the late Mr. Hewson imagined, that it concur- red with the thymus and lymphatic glands of the body in forming the red globules of the blood. All these opinions seem to be equally fanciful. The want of an excretory duct has occasioned the real use of this viscus to be still doubtful. Perhaps the blood undergoes some change in it, which may assist in the pre- paration of the bile. This is the opinion of the generality of modern physiologists; and the great quantity of blood with which it is supplied, together with the course of its veins into the vena portae, seem to render this no- tion probable. Sect. XI. Of the Glandule Renale-s, Kidneys, and Ureters. The glandulae renales, which were by the ancients supposed to secrete the atra bilis, and by them named capsula atrabilares, are two flat bodies of an irregular figure, one on each side between the kidney and the aorta. In the foetus they are as large as the kidneys: but they do not increase afterwards in propor- tion to those parts; and in adults and old peo- ple they are generally found shrivelled, and much wasted. They have their arteries and veins. Their arteries usually arise from the splenic or the emulgent, and sometimes from the aorta; and their veins go to the neighbour- ing veins, or to the vena cava. Their nerves are branches of the intercostal. 260 Of the Abdomen. The use of these parts is not yet perfectly known. In the foetus the secretion of urine must be in a very small quantity, and a part of the blood may perhaps then pass through these channels, which in the adult is carried to the kidneys to supply the matter of urine. The kidneys are two in number, situated one on the right and the other on the left side in the lumbar region, between the last false rib and the os ilium, by the sides of the verte- brae. Each kidney in its figure resembles a sort of bean, which from its shape is called kidney-bean. The concave part of each kidney is turned towards the aorta and vena cava as- cendens. They are surrounded by a good deal of fat, and receive a coat from the peri- tonaeum ; and when this is removed, a very fine membrane is found investing their sub- stance and the vessels which ramify through them. Each kidney has a considerable artery and vein, which are called the emulgent. The ar- tery is a branch from the aorta, and the vein passes into the vena cava. Their nerves, which every where accompany the blood- vessels, arise from a considerable plexus, which is derived from the intercostal. In each kidney, which in the adult is of a pretty firm texture, there are three substances to be distinguished.* The outer part is glan- dular or cortical, beyond this is the vascular * The kidneys in the foetus are distinctly lobulated ; but in the adult they become perfectly firm, smooth, and regular. Of the Abdomen. 261 or tubular substance, and the inner part is pa- pillary or membranous. It is in the cortical part of the kidney that the secretion is carried on; the urine being here received from the minute extremities of the capillary arteries, is conveyed out of this cortical substance by an infinite number of ve- ry small cylindrical canals or excretory ves- sels, which constitute the tubular part. These tubes, as they approach the inner substance of the kidney gradually unite together; and thus forming larger canals, at length termi- nate in ten or twelve little protuberances call- ed papilla, the orifices of which may be seen without the assistance of glasses. These pa- pillae open into a small cavity or reservoir called the pelvis of the kidney, and formed by a distinct membranous bag which embraces the papillae. From this pelvis the urine is conveyed through a membranous canal which passes out from the hollow side of the kidney, a little below the blood-vessels, and is called ureter. The ureters are each about as large as a common writing-pen. They are somewhat curved in their course from the kidneys, like the letter/, and at length terminate in the pos- terior and almost inferior part of the bladder, at some distance from each other. They pass into the bladder in the same manner as the ductus choledochus communis passes into the intestinum duodenum, not by a direct passage, but by an oblique course between the two coats; so that the discharge of urine into the bladder is promoted, whilst its return is pre- 262 Of the Abdomen. vented. Nor does this mode of structure pre- vent the passage of fluids only from the blad- der into the ureters, but likewise air:—for air thrown into the bladder inflates it, and it con- tinues to be distended if a ligature is passed round its neck; which seems to prove suffici- ently that it cannot pass into the ureters. Sect. XII. Of the Urinary Bladder. The urinary bladder is a membranous and muscular bag of an oblong roundish shape, situated in the pelvis, between the os pubis and intestinum rectum in men, and between the os pubis and uterus in women. Its up- per and widest part is usually called the bot- tom, its narrow part the neck of the bladder; the former only is covered by the peritonaeum. The bladder is formed of three coats, con- nected together by means of cellular mem- brane. The external or peritonaeal, is only a partial one, covering the upper and back part of the bladder. The middle, or muscular coat, is composed of irritable, and of course muscular fibres, which are most collected around the neck of the bladder, but not so as to form a distinct muscle, or sphincter, as the generality of anatomists have hitherto sup- posed. The inner coat, though much smoother, has been said to resemble the villous tunic of the intestines, and like that is provided with a mucus, which defends it against the acrimony of the urine. Of the Abdomen. 263 It will be easily conceived from what has been said, that the kidneys are two glandular bodies, through which a saline and excremen- titious fluid called urine is constantly filtering from the mass of blood. While only a small quantity of urine is col- lected in the bladder, it excites no kind of un- easiness ; but when a greater quantity is ac- cumulated, so that the bladder is distended in a certain degree, it excites in us a certain sen- sation, which brings on as it were a voluntary contraction of the bladder to promote its dis- charge.—But this contraction is not effected by the muscular fibres of the bladder alone: for all the abdominal muscles contract in obe- dience to our will, and press downwards all the viscera of the lower belly ; and these pow- ers being united, at length overcome the re- sistance of the fibres surrounding the neck of the bladder, which dilates and affords a pas- sage to the urine through the urethra. The frequency of this evacuation depends on the quantity of urine secreted; on the de- gree of acrimony it possesses; on the size of the bladder, and on its degree of sensibility. The urine varies much in its colour and contents. These varieties depend, on age, sex, climate, diet, and other circumstances. In in- fants it is generally a clear watery fluid, with- out smell or taste. As we advance in life, it acquires more colour and smell, and becomes more impregnated with salts. In old people it becomes still more acrid and fetid. In a healthy state it is nearly of a straw co- lour,—After being kept for some time, it de- 264 Of the Abdomen. posites a tartarous matter, which is found to be composed chiefly of earth and salt, and soon incrusts the sides of the vessel in which it is contained. While this separation is tak- ing place, appearances like minute fibres or threads of a whitish colour may be seen in the middle of the urine, and an oily scum observ- ed floating on its surface. So that the most common appearances of the urine are suffici- ent to ascertain that it is a watery substance, impregnated with earthy, saline, and oily par- ticles. The urine is not always voided of the same colour and consistence: for these are found to depend on the proportion of its watery part to that of its other constituent principles.—Its co- lour and degree of fluidity seem to depend on the quantity of saline and inflammable parti- cles contained in it: so that an increased pro- portion of those parts will constantly give the urine a higher colour, and add to the quanti- ty of sediment. The variety in the appearance of the urine, depends on the nature and quantity of solid and fluid aliment we take in; and it is like- wise occasioned by the different state of the urinary vessels, by which we mean the chan- nels through which it is separated from the blood, and conveyed through the pelvis into the ureters. The causes of calculous concre- tions in the urinary passages, are to be looked for in the natural constitution of the body, mode of life, he. It having been observed, that after drink- ing any light wine or Spa water, it very soon Of the Abdomen. 26 5 passed off by urine, it has been supposed by some, that the urine is not altogether convey- ed to the bladder by the ordinary course of circulation, but that there must certainly exist some other shorter means of communication, perhaps by certain vessels between the sto- mach and the bladder, or by a retrograde motion in the lymphatics. But it is certain, that if we open the belly of a dog, press out the urine from the bladder, pass a ligature round the emulgent arteries, and then sew up the abdo- men, and give him even the most diuretic li- quor to drink, the stomach and other chan- nels will be distended with it, but not a drop of urine will be found to have passed into the bladder; or the same thing happens when a ligature is thrown round the two ureters. This experiment then seems to be a sufficient proof, that all the urine we evacuate, is conveyed to the kidneys through the emulgent arteries, in the manner we have described.—It is true, that wine and other liquors promote a speedy eva- cuation of urine: but the discharge seems to be merely the effect of the stimulus they occa- sion ; by which the bladder and urinary parts are solicited to a more copious discharge of the urine, which was before in the body, and not immediately of that which was last drank ; and this increased discharge, if the supply is kept up, will continue : nor will this appear won- derful, if we consider the great capacity of the vessels that go to the kidneys ; the constant sup- ply of fresh blood that is essential to health ; and the rapidity with which it is incessantly L 1 266 Of the Abdomen. circulated through the heart to all parts of the body. Sect. XIII. Of Digestion. We are now proceeding to speak of diges- tion, which seems to be introduced in this place with propriety, after a description of the abdominal viscera, the greater part of which contribute to this function. By digestion is to be understood, the changes the aliment under- goes for the formation of chyle :—these chan- ges are effected in the mouth, stomach, and small intestines. The mouth, of which every body has a ge- neral knowledge, is the cavity between the two jaws, formed anteriorly and laterally by the lips, teeth, and cheeks, and terminating posteriorly in the throat. The lips and cheeks are made up of fat and muscles, covered by the cuticle, which is con- tinued over the whole inner surface of the mouth, like a fine and delicate membrane.— Besides this membrane, the inside of the mouth is furnished with a spongy and very vascular / substance called the gums, by means of which the teeth are secured in their sockets. A simi- lar substance covers the roof of the mouth, and forms what is called the velum pendulum palati, which is fixed to the extremity of the arch formed by the ossa maxillaria and ossa pa- lati, and terminates in a soft, small, and coni- cal body, named uvula ; which appears, as it were, suspended from the middle of the arch over the basis of the tongue. Of the Abdomen. 267 The velum pendulum palati performs the office of a valve between the cavity of the mouth and the pharynx, being moved by se- veral muscles.* The tongue is composed of several mus- clesf which enable it to perform a variety of motions for the articulation of the voice ; for the purposes of mastication; and for convey- ing the aliment into the pharynx. Its upper part is covered with papillae, which constitute the organ of taste, and are easily to be dis- tinguished; it is covered by the same mem- brane that lines the inside of the mouth, any its gela- tinous principle, it seems to be much -better calculated than a pure and watery fluid would be, for such it has been supposed to be by some anatomists. The mouths of the lymphatics and lacteals,hy acting as capillary tubes, seem to absorb the lymph and chyle somewhat in the same manner as a capillary tube of glass, when put into a ba- son of water, is enabled to attract the water into it to a certain height; but it is probable that they 284 Of the Abdomen. likewise possess a living power, which assists in performing this office. In the human body the lymph, or the chyle, is probably conveyed upon this principle as far as the first pair of valves, which seem to be placed not far from the orifice of the absorbing vessel, whether lymphatic or lacteal; and the fluid will then be propelled forwards, by a continuation of the absorption at the orifice. But this does not seem to be the only inducement to its progress towards the thoracic duct; these vessels have probably a muscular coat, which may serve to press the fluid forwards from one pair of valves to another ; and as the large lymphatic vessels and the thoracic duct are placed close to the large arteries, which have a considerable pul- sation, it is reasonable to suppose, that they derive some advantages from this situation. Sect. XV. Of the Generative Organs; of Conception, he. J. 1. The Male Organs. The male organs of generation have been usually divided into the parts which serve to prepare the semen from the blood, and those which are distended to convey it into the womb* But it seems to be more proper to distinguish them into the preparing, the containing, and the expelling parts, which are the different offices of the testes, the vesicula seminales, and the penis ; and this is the Order in which we propose to describe them. Of the Abdomen. 285 The testes are two glandular bodies, serving to secrete the semen from the blood. They are originally formed and lodged within the ca- vity of the abdomen; and it is not till after the child is born, or very near that time, that they begin to pass into the groin, and from thence into the scrotum.* By this disposition they are very wisely protected from the inju- ries to which they would be liable to be ex- posed, from the different positions of the child at the time of parturition. The testicles in this state are loosely at- tached to the psoae muscles, by means of the peritonaeum by which they are covered; and they are at this time of life connected in a very particular manner to the parietes of the abdomen, and likewise to the scrotum, by means of a substance which Mr. Hunter calls the ligament or gubernaculum testis, because it connects the testis with the scrotum, and di- rects its course in its descent. This guber- * It sometimes happens in dissecting ruptures, that the intes- tine is found in the same sac, and in contact with the testis. This appearance was at first attributed to a supposed laceration of the peritonaeum ; but later observations, by pointing out the situation of the testicles in the foetus, have led to prove, that the testis, as it descends into the scrotum, carries with it a por- tion or elongation of the peritonaeum, which becomes its tunica vaginalis, or a kind of sac, in which the testicle is lodged, as will be explained in the course of this section. The communica- tion between this sac and the cavity of the abdomen, is usually soon cut off; but in some subjects it continues open during life; and when an hernia or descent of the intestine takes place in such a subject, it does not push down a portion of the peritonaeum before it, as it must otherwise necessarily do, but passes at once through this opening, and comes in contact with the .naked tes- ticle, constituting that particular species of rupture called hernia congenita. 286 Of the Abdomen. naculum is of a pyramidal form, with its bul- bous head fixed to the lower end of the testis and epididymis, and loses its lower and slen- der extremity in the cellular membrane of the scrotum. It is difficult to ascertain what the structure and composition of this gubernacu- lum is, but it is certainly vascular and fibrous; and, from certain circumstances, it would seem to be in part composed of the cremaster mus- cle, running upwards to join the lower end of the testis. We are not to suppose that the testicle, when descended into the scrotum, is to be seen loose as a piece of gut or omentum would be in a common hernial sac. We have already observed, that during its residence in the cavity of the abdomen it is attached to the peritonaeum, which descends with it; so that when the sac is completed in the scrotum, the testicle is at first attached only to the posterior part of it, while the fore part of it lies loose, and for some time affords a communication with the abdomen. The spermatic chord, which is made up of the spermatic artery and vein, and of the vas deferens or excretory duct of the testis, is closely attached behind to the posterior part of this elongation of the perito- naeum- But the fore part of the peritoneal sac, which is at first loose and not attacfied to the testicle, closes after a certain time, and becomes united to the posterior part, and thus perfectly surrounds the testicle as it were in a purse. The testicles of the foetus differ only in their size and situation from those of the adult. In Of the Abdomen. 287 their passage from the abdomen they descend through the abdominal rings into the scrotum, where they are supported and defended by va- rious integuments. What the immediate cause of this descent is, has not yet been satisfactorily determined. It has been ascribed to the effects of respira- tion, but the testicles have sometimes been found in the scrotum before the child has breathed; and it does not seem to be occasi- oned by the action of the cremaster muscle, because the same effect would be liable to happen to the hedge-hog, and some other quadrupeds, whose testicles remain in the ab- domen during life. The scrotum, which is the external or com- mon covering of both testicles, is a kind of sac formed by the common integuments, and ex- ternally divided into two equal parts by a pro- minent line called raphe. In the inner part of the scrotum we meet with a cellular coat called dartos,* which by its duplicature divides the scrotum into two equal parts, and forms what is called septum scroti, which corresponds with the raphe. The collapsion which is so often observed to take place in the scrotum of the healthy sub- ject, when excited by cold or by the stimulus of venery, seems to be very properly attribute * The dartos has usually been considered as a muscle, and h described as such both by Douglas and Winslow. But there being no part of the scrotum of the human subject which can be said to consist of muscular fibres, Albinus and Haller have very properly omitted to describe the dartos. as a muscle, and consider it merelv as a cellular coat. 288 Of the Abdomen. ed to the contractile motion of the skin, and not to any muscular fibres, as is the case in dogs and some other quadrupeds. The scrotum, then, by means of its septum, is found to make two distinct bags, in which the testicles, invested by their proper tunics, are securely lodged and separated from each other. These coats are the cremaster, the tunica vaginalis, and the tunica albuginea. The first of these is composed of muscular fibres, and is to be considered only as a par- tial covering of the testis; for it surrounds only the spermatic chord, and terminates upon the upper and external parts of the tunica va- ginalis testis, serving to draw up and suspend the testicle.* The tunica vaginalis testis has already been described as being a thin pro- duction of the peritonaeum, loosely adhering every where to the testicle, which it includes as it were in a bag. The tunica albuginea is a firm, white, and very compact membrane of a glistening appearance, which immediate- ly invests the body of the testis and the epidi- dymus; serving in some measure to connect them to each other, but without extending itself at all to the spermatic chord. This tu- nica albuginea serves to confine the growth of the testis and epididymus within certain limits, and by giving them a due degree of firmness, enables them to perform their pro- per functions. * The cremaster muscle is composed of a few fibres from the obliquus internus abdominis, which uniting with a few from the transversalis, descend upon the spermatic chord, and are insen- sibly lost upon the tunica vaginalis of the testicle. It 9erves to suspend and draw up the testicle. Of the Abdomen. 289 Having removed this last tunic, we disco- ver the substance of the testicle itself, which appears to be made up of an infinite number of very elastic filaments, which may be best distinguished after macerating the testicle in water. Each testicle is made up of the sper- matic artery and vein, and the excretory ves- sels or tubuli seminiferi. There are likewise a great number of absorbent vessels, and some branches of nerves to be met with in the tes- ticles. The spermatic arteries arise one on each side from the aorta, generally about an inch below the emulgents. The right spermatic vein commonly passes into the vena cava; but the left spermatic vein usually empties itself into the emulgent on that side; and it is sup- posed to take this course into the emulgent, that it may avoid passing over the aorta, which it would be obliged to do in its way to the vena cava. The blood is circulated very slowly through the spermatic artery, which makes an infinite number of circumvolutions in the substance of the testicle, where it deposites the semen, which passes through the tubuli seminiferi. These tubuli seminiferi are seen running in short waves from the tunica albuginea to the axis of the testicle ; and are divided into dis- tinct portions by certain thin membranous pro- ductions, which originate from the tunica al- buginea. They at length unite, and by an infinite number of convolutions form a sort of Oo 29U Of the Abdomen. appendix to the testis called epididymis,* which is a vascular body of an oblong shape, situate upon the superior part of each testicle. These tubuli of the epididymis at length form an ex- cretory duct called vas deferens, which as- cends towards the abdominal rings, with the other parts that make up the spermatic chord, and then a separation takes place ; the nerves and blood-vessels passing on to their several terminations, and the vas deferens going to deposite its semen in the vesiculae seminales, which are two soft bodies of a white and con- voluted appearance externally, situated ob- liquely between the rectum and the lower part of the bladder, and uniting together at the lower extremity. From these reservoirs,f * The testicles were named didymi by the ancients, and the name of this part was given to it on account of its situation upon the testicle. ■f That the bags called vesicula seminales are reservoirs of se- men, is a circumstance which has been by anatomists universal- ly believed. Mr. J. Hunter, however, from several circumstan- ces, has been induced to think this opinion erroneous. He has examined these vesiculae in people who have died sud- denly, and he found their contents to be different in their proper- ties from the semen. In those who had lost one of the testicles, or the use of one of them, by disease, both the vesiculae were full, and their contents similar. And in a lusus nature, where there was no communication between the vasa deferentia and ve- siculae, nor between the vesiculae and penis, the same thing took place. From these observations, he thinks we have a presumptive proof, That the semen can be absorbed in the body of the testi- cle and in the epididymis, and that the vesiculae secrete a mucus which they are capable of absorbing when it cannot be made use of: That the semen is not retained in reservoirs after it is secret- ed, and kept there till it is used; but that it is secreted at the time, in consequence of certain affections of the mind stimulat- ing the testicles to this action. He corroborates his observations by the appearance on dissec- tion in other animals; and here he finds, That the shape and Of the Abdomen. 291 which are plentifully supplied with blood-ves- sels and nerves, the semen is occasionally dis- charged through two short passages, which open into the urethra close to a little eminence called verumontanum. contents of the vesiculae vary much in different animals, while the semen in most of them he has examined is nearly the same : That the vasa deferentia in many animals do not communicate with the vesiculae: That the contents of the vesiculae of castrated and perfect animals are similar, and nearly equal in quantity, in no way resembling the semen as emitted from the animal in coitu, or what is found in the vas deferens after death. He ob- serves likewise, that the bulb of the urethra of perfect males is considerably larger than in castrated animals. From the whole, he thinks the following inferences may be fairly drawn: That the bags called vesiculx seminales are not se- minal reservoirs, but glands secreting a peculiar mucus; and that the bulb of the urethra is properly speaking the receptacle of the semen, in which it is accumulated previous to ejection. But although he has endeavoured to prove that the vesiculae do not contain the semen, he has not been able to ascertain their particular use. He thinks, however, we may be allowed upon the whole to conclude, that they are, together with other parts, subservient to the purposes of generation. Although the author has treated this subject very ably, and made many ingenious observations, some things may be object- ed to what he has advanced ; of which the following are a few i That those animals who have bags called vesiculae seminales per- form copulation quickly ; whereas others that want them, as in the dog kind, are tedious in copulation: That in the human body, at least, there is a free communication between the vasa deferentia and vesiculae ; and in animals where the author has observed no communication between the vasa deferentia and ve- siculae, there may be a communication by vessels not yet disco- vered, and which may be compared to the hepato-cystk ducts in fowls and fishes : That the fluid in the end of the va$a defe- rentia and the vesiculae seminales are similar, according to the author's own observation.: That the vesiculae in some animals increase and decrease with the testicle at particular seasons : That in birds and certain fishes, there is a dilatation of the ends of the vasa deferentia, which the author himSelf allows to be a reservoir for the semen. With respect to the circumstance of the bulb of the urethra answering tire purpose of a reservoir, the author has mentioned no facts which tend to establish this opinion. See Observations on certain Parts of the Animal Occonomy. 292 Of the Abdomen. Near this eminence we meet with the pros- tate, which is situated at the neck of the blad- der, and is described as being of a glandular structure. It is shaped somewhat like a heart with its small end foremost, and invests the origin of the urethra. Internally it appears to be of a firm substance, and composed of seve- ral follicles, secreting a whitish viscid fluid, that is discharged by ten or twelve excretory ducts into the urethra, on each side of the openings of the vesiculae seminales at the same time, and from the same causes that the semen is expelled. As this latter fluid is found to be exceedingly limpid in the vesiculae seminales of the dead subject, it probably owes its white- ness and viscidity to this liquor of the pros- tate. The penis, which is to be considered as the vehicle or active organ of procreation, is composed of two columns, the corpora caver- nosa, and corpus spongiosum. The corpora cavernosa, which constitute the greatest part of the penis, may be described as two cylin- drical ligamentous tubes, each of which is composed of an infinite number of minute cells of a spongy texture, which communicate with each other. These two bodies are of a very pliant texture, and capable of considerable distention; and being united laterally to each other, occasion by this union a space above and another below. The uppermost of these spaces is filled by the blood-vessels, and the lower one, which is larger than the other, by the urethra and its corpus spongiosum. These two cavernous bodies are at first only separat- Of the Abdomen. 293 ed by a partition of tendinous fibres, which allow them to communicate with each other; but they afterwards divaricate from each other like the branches of the letter Y, and diminish- ing gradually in size, are attached, one on each side, by means of the ligamentum sus- pensorium penis to the ramus ischii, and to the inferior portion of the os pubis. The corpus spongiosum penis, or corpus spongiosum urethrae, as it is styled by some autnors, begins as soon as the urethra has passed the prostate, with a thick origin almost like a heart, first under the urethra, and af- terwards above it, becoming gradually thin- ner, and surrounding the whole canal of the urethra, till it terminates in a considerable ex- pansion, and constitutes what is called the glans penis, which is exceedingly vascular, and covered with papillae like the tongue. The cuticle which lines the inner surface of the ure- thra, is continued over the glans in the same manner as it is spread over the lips. The penis is invested by the common inte- guments, but the cutis is reflected back every where from the glans as it is in the eye-lids; so that it covers this part, when the penis is in a relaxed state, as it were with a hood, and from this use is called prepuce. The prepuce is tied down to the under part of the glans by a small ligament catted franum, which is in fact only a continuation of the cu- ticle and cutis. There are many simple seba- ceous follicles called glandula odorifera, placed round the basis of the glans ; and the fluid they secrete serves to preserve the exqui- 294 Of the Abdomen. site sensibility of this part of the penis, and to prevent the ill effects of attrition from the pre- puce. The urethra may be defined to be a mem- branous canal, passing from the bladder through the whole extent of the penis, Seve- ral very small openings, called lacuna, com- municate with this canal, through which a mu- cus is discharged into, it; and besides these, there are two glands, first described by Cow- per, as secreting a fluid for lubricating the urethra, and called Cowper's glatids ;* and Littref speaks of a gland situated near the prostate, as being destined for the same use. The urethra being continued from the neck of the bladder, is to be considered as making part of the urinary passage; and it likewise affords a conveyance to the semen, which we have observed is occasionally discharged into it from the vesiculae seminales. The direc- tion of this canal being first under and then before the pubis, occasions a winding in its course from the bladder to the penis not unlike the turns of the letter S. The penis has three pair of muscles, the erectores, acceleratores, and transversales. They push the blood from the crura to the fore part of the corpora cavernosa. The first origi- nate from the tuberosity of the ischium, and terminate in the corpora cavernosa. The ac- celeratores arise from the sphincter, and by * Both Hefster and Morgagni observe, that they have some- times not been able to find these glands: so that they do not seem to exist in all subjects. f Memoires de 1' Acad. Royale des Sciences, 1700. Of the Abdomen. 295 their insertion serve to compress the bulbous part of the urethra; and the transversales are destined to afford a passage to the semen, by dilating the canal of the urethra. The arteries of the penis are chiefly deriv- ed from the internal iliacs. Some of them are supposed to terminate by pabulous orifices within the corpora cavernosa and corpus spon- giosum ; and others terminate in veins, which at last make up the vena magna dorsi penis, and other smaller veins, which are in general distributed in like order with the arteries. Its nerves are large and numerous. They arise from the great sciatic nerve, and accom- pany the arteries in their course through the penis. We have now described the anatomy of this organ ; and there only remains to be explain- ed, how it is enabled to attain that degree of firmness and distention which is essential to the great work of generation. The greatest part of the penis has been spoken of as being of a spongy and cellular texture, plentifully supplied with blood-vessels and nerves, and as having muscles to move it in different directions. Now, the blood is constantly passing into its cells through the small branches of the arteries which open into them, and is from thence as constantly return- ed by the veins, so long as the corpora caver- nosa and corpus spongiosum continue to be in a relaxed and pliant state. But when, from any nervous influence, or other means, which it is not necessary here to define or explain, the erectores penis, ejaculatores seminis, le- 296 Of the Abdomen. vatores ani, he. are induced to contract, the veins undergo a certain degree of compres- sion, and the passage of the blood through them is so much impeded, that it collects in them in a greater proportion than they are enabled to carry off, so that the penis gradu- ally enlarges ; and being more and more for- cibly drawn up against the os pubis, the vena magna itself is at length compressed, and the penis becomes fully distended. But as the causes which first occasioned this distention subside, the penis gradually returns to its state of relaxation. J. 2. Female Organs of Generation. Anatomical writers visually divide the fe- male organs of generation into external and in- ternal. In the first division they include the mons veneris, labia pudendi, perinaum, clito- ris, nympha, and caruncula myrtiformes ; and in the latter, the vagina, with the uterus and its appendages. The mons veneris, which is placed on the upper part of the symphysis pubis, is inter- nally composed of adipose membranes, which makes it soft and prominent: it divides into two parts called labia pudendi, which descend- ing towards the rectum, from which they are divided by the perinaeum, form what is called the fourchette. The perinaeum is that fleshy space which extends about an inch and an half from the fourchette to the anus, and from thence about two inches to the coccyx. Of the Abdomen. 297 The labia pudendi being separated, we ob- serve a sulcus called fossa magna; in the up- per part of which is placed the clitoris, a small round spongy body, in some measure resem- bling the male penis, but impervious, com- posed of two corpora cavernosa, arising from the tuberosities of the ossa ischii; furnished with two pair of muscles, the erectores clito- ridis, and the sphincter or constrictor ostii va- ginae ; and terminating in a glans, which is covered with its prepuce. From the lower part, on each side of the fossa, pass the nym- phae, two membranous and spongy folds which seem destined for useful purposes in parturi- tion, by tending to enlarge the volume of the vagina as the child's head passes through it. Between these, about die middle of the fossa magna, we perceive the orifice of the vagina or os externum, closed by folds and wrinkles; and about half an inch above this, and about an inch below the clitoris, appears the meatus urinarius or orifice of the urethra, much short- er, though somewhat larger, than in men, with a little prominence at its lower edge, which facilitates the introduction of the ca- theter. The os externum is surrounded internally by several membranous folds called caruncula myrtiformes, which are partly the remains of a thin membrane called hymen, that covers the vagina in children. In general the hymen is sufficiently open to admit the passage of the menses, if it exists at the time of their appear- ance ; sometimes, however, it has been found perfectly closed. pP 298 Of the Abdomen. The vagina, situated between the urethra and the rectum, is a membranous cavity, sur- rounded especially at its external extremity with a spongy and vascular substance, which is covered by the sphincter ostii vaginx. It terminates in the uterus, about half an inch above the os tincae, and is wider and shorter in women who have had children than in vir- gins. All these parts are plentifully supplied with blood-vessels and nerves. Around the nym- phae there are sebaceous follicles, which pour out a fluid to lubricate the inner surface of the vagina; and the meatus urinarius, like the urethra in the male subject, is constantly moistened by a mucus, which defends it against the acrimony of the urine. The uterus is a hollow viscus, situated in the hypogastric region, between the rectum and bladder. It is destined to receive the first rudiments of the foetus, and to assist in the developement of all its parts, till it arrives at a state of perfection, and is fitted to enter into the world, at the time appointed by the wise Author of nature. The uterus, in its unimpregnated state, re- sembles a pear in shape, somewhat flattened, with its fundus or bottom part turned towards the abdomen, and its cervix or neck surround- ed by the vagina. The entrance into its ca- vity forms a little protuberance, which has been compared to the mouth of a tench, and is therefore called os tinea. The substance of the uterus, which is of a considerable thickness, appears to be com- Of the Abdomen. 299 posed of muscular and small ligamentous fi- bres, small branches of nerves, some lym- phatics, and with arteries and veins innumera- ble. Its nerves are chiefly derived from the intercostal, and its arteries and veins from the hypogastric and spermatic. The membrane which lines its cervix, is a continuation of the inner membrane of the vagina; but the outer surface of the body of the uterus is covered with the peritonaeum, which is reflected over it, and descends from thence to the intestinum rectum. This duplicature of the peritonaeum, by passing off from the sides of the uterus to the sides of the pelvis, is there firmly con- nected, and forms what are called ligamenta uteri lata; which not only serve to support the uterus, but to convey nerves and blood- vessels to it. The ligamenta uteri rotunda arise from the sides of the fundus uteri, and passing along within the fore-part of the ligamenta lata, de- scend through the abdominal rings, and ter- minate in the substance of the mons veneris. The substance of these ligaments is vascular, and although both they and the ligamenta lata admit the uterus in the virgin state, to move only about an inch up and down, yet in the course of pregnancy they admit of consider- able distention, and after parturition return nearly to their original state with surprising quickness. On each side of the inner surface of the uterus, in the angle near the fundus, a small orifice is to be discovered, which is the begin- ning of one of the tubae Fallopianse. Each of 300 Of the Abdomen. these tubes, which are two in number, passing through the substance of the uterus, is extend- ed along the broad ligaments, till it reaches the edge of the pelvis, from whence it reflects back; and turning over behind the ligaments, about an inch of its extremity is seen hanging loose in the pelvis, near the ovarium. These extremities, having a jagged appearance, are called fimbria, or morsus diaboli. Each tuba Fallopiana is usually about three or four inches long. Their cavities are at first very small, but become gradually larger, like a trumpet, as they approach the fimbriae. Near the fimbriae of each tuba Fallopiana, about an inch from the uterus, is situated an oval body called ovarium, of about half the size of the male testicle. Each of these ovaria is covered by a production of the peritonaeum, and hangs loose in the pelvis. They are of a flat and angular form, and appear to be com- posed of a white and cellular substance, in which we are able to discover several minute vesicles filled with a coagulable lymph, of an uncertain number, commonly exceeding 12 in each ovary. In the female of riper years, these vesicles become exceedingly turgid, and a kind of yellow coagulum is gradually formed within one of them, which increases for a cer- tain time. In conception, one of these mature ova is supposed to be impregnated with the male semen, and to be squeezed out of its nidus into the Fallopian tube ; after which the ruptured part forms a substance which in some animals is of a yellow colour, and is therefore called corpus luteum; and it is observable, that Of the Abdomen. 301 the number of these scars or fissures in the ovarium, constantly corresponds with the num- ber of foetuses excluded by the mother. q. 3. Of Conception. Man, being ever curious and inquisitive, has naturally been led to inquire after the origin of his existence ; and the subject of generation has employed the philosophical world in all ages : but in following nature up to her mi- nute recesses, the philosopher soon finds him- self bewildered, and his imagination often sup- plies that which he so eagerly wishes to disco- ver, but which is destined perhaps never to be revealed to him. Of the many theories which have been formed on this subject, that of the ancient philosophers seems to have been the most simple : they considered the male semen as alone capable of forming the foetus, and be- lieved that the female only afforded it a lodging in the womb, and supplied it with nourishment after it was perfectly formed. This opinion, however, soon gave place to another, in which the female was allowed a more considerable share in conception. This second system considered the foetus as being formed by the mixture of the semi- nal liquor of both sexes, by a certain arrange- ment of its several particles in the uterus. But in the 16th century, vesicles or eggs were discovered in the ovaria or female testicles ; the foetus had been found sometimes in the ab- domen, and sometimes in the Fallopian tubes ; 302 Of the Abdomen. and the two former opinions were exploded in favour of a new doctrine. The ovaria were compared to a bunch of grapes, being suppos- ed to consist of vesicles, each of which had a stalk ; so that it might be disengaged without hurting the rest, or spilling the liquor it con- tained. Each vesicle was said to include a lit- tle animal, almost complete in all its parts ; and the vapour of the male semen being conveyed to the ovarium, was supposed to produce a fer- mentation in the vesicle, which approached the nearest to maturity; and thus inducing it to disengage itself from the ovarium, it passed into the tuba Fallopiana, through which it was conveyed to the uterus. Here it was supposed to take root like a vegetable seed, and to form, with the vessels originating from the uterus, what is called the placenta; by means of which the circulation is carried on between the mo- ther and the foetus. This opinion, with all its absurdities, con- tinued to be almost universally adopted till the close of the same century, when Lieuwen- hoeck, by means of his glasses, discovered certain opake particles, which he described as so many animalcula, floating in the seminal fluid of the male. This discovery introduced a new schism among the philosophers of that time, and gave rise to a system which is not yet entirely explod- ed. According to this theory the male semen passing into the tubae Fallopianae, one of the animalcula penetrates into the substance of the ovarium, and enters into one of its vesicles or ova. This impregnated ovum is then squeez- Of the Abdomen. 303 ed from its husk, through the coats of the ova- rium, and being seized by the fimbriae, is con- ducted through the lube to the uterus, where it is nourished till it arrives at a state of per- fection. In this system there is much ingenu- ity ; but there are certain circumstances sup- posed to take place, which have been hitherto inexplicable. A celebrated modern writer, M. Buffon, endeavours to restore, in some mea- sure, the most ancient opinion, by allowing the female semen a share in this office ; asserting, that animalcula or organic particles are to be discovered in the seminal liquor of both sexes : he derives the female semen from the ovaria, and he contends that no ovum exists in those parts. But in this idea he is evidently mista- ken ; and the opinion now most generally adopted is, that an impregnation of the ovum, by the influence of the male semen, is essential to conception.* That the ovum is to be im- pregnated, there can be no doubt; but as the manner in which such an impregnation is sup- posed to take place, and the means by which the ovum afterwards gets into the Fallopian tube, and from thence into the uterus, are still found- ed chiefly on hypothesis, we will not attempt to extend farther the investigation of a sub- ject concerning which so little can be advanc- ed with certainty. * The learned Abbe Spallan/.ani has thrown much light on this curious subject, and has proved by a variety of experiments, that the animalcule exists entire in the female ovum and that the male seed is only necessary to vivify and pit it in motion.—His experiments and observations are worthy the attentive perusal of every physiologist. ,304 Of the Abdomen, \. 4. Of the Foetus in Utero. Opportunities of dissecting the human gravid uterus occurring but seldom, the state of the embryo* immediately after conception cannot be perfectly known. When the ovum descends into the uterus, it is supposed to be very minute ; and it is not till a considerable time after conception that the rudiments of the embryo begin to be ascertain- ed. About the third or fourth week the eye may discover the first lineaments of the foetus ; but these lineaments are as yet very imperfect, it being only about the size of a house-fly. Two little vessels appear in an almost transpa- rent jelly ; the largest of which is destined to become the head of the foetus, and the other smaller one is reserved for the trunk. But at this period no extremities are to be seen; the umbilical cord appears only as a very minute thread, and the placenta does not as yet absorb the red particles of the blood. At six weeks, not only the head but the features of the face begin to be developed. The nose appears like a small prominent line, and we are able to dis- cover another line under it, which is destined for the separation of the lips. Two black points appear in the place of eyes, and two minute holes mark the ears. At the sides of * The rudiments of the child are usually distinguished by this name till the human figure can be distinctly ascertained, and then it has the appellation of /ictus. Of the Abdomen. 305 the trunk, both above and below, we see four minute protuberances, which are the rudiments of the arms and legs. At the end of eight weeks the body of the foetus is upwards of an inch in length, and both the hands and feet are to be distinguished. The upper extremities are found to increase faster than the lower ones, and the separation of the fingers is accomplish- ed sooner than that of the toes. At this period the human form may be de- cisively ascertained ;—all the parts of the face may be distinguished, the shape of the body is clearly marked out, the haunches and the abdomen are elevated, the fingers and toes are separated from each other, and the intestines appear like minute threads. At the end of the third month, the foetus measures about three inches ; at the end of the fourth month, five inches ; in the fifth month, six or seven inches ; in the sixth month, eight or nine inches ; in the seventh month, eleven or twelve inches; in the eighth month, fourteen or fifteen inches ; and at the end of the ninth month, or full time, from eighteen to twenty- two inches. But as we have not an opportu- nity of examining the same foetus at different periods of pregnancy, and as their size and length may be influenced by the constitution and mode of life of the mother, calculations of this kind must be very uncertain. The foetus during all this time assumes an oval figure, which corresponds with the shape of the uterus. Its chin is found reclining on its breast with its knees drawn up towards its chin, and its arms folded over them. But it Qq 306 Of the Abdomen. seems likely, that the posture of some of these parts is varied in the latter months of pregnan- cy, so as to cause those painful twitches which its mother usually feels from time to time. In natural cases, its head is probably placed to- wards the os tincse from the time of conception to that of its birth; though formerly it was con- sidered as being placed towards the fundus ute- ri till about the eighth or ninth month, when the head, by becoming specifically heavier than the other parts of the body, was supposed to be turned downwards. The capacity of the uterus increases in pro- portion to the growth of the foetus, but with- out becoming thinner in its substance, as might naturally be expected. The nourishment of the foetus, during all this time, seems to be de- rived from the placenta, which appears to be originally formed by that part of the ovum which is next the fundus uteri. The remain- ing part of the ovum is covered by a mem- brane called spongy chorion \* within which is another called true chorion, which includes a third termed amnios ;f this contains a watery * Tit. Hunter has described this as a lamella from the inner surface of the uterus. In the latter months of pregnancy it be- comes gradually thinner and more connected with the chorion : he has named it membrana caduca, or decidua, as it is cast off with the placenta. Signior Scarpa, with more probability, considers it as being composed of an inspissated coagulable lymph. f In some quadrupeds, the urine appears to be conveyed from the bladder through a canal called urachus to the allantois, which is a reservoir, resembling a long and blind gut, situated between the chorion and amnios. The human foetus seems to have no such reservoir, though fome writers have supposed that it does exist. From the top of the bladder a few longitudinal fibres are extended to the umbilical chord ; and these fibres have been con- sidered as the urachus, though without having been ever found pervious. Of the Abdomen. 307 fluid which is the liquor amnii,* in which the foetus floats till the time of its birth. On the side next the foetus, the placenta is covered by the amnios and true chorion; on the side next the mother it has a production continued from the spongy chorion. The amnios and chorion are remarkably thin and transparent, having no blood-vessels entering into their composi- tion. The spongy chorion is opake and vas- cular. In the first months of pregnancy, the involu- cra bear a large proportion to their contents ; but this proportion is afterwards reversed, as the foetus increases in bulk. The placenta, which is the medium through which the blood is conveyed from the mother to the foetus, and the manner in which this conveyance takes place, deserve next to be considered. The placenta is a broad, flat, and spongy substance, like a cake, closely adhering to the inner surface of the womb, usually near the fundus, and appearing to be chiefly made up of the ramifications of the umbilical arteries and vein, and partly of the extremities of the uterine vessels. The arteries of the uterus discharge their contents into the substance of * The liquor amnii coagulates like the lymph. It has been supposed to pass into the oesophagus, and to afford nourishment to the foetus; but this does not feem probable. Children have come into the world without an oesophagus or any communica- tion between the stomach and the mouth ; but there has been no well attested instance of a child's having been born without a pla- centa ; and it does not seem likely, that any of the fluid can be absorbed through the pores of the skin, the skin in the foetus be- ing every where covered with a great quantity of mucus. 308 Of the Abdomen. this cake ; and the veins of the placenta, re- ceiving the blood either by a direct communi- cation of vessels, or by absorption, at length form the umbilical vein, which passes on to the sinus of the vena portse, and from thence to the vena cava, by means of the canalis veno- sus, a communication that is closed in the adult. But the circulation of the blood through the heart is not conducted in the foetus as in the adult: in the latter, the blood is carried from the right auricle of the heart through the pulmonary artery, and is returned to the left auricle by the pulmonary vein; but a dilatation of the lungs is essential to the passage of the blood through the pulmonary vessels, and this dilatation cannot take place till after the child is born and has respired. This deficiency, how- ever, is supplied in the foetus by the immedi- ate communication between the right and left auricle, through an oval opening, in the septum which divides the two auricles, called foramen ovale. The blood is likewise transmitted from the pulmonary artery to the aorta, by means of a duct called canalis arteriosus, which, like the canalis venosus, and foramen ovale, gra- dually closes after birth. The blood is returned again from the foetus through two arteries called the umbilical arte- ries, which arise from the iliacs. These two vessels taking a winding course with the vein, form with that, and the membranes by which they are surrounded, what is called the um- bilical chord. These arteries, after ramifying through the substance of the placenta, dis- charge their blood into the veins of the uterus; \\.\T <>\l Y X he, t v V^/O^ u + ^^ l'|..lr\\\ t^j lr ,/>«+, PM,,(. Of the Abdomen. 309 in the same manner as the uterine arteries dis- charged their blood into the branches of the umbilical vein. So that the blood is constant- ly passing in at one side of the placenta and out at the other; but in what particular man- ner it gets through the placenta is a point not yet determined. EXPLANATION of PLATES XXV. XXVI. and XXVII. Plate XXV. Fig. 1. Shows the Contents of the Thorax and Abdomen in situ. 1. Top of the trachea, or windpipe. 2 2, The internal jugular veins. 3 3, The subcla- vian veins. 4, The vena cava descendens. 5, The right auricle of the heart. 6, The right ventricle. 7, Part of the left ventricle. 8, The aorta descendens. 9, The pulmonary artery. 10, The right lung, part of which is cut off to show the great blood-vessels. 11, The left lung entire. 12 12, The anterior edge of the diaphragm. 13 13, The two great lobes of the liver. 14, The ligamentum rotundum. 15, The gall-bladder. 16, The stomach. 17 17, The jejunum and ilium. 18, The spleen. 310 Of the Abdomen. Fig. 2. Shows the Organs subservient of the Chylopoietic Viscera,—with those of Urine and Generation. 1 1, The under side of the two great lobes of the liver, a, Lobulus Spigelii. 2, The li- gamentum rotundum. 3, The gall-bladder. 4, The pancreas. 5, The spleen. 6 6, The kidneys. 7, The aorta descendens. 8, Ve- na cava ascendens. 9 9, The renal veins co- vering the arteries. 10, A probe under the spermatic vessels and a bit of the inferior me- senteric artery, and over the ureters. 1111, The ureters. 12 12, The iliac arteries and veins. 13, The rectum intestinum. 14, The bladder of urine. Fig. 3. Shows the Chylopoietic Viscera, and Organs subservient to them, taken out of the Body entire. A A, The under side of the two great lobes of the liver. B, Ligamentum rotundum. C, The gall-bladder. D, Ductus cysticus. E, Ductus hepaticus. F, Ductus communis cho- ledochus. G, Venaportarum. H, Arteria hep- atica. I I, The stomach. K K, Venae and arte- riae gastro-epiploicae, dextrae h sinistra. L L, Venae & arteriae coronariae ventriculi. M, The spleen. N N, Mesocolon, with its vessels. OOO, Intestinum colon. P, One of the li- gaments of the colon, which is a bundle of longitudinal muscular fibres. Q.QQ.Q, Jeju- Of the Abdomen. 511 num and ilium. R R, Sigmoid flexure of the colon with the ligament continued, and over S, The rectum intestinum. T T, Levatores ani. U, Sphincter ani. V, The place to which the prostate gland is connected. W, The anus. Fig. 4. Shows the Heart of a Foetus at the full time, with the Right Auricle cut open to show the Foramen Ovale, or passage be- tween both Auricles. a, The right ventricle, b, The left ventri- cle, c c, The outer side of the right auricle stretched out. d d, The posterior side, which forms the anterior side of the septum, e, The foramen ovale, with the membrane or valve which covers the left side, f, Vena cava in- ferior passing through g, A portion of the dia- phragm. Fig. 5. Shows the Heart and Large Vessels of a Foetus at the full time. a, The left ventricle, b, The right ventri- cle, c, A part of the right auricle. d, Left auricle, e e, The right branch of the pulmo- nary artery, f, Arteria pulmonalis. g g, The left branch of the pulmonary artery, with a number of its largest branches dissected from the lungs, h, The canalis arteriosus, i, The arch of the aorta, kk, The aorta descendens. 1, The left subclavian artery, m, The left ca- rotid artery, n, The right carotid artery, o, The right subclavian artery, p, The origin 312 Of the Abdomen, of the right carotid arid right subclavian arteries in one common trunk, q, The vena cava su- perior or descendens. r, The right common subclavian vein. s, The left common sub- clavian vein. N. B. All the parts described in this figure are to be found in the adult, except the cana- lis arteriosus. Plate XXVI. Fig. 1. Exhibits the more superficial Lympha- tic Vessels of the Lower Extremity. A, The spine of the os ilium. B, The os pubis. C, The iliac artery. D, The knee. E, E, F, Branches of the crural artery. G, The musculus gastrocnemius. IT, The tibia. I, The tendon of the musculus tibialis anticus. On the out-lines, a, A lymphatic vessel be- longing to the top of the foot, b, Its first di- vision into branches. c, c, c, Other divisions of the same lymphatic vessel. d, A small lymphatic gland. e, The lymphatic vessels which lie between the skin and the muscles of the thigh, f, f, Two lymphat^ glands at the upper part of the thigh below the groin, g, g, Other glands. h, A lymphatic vessel which passes by the side of those glands with- out communicating with them ; and, bending towards the inside of the groin at (i), opens into the lymphatic gland (k). 1,1, Lympha- tic glands in the groin, which are common to the lymphatic vessels of the genitals and those \ V Of the Abdomen, 313 of the lower extremity. m, n, A plexus of lymphatic vessels passing on the inside of the iliac artery. Fig. 2. Exhibits a Back View of the lower Ex- tremity, dissected so as to show the deeper- seated Lymphatic Vessels which accompany the Arteries. A, The os pubis. B, The tuberosity of the ischium. C, That part of the os ilium which was articulated with the os sacrum. D, The extremity of the iliac artery appearing above the groin. E, The knee. F F, The two cut surfaces of the triceps muscle, which was divided to show the lymphatic vessels that pass through its perforation along with the cru- ral artery. G, The edge of the musculus gra- cilis. H, The gastrocnemius and soleus, much shrunk by being dried, and by the soleus be- ing separated from the tibia to expose the ves- sels. I, The heel. K, The sole of the foot. L, The superficial lymphatic vessels passing over the knee, to get to the thigh. On the out-lines ; M, The posterior tibial artery, a, A lymphatic vessel accompanying the posterior tibial artery, b, The same vessel crossing the artery, c, A small lymphatic gland, through which this deep-seated lymphatic vessel passes. d, The lymphatic vessel passing under a small part of the soleus, which is left attached to the bone, the rest being removed. e, The lym- phatic vessel crossing the popliteal artery, f, g, h, Lymphatic glands in the ham, through R r 314 Of the Abdomen. which the lymphatic vessel passes, i, The lymphatic vessel passing with the crural ar- tery, through the perforation of the triceps muscle, k, The lymphatic vessel, after it has passed the perforation of the triceps, dividing into branches which embrace the artery (1). m, A lymphatic gland belonging to the deep- seated lymphatic vessel. At this place those vessels pass to the fore part of the groin, where they communicate with the superficial lym- phatic vessels, n, A part of the superficial lymphatic vessel appearing on the brim of the pelvis. Fig. 3. Exhibits the Trunk of the Human Subject, prepared to show the Lymphatic Vessels and the Ductus Thoracicus. A, The neck. B B, The two jugular veins. C, The vena cava superior. D D D I>, The subclavian veins. E, The beginning of the aorta, pulled to the left side by means of a li- gature, in order to show the thoracic duct be- hind it. F, The branches arising from the curvature of the aorta. G G, The two caro- tid arteries. H H, The first ribs. I I, The trachea. K K, The spine. L L, The vena azygos. M M, The descending aorta. N, Ttie coeliac artery, dividing into three branch- es. O, The superior mesenteric artery. P, The right crus diaphragmatis. Q Q, The two kidneys. R, The right emulgent artery. S S, The external iliac arteries, g d, The musculi psoae. T, The internal iliac artery. Of the Abdomen. 315 ^auricles : That the use of these and the other valves of the heart may be understood, it must be observed, that four large vessels pass out from the basis of the heart, viz. two arteries and two veins : and that each of these vessels is furnished with a thin membranous production, which is attached all round to the borders of their several orifices, from whence hanging loosely down they appear to be divid- ed into two or three distinct portions. But as * Authors differ about the course and distinctions of these fibres ; and it seems right to observe, that the structure of the heart being more compact than that of other muscles, its fibres are not easily separated. 340 Of the Thorax. their uses in the arteries and veins are differ- ent, so are they differently disposed. Those of the arteries are intended to give way to the passage of the blood into them from the ven- tricles, but to oppose its return: and, on the contrary, the valves of the veins are construct- ed so as to allow the blood only to pass into the heart. In consequence of these different uses, we find the valves of the pulmonary ar- tery and of the aorta attached to the orifices of those vessels, so as to have their concave surfaces turned towards the artery; and their convex surfaces, which mutually meet toge- ther, being placed towards the ventricle, only permit the blood to pass one way, which is into the arteries. There are usually three of these valves belonging to the pulmonary artery, and as many to the aorta ; and from their figure they are called valvula semilunares. The com- munication between the two great veins and the ventricles is by means of the two appen- dages or auricles into which the blood is dis- charged ; so that the other valves which may be said to belong to the veins, are placed in each ventricle, where the auricle opens into it. The valves in the right ventricle are usually three in number, and are named valvula tri- cuspides ; but in the left ventricle we common- ly observe only two, and these are the valvula mitrales. The membranes which form these valves in each cavity are attached so as to pro- ject somewhat forward; and both the tricus- pides and the mitrales are connected with the tendinous strings, which were described as arising from the fleshy columna. By the con- Of the Thorax. 341 traction of either ventricle, the blood is driv- en into the artery which communicates v.ith that ventricle; and these tendinous strings being gradually relaxed as the sides of the ca- vity are brought nearer to each other, the valves naturally close the opening into the au- ricle, and the blood necessarily directs its course into the then only open passage, which is into the artery; but after this contraction, the heart becomes relaxed, the tendinous strings are again stretched out, and, drawing the valves of the auricle downwards, the blood is poured by the veins into the ventricle, from whence, by another contraction, it is again thrown into the artery, as will be described hereafter. The right ventricle is not quite so long, though somewhat larger, than the left; but the latter has more substance than the other: and this seems to be, because it is in- tended to transmit the blood to the most dis- tant parts of the body, whereas the right ven- tricle distributes it only to the lungs. The heart receives its nerves from the par vagum and the intercostals. The arteries which serve for its nourishment are two in number, and arise from the aorta. They sur- round in some measure the basis of the heart, and from this course are called the coronary arteries. From these arteries the blood is re- turned by veins of the same name into the au- ricles, and even into the ventricles. The muscular bags called the auricles are situated at the basis of the heart, at the sides of each other; and, corresponding with the two ventricles, are like those two cavities dis- 342 Of the Thorax. tinguished into right and left. These sacs, which are interiorly unequal, have externally a jagged appendix; which, from its having been compared to the extremity of an ear, has given them their name of auricles. Sect. XI. Angiology, or a Description of the Blood-vessels. The heart has been described as contracting itself, and throwing the blood from its two ven- tricles into the pulmonary artery and the aor- ta, and then as relaxing itself and receiving a fresh supply from two large veins, which are the pulmonary vein and the vena cava. We will now point out the principal distributions of these vessels. The pulmonary artery arises from the right ventricle by a large trunk, which soon divides into two considerable branches, which pass to the right and left lobes of the lungs ; each of these branches is afterwards divided and sub- divided into an infinite number of branches and ramifications, which extend through the whole substance of the lungs ; and from these branch- es the blood is returned by the veins, which, contrary to the course of the arteries, begin by very minute canals, and gradually become larger, forming at length four large trunks called pulmonary veins, which terminate in the left auricle by one common opening, from whence the blood passes into the left ventricle. From this same ventricle arises the aorta or great artery, which at its' beginning is nearly Of the Thorax. 343 an inch in diameter; it soon sends off two branches, the coronaries, which go to be dis- tributed to the heart and its auricles. After this, at or about the third or fourth vertebra of the back, it makes a considerable curva- ture ; from this curvature * arise three arte- ries; one of Which soon divides into two branch- es. The first two are the left subclavian and the left carotid, and the third is a com- mon trunk to the right subclavian and right carotid ; though sometimes both the carotids arise distinctly from the aorta. The two carotids ascend within the subcla- vian, along the sides of the trachea; and when they have reached the larynx, divide into two principal branches, the internal and external carotid. The first of these runs a lit- tle way backwards in a bending direction ; and having reached the under part of the ear, pass- es through the canal into the os petrosum, and entering into the cavity of the cranium, is distributed to the brain and the membranes which envelope it, and likewise to the eye. The external carotid divides into several branches, which are distributed to the larynx, pharynx, and other parts of the neck; and to the jaws, lips, tongue, eyes, temples, and all the external parts of the head. * Anatomists usually call the upper part of this curvature aorta ascendens; and the other part of the artery to its division at the iliacs, aorta descendens : but they differ about the place where this distinction is to be introduced ; and it seems sufficiently to answer every purpose, to speak only of the aorta and its curva- ture. 344 Of the Thorax. Each subclavian is likewise divided into a great number of branches. It sends off the vertebral artery, which passes through the openings we see at the bottom of the transverse processes of the vertebrae of the neck, and in its course sends off many ramifications to the neighbouring parts. Some of its branches are distributed to the spinal marrow, and after a considerable inflection it enters into the crani- um, and is distributed to the brain. The sub- clavian likewise sends off branches to the mus- cles of the neck and scapula ; and the medias- tinum, thymus, pericardium, diaphragm, the breasts, and the muscles of the thorax, and even of the abdomen, derive branches from the subclavian, which are distinguished by different names, alluding to the parts to which they are distributed; as the mammary, the phrenic, the intercostal, he. But notwithstand- ing the great number of branches which have been described as arising from the subclavian, it is still a considerable artery when it reach- es the axilla, where it drops its former name, which alludes to its passage under the clavi- cle, and is called the axillary artery; from which a variety of branches are distributed to the muscles of the breast, scapula, and arm.— But its main trunk taking the name of brachi- alis, runs along on the inside of the arm near the os humeri, till it reaches the joint of the fore-arm, and then it divides into two branch- es. This division however is different in dif- ferent subjects ; for in some it takes place high- er up and in others lower down. When it happens to divide above the joint, it may be Of the Thorax, 345 considered as a happy disposition in case of an accident by bleeding; for supposing the artery to be unfortunately punctured by the lancet, and that the haemorrhage could only be stop- ped by making a ligature on the vessel, one branch would remain unhurt, through which the blood would pass uninterrupted to the fore- arm and hand. One of the two branches of the brachialis plunges down under the flexor muscles, and runs along the edge of the ulna; while the other is carried along the outer sur- face of the radius, and is easily felt at the wrist, where it is only covered by the common integuments. Both these branches commonly unite in the palm of the hand, and form an ar- terial arch from whence branches are detached to the fingers. The aorta, after having given off at its cur- vature the carotids and subclavians which con- vey blood to all the upper parts of the body, descends upon the bocjies of the vertebrae a lit- tle to the left, as far as the os sacrum, where it drops the name of aorta, and divides into two considerable branches. In this course, from its curvature to its bifurcation, it sends off several arteries in the following order: 1. One or two little arteries, first demonstrated by Ruysch as going to the bronchi, and called arteria bronchinales Ruyschii. 2. The arteriae oesophageae. These are commonly three or four in number. They arise from the fore- part of the aorta, and are distributed chiefly to the oesophagus. 3. The inferior intercostal arteries, which are distributed between the ribs in the same manner as the arteries of the X x 346 Of the Thorax. three or four superior ribs are, which are de- rived from the subclavian. These arteries send off branches to the medulla spinalis. 4, The diaphragmatic or inferior phrenic arteries, which go to the diaphragm, stomach, omentum, duodenum, pancreas, spleen, liver, and -gall- bladder. 5. The coeliac, which sends off the coronary-stomachic, the splenic, and the hepa- tic artery. 6. The superior mesenteric artery, which is distributed to the mesentery and small intestines, 7. The emulgents, which go to the kidneys. 8. The arteries, which are distri- buted to the glandulae renales. 9. The sper- matic. 10. The inferior mesenteric artery, which ramifies through the lower portion of the mesentery and the large intestines.—A branch of this artery which goes to the rectum is call* ed the internal hamorrhoidal. 11. The lum- bar arteries, and a very small branch called the sacra, which are distributed to the mus- cles of the loins and abdomen, and to the os sacrum and medulla spinalis. The trunk of the aorta, when it has reached the last vertebra lumborum, or the os sacrum, drops the name of aorta, and separates into two forked branches called the iliacs. Each of these soon divides into two branches; one of which is called the internal iliac, or hypogas- tric artery, and is distributed upon the contents of the pelvis and upon the muscles on its out- er side. One branch, called pudenda commu- nis, sends small ramifications to the end of the rectum under the name of hamorrhoidales externa, and is afterwards distributed upon the penis. The other branch, the external ili- Of the Thorax. 347 ac, after having given off the circumflex artery of the os ilium and the epigastric, which is distributed to the recti-muscles, passes out of the abdomen under Poupart's ligament, and takes the name of crural artery. It descends on the inner part of the thigh close to the os femoris, sending off branches to the muscles, and then sinking deeper in the hind part of the thigh, reaches the ham, where it takes the name of popliteal: after this it separates into two considerable branches; one of which is called the anterior tibial artery ; the other di- vides into two branches, and these arteries all go to be distributed to the leg and foot. The blood, which is thus distributed by the aorta to all parts of the body, is brought back by the veins^ which are supposed to be conti- nued from the ultimate branches of arteries ; and uniting together as they approach the heart, at length form the large trunks, the ve- na cava ascendens, and vena cava descendens. All the veins which bring back the blood from the upper extremities, and from the head and breast, pass into the vena cava descendens ; and those which return it from the lower parts of the body terminate in the vena cava ascen- dens .-, and these two eavas uniting together as they approach the heart, open by one common orifice into the left auricle. It does not here seem to be necessary to fol- low the different divisions of the veins as we did those of the arteries; and it will be suffi- cient to remark, that in general every artery is accompanied by its vein, and that both are dis- tinguished by the same name. But, like many 548 Of the Thorax. other general rules, this too has its excep- tions.* The veins for instance, which accom- pany the external and internal carotid, are not called the carotid veins, but the external and in- ternal jugular.—In the thorax, there is a vein distinguished by a proper name, and this is the azygos, or vena sine pari. This vein, which is a pretty considerable one, runs along by the right side of the vertebrae of the back, and is chiefly destined to receive the blood from the intercostals on that side, and from the lower half of those on the left side, and to convey it into the vena cava descendens. In the abdo- men we meet with a vein, which is still a more remarkable one, and this is the vena porta, which performs the office both of an artery and a vein. It is formed by a re-union of all the veins which come from the stomach, intestines, omentum, pancreas, and spleen, so as to com- pose one great trunk, which goes to ramify through the liver; and after having deposited the bile, its ramifications unite and bring back into the vena cava, not only the blood which the vena portae had carried into the liver, but likewise the blood from the hepatic artery. Every artery has a vein which corresponds with it; but the trunks and branches of the veins are more numerous than those of the arteries. —The reasons for this disposition are perhaps more difficult to be explained ; the blood in its course through the veins is much farther re- moved from the source and cause of its mo- * Tn the extremities, some of the deep-seated veins, and all the superficial ones, take a course different from that of the arteries. Of the Thorax. 349 tion, which are in the heart, than it was when in the arteries ; so that its course is consequent- ly less rapid, and enough of it could not pos- sibly be brought back to the heart in the mo- ment of its dilatation, to equal the quantity which is driven into the arteries from the two ventricles, at the time they contract; and the equilibrium which is so essential to the conti- nuance of life and health would consequently be destroyed, if the capacity of the veins did not exceed that of the arteries, in the same pro- portion that the rapidity of the blood's motion through the arteries exceeds that of its return through the veins. A large artery ramifying through the body, and continued to the minute branches of veins, which gradually unite together to form a large trunk, may be compared to two trees united to each other at their tops ; or rather as having their ramifications so disposed that the two trunks terminate inone common point; and if we farther suppose, that both these trunks and their branches are hollow, and that a fluid is inces- santly circulated through them, by entering in- to one of the trunks and returning through the other, we shall be enabled to conceive how the blood is circulated through the vessels of the human body. Every trunk of an artery, before it divides, is nearly cylindrical, or of equal diameter through its whole length, and so are all its branches when examined separately. But eve- ry trunk seems to contain less blood than the many branches do into which that trunk sepa- rates ; and each of these branches probably 350 Of the Thorax. contains less blood than the ramifications do into which it is subdivided : and it is the same with the veins; the volume of their several ramifications, when considered together, being found to exceed that of the great trunk which they form by their union. The return of the blood through the veins to the heart, is promoted by the action of the muscles, and the pulsation of the arteries. And this return is likewise greatly assisted by the valves which are to be met with in the veins, and which constitute one of the great distinctions between them and the arteries. These valves, which are supposed to be form- ed by the inner coat of the veins, permit the blood to flow from the extremities towards the heart, but oppose its return. They are most frequent in the smaller veins. As the column of blood increases, they seem to become less necessary; and therefore in the vena cava as- cendens, we meet with only one valve, which is near its origin. The arteries are composed of several tunics. Some writers enumerate five of these tunics; but perhaps we may more properly reckon only three, viz. the nervous, muscular, and cuticular coats. The veins are by some anatomists de- scribed as having the same number of coats as the arteries; but as they do not seem to be ir- ritable, we cannot with propriety suppose them to have a muscular tunic. We are aware of Dr. Verschuir's* experiments to prove that the jugular and some other veins possess a * De Arteriarum et Venarum vi irritabili, 4to. Of the Thorax. 351 certain degree of irritability; but it is cer- tain, that his experiments, repeated by others, have produced a different result; and even he himself allows, that sometimes he was unable to distinguish any such property in the veins. Both these series of vessels are nourished by still more minute arteries and veins, which are seen creeping over their coats, and ramifying through their whole substance, and are called vasa vasorum: they have likewise many mi- nute branches of nerves. The arteries are much stronger than the veins, and they seem to require this force to be enabled to resist the impetus with which the blood circulates through them, and to impel it on towards the veins. When the heart contracts, it impels the blood into the arteries, and sensibly distends them; and these vessels again contract, as the heart becomes relaxed to receive more blood from the auricles; so that the cause of the contrac- tion and dilatation of the arteries seems to be easy to be understood, being owing in part to their own contractile power, and in part to the action of the heart; but in the veins, the effects of this impulse not being so sensibly felt, and the vessels themselves having little or no con- tractile power, the blood seems to flow in a constant and equal stream: and this, together with its passing gradually from a small channel into a larger one, seems to be the reason why the veins have no pulsatory motion, except the large ones near the heart; and in these it seems to be occasioned by the motion of the dia- phragm, and by the regurgitation of the blood in the cavas. 3 52 Of the Thorax. Sect. XII. Of the Action of the Heart, Auricles, and Arteries. The heart, at the time it contracts, drives the blood from its ventricles into the arteries; and the arteries being thus filled and distended, are naturally inclined to contract the moment the heart begins to dilate, and ceases to supply them with blood. These alternate motions of contraction and dilatation of the heart and ar- teries, are distinguished by the names of systole and diastole. When the heart is in a state of contraction or systole, the arteries are at that instant distended with blood, and in their diastole ; and it is in this state we feel their pulsatory motion which we call the pulse. When the heart dilates, and the arteries contract, the blood is impelled onwards into the veins through which it is returned back to the heart. While the heart, however, is in its systole, the blood cannot pass from the veins into the ventricles, but is detained in the auricles, which are two reservoirs formed for this use, till the diastole, or dilatation of the heart, takes place; and then the distended auricles contract, and drive the blood into the ventricles ; so that the auricles have an alternate systole and diastole as well as the heart. Although both the ventricles of the heart contract at the same time, yet the blood pass- es from one to the other. In the same mo- ment, for instance, that the left ventricle drives the blood into the aorta, the right ven- tricle impels it into the pulmonary artery, Of the Thorax. 353 which is distributed through all the substance of the lungs. The blood is afterwards brought back into the left ventricle by the pulmonary vein, at the same time that the blood is returned by the cavas, into the right ventricle, from ail the other parts of the body. This seems to be the mode of action of the heart and its vessels ; but the cause of this action has, like all other intricate and interesting sub- jects, been differently explained. It seems to depend on the stimulus made on the different parts of the heart by the blood itself, which by its quantity and heat, or other properties,* is perhaps capable of first exciting that motion, which is afterwards continued through life, in- dependent of the will, by a regular return of blood to the auricles, in a quantity proportion- ed to that which is thrown into the arteries. The heart possesses the vis insita, or prin- ciple of irritability, in a much greater degree than any other muscle of the body. The pulse is quicker in young than in old subjects, be- cause the former are cat. par. more irritable than the latter. Upon the same principle we may explain, why the pulse is constantly quick- er in weak than in robust persons. Y y * Dr. Harvey long ago suggested, that the blood is possessed of a living principle ; and Mr. J. Hunter has lately endeavoured to revive this doctrine : in support of which he has adduced many ingenious arguments. The subject is a curious one, and deferves to be prosecuted as an inquiry which cannot but be interesting to physiologists. 354 Of the Thorax* Sect. XIII. Of the Circulation. After what has been observed of the structure and action of the heart and its auricles, and likewise of the arteries and veins, there seem to be but very few arguments required to demonstrate the circulation of the blood, which has long since been established as a medical truth.- This circulation may be defined to be a perpetual motion of the blood, in consequence of the action of the heart and arteries, which impel it through all the parts of the body, from whence it is brought back by the veins of the heart. A very satisfactory proof of this circulation, and a proof easy to be understood, may be de- duced from the different effects of pressure on an artery and a vein. If a ligature, for instance, is passed round an artery, the vessel swells considerably between the ligature and the heart; whereas if we tie up a vein, it only be- comes filled between the extremity and the ligature, and this is what we every day observe in bleeding. The ligature we pass round the arm on these occasions, compresses the super- ficial veins; and the return of the blood through them being impeded, they become dis- tended. When the ligature is too loose, the veins are not sufficiently compressed, and the blood continues its progress towards the heart; and, on the contrary, when it is made too tight, the arteries themselves become compressed ; Of the Thorax. 355 and the flow of the blood through them being impeded, the veins cannot be distended- Another phaenomenon, which effectually proves the circulation, is the loss of blood that every living animal sustains by opening only a single artery of a moderate size ; for it con- tinues to flow from the wounded vessel till the equilibrium is destroyed which is essential to life. This truth was not unknown to the an- cients ; and it seems strange that it did not lead, them to a knowledge of the circulation, as it sufficiently proves, that all the other vessels must communicate with that which is opened. Galen, who lived more than 1500 years ago, drew this conclusion from it; and if we far- ther observe, that he describes (after Erasis- tratus, who flourished about 450 years before him) the several valves of the heart, and de- termines their disposition and uses, it will ap- pear wonderful, that a period of near 2Q00 years should afterwards elapse before the true course of the blood was ascertained. This discovery, for which we are indebted to the immortal Harvey, has thrown new lights on physiology and the doctrine of diseases, and constitutes one of the most important periods of anatomical history. Sect. XIV. Of the Nature of the Blood. Blood, recently drawn from a vein into a bason, would seem to be an homogeneous fluid 356 Of the Thorax. of a red colour ;* but when suffered to rest, it Soon coagulaies, and divides into two parts, which are distinguished by the names of cras- samentum and serum. The crassamentum is the red coagulum, and the serum is the water in which it floats. Each of these may be again separated into two others; for the crassamen- tum, by being repeatedly washed in warm wa- ter, gives out all its red globules, and what remains appears to be composed of the coagu- lable lymph,f which is a gelatinous substance, capable of being hardened by fire till it becomes perfectly horny: and if we expose the serum to a certain degree of heat, part of it will be found to coagulate like the white of an egg, and there will remain a clear and limpid water, resembling urine both in its appearance and smell. The serum and crassamentum differ in their proportion in different constitutions; in a strong person, the crassamentum is in a great- er proportion to the serum than in a weak one ;$ and the same difference is found to take place in diseases.J * The blood, as it flows through the arteries, is observed to be more florid than it is in the veins ; and this redness is acquir- ed in its passage through the lungs. Vid. feet. vii. f It may not be improper to observe, that till of late the coa- gulable lymph has been confounded with the serum of the^ blood, which contains a substance that is likewise coagulable, though only when exposed to heat, or combined with certain chemical substances ; whereas the other coagulates spontaneously when exposed to the air or to rest. % Hewson's Experim. Enq. Part I. $ When the blood separates into strum and crassamentum, if the latter be covered with a crust of a whitish or buff colour, it has been usually considered as a certain proof of the blood's being in a state of too great viscidity. This appearance commonly taking place in inflammatory diseases, has long served to confirm the Of the Thorax. 357 Sect. XV. Of Nutrition. The variety of functions which we have de- scribed as being incessantly performed by the living body, and the continual circulation of the blood through it, must necessarily occasion a constant dissipation of the several parts which enter into its composition. In speaking of the insensible perspiration, we observed how much was incessantly passing off from the lungs and the surface of the skin. The discharge by urine is likewise every day considerable; and great part of the bile, saliva, he. are excluded by stool. But the solid, as well as the fluid parts of the body, require a constant renewal of nutritious particles. They are exposed to the attrition of the fluids which are circulated through them; and the contraction and relax- ation they repeat so many thousand times in every day, would necessarily occasion a disso- lution of the machine, if the renewal was not proportioned to the waste. It is easy to conceive how the chyle formed from the aliment is assimilated into the nature theory which ascribes the cause of inflammation to lentor and ob- structions. But from the late Mr. Hewson's experiments it ap- pears, that when the action of the arteries is increased, the blood, instead of being more viscid, is, on the contrary, more fluid than in the ordinary state, previous to inflammation ; and that in con- sequence of this, the coagulable lymph suffers the red globules, which are the heaviest part of the blood, to fall down to the bot- tom before it coagulates ; so that the crassamentum is divided into two parts: one of which is found to consist of the coagula- ble lymph alone (in this case termed the buff) ; and the other, partly of this and partly of the red globules. 358 Of the Thorax. of blood, and repairs the loss of the fluid parts of our body ; but how the solids are renewed, has never yet been satisfactorily explained. The nutritious parts of the blood are probably deposited by the arteries by exsudation through their pores into the tela celluloa; and as the solid parts of the body are in the embryo only a kind of jelly, which gradually acquires the degree of consistence they are found to have when the body arrives at a more advanced age; and these same parts which consist of bones, cartilages, ligaments, muscles, he. are sometimes reduced again by disease to a gela- tinous state; we may, with some degree of probability, consider the coagulable lymph as the source of nutrition. If the supply of nourishment exceeds the degree of waste, the body increases; and this happens in infancy and in youth : for at those periods, but more particularly the former one, the fluids bear a large proportion to the solids ; and the fibres being soft and yielding, are pro- portionably more capable of extension and in- crease. But when the supply of nutrition only equals the waste, we neither increase nor de- crease ; and we find this to be the case when the body has attained its full growth or acme : for the solids having then acquired a certain degree of firmness and rigidity, do not permit a farther increase of the body. But as we ap- proach to old age, rigidity begins to be in ex- cess, and the fluids * bear a much less propor- * As the fluids become less in proportion to the solids, their acrimony is found to increase ; and this may perhaps compen- sate for the want of fluidity in the blood by diminishing its co- hesion. Of the Thorax. 359 tion to the solids than before. The dissipation of the body is greater than the supply of nour- ishment ; many of the smaller vessels become gradually impervious ;* and the fibres losing their moisture and their elasticity, appear flac- cid and wrinkled. The lilies and the roses dis- appear, because the fluids by which they were produced can no longer reach the extremities of the capillary vessels of the skin. As these changes take place, the nervous power being proportionably weakened, the irritability and sensibility of the body, which were formerly so remarkable, are greatly diminished ; and in advanced life, the hearing, the eye-sight, and all the other senses, become gradually im- paired. Sect. XVI. Of the Glands and Secretions. The glands are commonly understood to be small, roundish, or oval bodies formed by the convolution of a great number of vessels, and destined to separate particular humours from the mass of blood. They are usually divided into two classes; but it seems more proper to distinguish three kinds of glands, viz. the mucous, conglobate, and conglomerate. The mucous glands, or follicles, as they are most commonly called, are small cylindrical * In infancy, the arteries are numerous and large in respect to the veins, and the lymphatic glands are larger than at any other time of life ; whereas, in old age, the capacity of the ve- nous system exceeds that of the arteries and the lymphatic sys- tem almost disappear:-. 360 Of the Thorax, tubes continued from the ends of arteries. In some parts of the body, as in the tonsils, for example, several of these follicles may be seen folded together in one common covering, and opening into one common sinus. These folli- cles are the vessels that secrete and pour out mucus in the mouth, oesophagus, stomach, in- testines, and other parts of the body. The conglobate glands are peculiar to the lymphatic system. Every lymphatic vein pass- es through a gland of this kind in its way to the thoracic duct. They are met with in dif- ferent parts of the body, particularly in the axilla, groin, and mesentery, and are either solitary or in distinct clusters. The conglomerate glands are of much great- er bulk than the conglobate, and seem to be an assemblage of many smaller glands. Of this kind are the liver, kidneys, he. Some of them, as the pancreas, parotids, he. have a granu- lated appearance. All these conglomerate glands are plentifully supplied with blood-ves- sels ; but their nerves are in general very mi- nute, and few in number. Each little granu- lated portion furnishes a small tube, which unites with other similar ducts, to form the common excretory duct of the gland. The principal glands, and the humours they secrete, have been already described in different parts of this work; and there only remains for us to examine the general structure of the glands, and to explain the mechanism of secretion. On the first of these subjects two different systems have been formed; each of which has had, and still continues to have, Of the Thorax. 361 its adherents. One of these systems was ad- vanced by Malpighi, who supposed that an ar- tery entering into a gland ramifies very mi- nutely through its whole substance ; and that its branches ultimately terminate in a vesicu- lar cavity or follicle, from whence the secreted fluid passes out through the excretory duct. This doctrine at first met with few opponents ; but the celebrated Ruysch, who first attempt- ed minute injections with wax, afterwards dis- puted the existence of these follicles, and as- serted, that every gland appears to be a con- tinued series of vessels, which after being re- peatedly convoluted in their course through its substance, at length terminate in the ex- cretory duct. Anatomists are still divided be- tween these two systems: that of Malpighi, however, seems to be the best founded. The mode of secretion has been explained in a variety of ways, and they are all perfectly hypothetical. In such an inquiry it is natural to ask, how one gland constantly separates a particular humour, while another gland se- cretes one of a very different nature from the blood? The bile, for instance, is separated by the liver, and the urine by the kidneys. Are these secretions to be imputed to any particu- lar dispositions in the fluids, or is their cause to be looked for in the solids ? It has been supposed, that every gland con- tains within itself a fermenting principle, by which it is enabled to change the nature of the blood it receives, and to endue it with a par- ticular property. So that, according to this system, the blood, as it circulates through the Zz 362 Of the Thorax. kidneys, becomes mixed with the fermenting principle of those glands, and a part of it is converted into urine ; and again, in the liver, in the salival and other glands, the bile, the saliva, and other juices, are generated from a similar cause. But it seems to be impossible for any liquor to be confined in a place expos- ed to the circulation, without being carried away by the torrent of blood, every part of which would be equally affected ; and this sys- tem of fermentation has long been rejected as vague and chimerical. But as the cause of se- cretion continued to be looked for in the fluids, the former system was succeeded by another, in which recourse was had to the analogy of the humours. It was observed, that if paper is moistened with water, and oil and water are afterwards poured upon it, that the water only will be permitted to pass through it; but that, on the other hand, if the paper has been pre- viously soaked in oil instead of water, the oil only, and not the water will be filtered through it. These observations led to a supposition, that every secretory organ is originally furnish- ed with a humour analogous to that which it is afterwards destined to separate from the blood; and that in consequence of this disposition, the secretory vessels of the liver, for instance, will only admit the bilious particles of the blood, while all the other humours will be excluded. This system is an ingenious one, but the dif- ficulties with which it abounds are unanswer- able ; for oil and water are immiscible ; where- as the blood, as it is circulated through the body, appears to be an homogeneous fluid. Of the Thorax. 363 Every oil will pass through a paper moistened only with one kind of oil; and wine, or spirits miAed with water, will easily be filtered through a paper previously soaked in water. Upon the same principle, all our humours, though differing in their other properties, yet agree- ing in that of being perfectly miscible with each other, will all easily pass through the same nitre.—But these are not all the objec- tions to this system. The humours which are supposed to be placed in the secretory vessels for the determination of similar particles of the blood, must be originally separated without any analogous fluid; and that which happens once, may as easily happen always. Again, it sometimes happens from a vicious disposi- tion, that humours are filtered through glands which are naturally not intended to afford them a passage, and when this once has hap- pened, it ought, according to this system, to be expected always to do so: whereas this is not the case; and we are, after all, naturally led to seek for the cause of secretions in the solids. It does not seem right to ascribe it to any particular figure of the secretory vessels ; because the soft texture of those parts does not permit them to preserve any constant shape, and our fluids seem to be capable of accommodating themselves to every kind of figure. Some have imputed it to the differ- ence of diameter in the orifices of the differ- ent secretory vessels. To this doctrine ob- jections have likewise been raised; and it has been argued, that the vessels of the liver, for instance, would, upon this principle, aiiord a 364 Of the Thorax. passage not only to the bile, but to all the other humours ofless consistence with it. In reply to this objection, it has been supposed, that secondary vessels exist, which originate from the first, and permit all the humours thinner than the bile to pass through them. Each of these hypotheses is probably very remote from the truth. EXPLANATION of PLATE XXVIII. This plate represents the Heart in situ, all the large Arteries and Veins, with some of the Muscles, he. Muscles, he.—Superior Extremity.—a, Masseter. b, Complexus. c, Digastricus. d, Os hyoides. e, Thyroid gland, f, Levator scapulae, g, Cucullaris. h h," The clavicles cut. i, The deltoid muscle, k. Biceps flexor cubiti cut. . 1, Caraco-brachialis. m, Triceps extensor cubiti. n, The heads of the pronator teres, flexor carpi radialis, and flexor digito- rum sublimis, cut. o, The flexor carpi ulna- ris, cut at its extremity, p, Flexor digitorum profundus, q, Supinator radii longus, cut at its extremity, r, Ligamentum carpi transver- sale. s, Extensores carpi radiales. t, Latis- simus dorsi. u, Anterior edge of the serratus anticus major, v v, The inferior part of the diaphragm, w w, Its anterior edge cut. x x, The kidneys, y, Transversus abdominis, z, Os ilium. / Of the Thorax. 365 Inferior Extremity.—a, Psoas magnus, b, Iliacus internus. c, The fleshy origin of the tensor vaginae femoris. d d, The ossa pubis cut from each other, e, Musculus pectineus cut from its origin. /, Short head of the tri- ceps abductor femoris cut. g, The great head of the triceps, h, The long head cut. i, Vas^ tus internus. k, Vastus externus. /, Crure- us. m, Gemellus, n, Soleus. o, Tibia. py Peronaeus longus. q, Peronaeus brevis. r, Fibula. Heart and Blood-vessels.—A, The heart, with the coronary arteries and veins. B, The right auricle of the heart. C, The aorta ascen- dens. D, The left subclavian artery. E, The left carotid artery. F, The common trunk which sends off the right subclavian and right carotid arteries. G, The carotis externa. H, Arteria facialis, which sends off the coronary arteries of the lips. I, Arteria temporalis pro- funda. K, Aorta descendens. L L, The iliac arteries,—which send off M M, The fe- moral or crural arteries. N. B. The other arteries in this figure have the same distri- bution as the veins of the same name:— And generally, in the anatomical plates, the description to be found on the one side, points out the same parts in the other. 1, The fron- tal vein. 2, The facial vein. 3, Vena tem- poralis profunda. 4, Vena occipitalis. 5, Vena jugularis externa. 6, Vena jugularis interna, covering the arteria carotis commu- nis. 7. The vascular arch on the palm of the hand, which is formed by, 8, The radial ar- tery and vein, and, 9, The ulnar artery and 366 Of the Brain and Nerves. vein. 10 10, Cephalic vein. 11, Basilic vein, that on the right side cut. 12, Median vein. 13, The humeral vein, which, with the me- dian, covers the humeral artery. 14 14, The external thoracic or mammary arteries and veins. 15, The axillary vein, covering the artery. 16 16, The subclavian veins, which, with (6 6) the jugulars, form, 17, The vena cava superior. 18, The cutaneous arch of veins on the fore part of the foot. 19, The vena tibialis antica, covering the artery. 20, The vena profunda femoris, covering the ar- tery, 21, The upper part of the vena saphena major. 22, The femoral vein. 23 23, The iliac veins. 24 24, Vena cava inferior. 25 25, The renal veins covering the arteries. 26 26, The diaphragmatic veins. PART V. OF THE BRAIN AND NERVES. Sect. I. Of the Brain and its Integuments-. THE bones of the cranium were described in the osteological part of this work, as inclosing the brain, and defending it from ex- ternal injury: but they are not its only pro- tection ; for when we make an horizontal sec- tion through these bones, we find this mass Of the Brain and Nerves, 367 every wheyre surrounded by two membranes,* the dura and pia mater.—The first of these lines the interior surface of the cranium, to which it every where adheres strongly,! but more particularly at the sutures, and at the many foramina through which vessels pass between it and the pericranium. The dura mater t is perfectly smooth and inelastic, and its inner surface is constantly bedewed with a fine pellucid fluid, which every where sepa- rates it from the pia mater. The dura mater sends off several considerable processes, which divide the brain into separate portions, and prevent them from compressing each other. Of these processes there is one superior and longitudinal, called the falx or falciform pro- cess, from its resemblance to a scythe. It arises from the spine of the os frontis, near the crista galli, and extending along in the direction of the sagittal suture, to beyond the lambdoidal suture, divides the brain into two hemispheres. A little below the lambdoidal * The Greeks called these membranes meninges ; but the Ara- bians, supposing them to be the source of all the other mem- branes of the body, afterwards gave them the names of dura and pia mater ; by which they are now usually distinguished. •j- In young subjects this adhesion is greater than in adults; but even then, in the healthy subject, it is no where easily sepa- rable, without breaking through some of the minute vessels by means of which it is attached to the bone. £ This membrane is commonly described as consisting of two laminae ; of which the external one is supposed to perform the of- fice of periosteum internum to the cranium, while the internal one forms the folds and processes of the dura mater. In the na- tural state, however, no such separation is apparent ; like other membranes, we may indeed divide it, not into two only, but many laminae; but this division is artificial, and depends on the dexterity of the anatomist. 568 Of the Brain and Nerves. suture, it divides into two broad wings or ex- pansions called the transverse or lateral process- es, which prevents the lobes of the cerebrum from pressing on the cerebellum. Besides these there is a fourth, which is situated under the transverse processes, and being continued to the spine of the occiput, divides the cere- bellum into two lobes. The blood, after being distributed through the cavity of the cranium by means of the ar- teries, is returned, as in the other parts of the body, by veins which all pass on to certain channels, situated behind these several pro- cesses. These canals or sinuses communicate with each other, and empty themselves into the in- ternal jugular veins, which convey the blood into the vena cava. They are in fact triangu- lar veins, running through the substance of the dura mater, and, like the processes, are dis- tinguished into longitudinal and lateral; and where these three meet, and where the fourth process passes off, we observe a fourth sinus, •which is called torcular; Herophilus, who first described it, having supposed that the blood at the union of these two veins, is, as it were, in a press. Besides these four canals, which were known to the ancients, modern anatomists enu- merate many others, by giving the appellation of sinuses to other veins of the dura mater, which for the most part empty themselves into some of those we have just now described. There are the inferior longitudinal sinus, the superior a#d inferior petrous sinuses, the ca- Of the Brain and Nerves. 369 vernous sinuses, the circular sinus, and the anterior and posterior occipital sinuses. These sinuses or veins, by being conveyed through a thick dense membrane, firmly sus- pended, as the dura mater is, within the cra- nium, are less liable to rupture ; at the same time they are well supported, and by running every where along the inner surface of the bones, they are prevented from pressing on the substance of the brain. To prevent too great a dilatation of them, we find filaments (called chorda Willisii, from their having been first noticed by Willis) stretched across their cavi- ties ; and the oblique manner in which the veins from the brain run through the substance of the brain into these channels, serves the purpose of a valve, which prevents the blood from turning back into the smaller and weaker vessels of the brain. The pia mater is a much softer and finer membrane than the dura mater; being exceed- ingly delicate, transparent, and vascular. It in- vests every part of the brain, and sends off an infinite number of elongations, which insinu- ate themselves between the convolutions, and even into the substance of the brain. This membrane is composed of two laminae ; of which the exterior one is named tunica arachnoidea, from its thinness, which is equal to that of a spider's web. These two laminae are intimate- ly adherent to each other at the upper part of the brain, but are easily separable at the basis of the brain, and through the whole length of the medulla spinalis. The external layer, or tunica arachnoidea, appears to be spread uni- 3 A 370 Of the Brain and Nerves. formly over the surface of the brain, but with- out entering into its furrows as the inner layer does ; the latter being found to insinuate itself between the convolutions, and even into the interior cavities of the brain. The blood-ves- sels of the brain are distributed through it in their way to that organ, and are therefore di- vided into very minute ramifications, before they penetrate the substance of the brain. There are several parts included under the general denomination of brain. One of these, which is of the softest consistence, and fills the greatest part of the cavity of the cranium, is the cerebrum, or brain properly so called. Another portion, which is seated in the infe- rior and posterior part of the head, is the cere- bellum ; and a third, which derives its origin from both these, is the medulla oblongata. The cerebrum is a medullary mass of a mo- derate consistence, filling up exactly all the upper part of the cavity of the cranium, and divided into two hemispheres by the falx of the dura mater. Each of these hemispheres is usually distinguished into an interior, a mid- dle, and a posterior lobe. The first of these is lodged on the orbital processes of the os fron- tis : the middle lobes lie on the middle fossae of the basis of the cranium, and the posterior lobes are placed on the transverse septum of the os occipitis, immediately over the cerebel- lum, from which they are separated by the la- teral processes of the dura mater. These two portions afford no distinguishing mark of se- paration ; and on this account Haller, and many other modern anatomists, omit the distinction Of the Brain and Nerves. 371 ei middle lobe, and speak only of the anterior and posterior lobes of the bra:n. The cerebrum appears to be composed of two distinct substances. Of these, the exte- rior one, which is of a greyish or ash-colour, is called the cortex, and is somewhat softer than the other, which is very white, and is call- ed medulla or substantia alba. After having removed the falx, and separat- ed the two hemispheres from each other, we perceive a white convex body, the corpus cal- losum, which is a portion of the medullary substance, uniting the two hemispheres to each other, and not invested by the cortex. By making an horizontal incision in the brain, on a level with this corpus callosum, we dis- cover two oblong cavities, named the anterior or lateral ventricles, one in each hemisphere. These two ventricles, which communicate with each other by a hole immediately under the plexus choroides, are separated laterally by a very fine medullary partition, called septum lucidum, from its thinness and transparency. The lower edge of this septum is fixed to the fornix, which is a kind of medullary arch (as its name implies) situated under the corpus callosum, and nearly of a triangular shape. Anteriorly the fornix sends off two medullary chords, called its anterior crura; which seem to be united to each other by a portion of me- dullary substance, named commissiira anterior cerebri. These crura diverging from one ano- ther, are lost at the outer side of the lower and fore-part of the third ventricle. Posteri- orly the fornix is formed into two other crura, 372 Of the Brain and Nerves. which unite with two medullary protuberances called pedes hippocampi, and sometimes cornua ammonis, that extend along the back part of the lateral ventricles. The concave edge of the pedes hippocampi is covered by a medul- lary lamina, called corpus fimbriatum. Neither the edges of the fornix, nor its pos- terior crura, can be well distinguished, till we have removed the plexus choroides. This is a production of the pia mater, which is spread over the lateral ventricles. Its loose edges are collected, so as to appear like a vascular band on each side. When we have removed this plexus, we dis- cover several other protuberances included in the lateral ventricles. These are the corpora striata, the thalami nervorum opticorum, the tubercula quadrumgemina, and the pineal gland. The corpora striata are two curved oblong eminences, that extend along the anterior part of the lateral ventricles. They derive their name from their striated appearance, which is owing to an intermixture of the cortical and medullary substances of the brain. The thala- mi nervorum opticorum, are so called, because the optic nerves arise chiefly from them, and they are likewise composed both of the cortex and medulla. They are separated from the corpora striata only by a kind of medullary chord, the geminum centrum semi-circulare. The thalami are nearly of an oval shape, and are situated at the bottom of the upper cavity of the lateral ventricles. They are closely Of the Brain and Nerves. 378 united, and at their convex part seem to be- come one body. Anteriorly, in the space between the thala- mi, we observe an orifice by which the lateral ventricles communicate, and another leads down from this, under the different appella- tions oi foramen commune anterius, vulva iter ad infundibulum, but more properly iter ad ter- tium ventriculum; and the separation of the thalami from each other posteriorly, forms another opening or interstice called anus. This has been supposed to communicate with the third ventricle; but it does not, the bottom of it being shut up by the pia mater. The back part of the anus is formed by a kind of medul- lary band, which connects the thalami to each other, and is called commissiira posterior ce- rebri. Behind the thalami and commissura poste- rior, we observe a small, soft, greyish, and oval body, about the size of a pea. This is the glandula pinealis ; it is described by Galen under the name of conarion, and has been ren- dered famous by Descartes, who supposed it to be the seat of the soul. Galen seems for- merly to have entertained the same opinion. Some modern writers have, with as little rea- son, imagined that the soul is placed in the corpus callosum. The pineal gland rests upon four remarka- ble eminences, disposed in pairs, and seated immediately below it. These tubercles, which by the ancients were called testes and nates, have, since the time of Winslow, been more commonly named tubercula quadrugemina. 374 Of the Brain and Nerves. Under the thalami we observe another ca- vity, the third ventricle, which terminates ai> tenorly in a small medullary canal, the infun- dibulum, that leads to the glandula pituitaria. It has been doubted, whether the iufundi- bulum is really hollow ;> but some late experi- ments on this part of the brain * by Professor Murray of Upsal, clearly prove it to be a me- dullary canal, surrounded by both laminae of the pia mater. After freezing the brain, this channel was found filled with ice; and de Haen tells f us, he found it dilated, and filled with a calcareous matter.! The soft spongy body in which the infundi- bulum terminates, was by the ancients sup- posed to be of a glandular structure, and des- tined to filter the serosity of the brain. Spi- gelius pretended to have discovered its excre- tory duct, but it seems certain that no such duct exists. It is of an oblong shape, com- posed, as it were, of two lobes. In ruminant animals it is much larger than in man. From the posterior part of the third ventri- cle, we see a small groove or channel, de- scending obliquely backwards. This channel, which is called the aqueduct of Sylvius, though it was known to the ancients, opens into ano- ther cavity of the brain, placed between the cerebellum and medulla oblongata, and called the fourth ventricle. * Disp. de Infundibulo Cerebri. f Ratio Med. torn. vi. p. 27*1. j: The under part of it, however, appears to be impervious; at least no injection that can be depended on has been made to pass from it into the glandula pituitaria without laceration of parts. Of the Brain and Nerves, 375 The cerebellum, which is divided into two lobes, is commonly supposed to be of a firmer texture than the cerebrum; but the truth is, that in the greater number of subjects, there appears to be no sensible difference in the con- sistence of these two parts. It has more of the cortical than of the medullary substance in its composition. The furrow that divides the two lobes of the cerebellum leads anteriorly to a process, composed of medullary and cortical substan- ces, covered by the pia mater \ and which, from its being divided into numerous furrows? resembling the rings of the earth-worm, is named processus vermiformis. This process forms a kind of ring in its course between the lobes. The surface of the cerebellum does not af- ford those circumvolutions which appear in the cerebrum; but instead of these, we observe a great number of minute furrows, running pa- rallel to each other, and nearly in a transverse direction. The pia mater insinuates itself into these furrows. When we cut into the substance of the cere- bellum, from above downwards, we find the medullary part running in a kind of ramifying course, and exhibiting an appearance that has gotten the name of arbor vita. These ramifi- cations unite to form a medullary trunk; the middle, anterior, and most considerable part of which forms two processes, the crura cere- belli, which unite with the crura cerebri, to form the medulla oblongata. The last furnish- es two other processes, which lose themselves 376 Of tiie Brain and Nerves. under the nates, and thus unite the lobes of the cerebellum to the posterior part of the ce- rebrum. Under the nates we observe a trans- verse medullary line, or linea alba, running from one of these processes to the other; and between them we find a very thin medullary lamina, covered with the pia mater, which the generality of anatomists have (though seemingly without reason) considered as a valve formed for closing the communication between the fourth ventricle and the aquaeduc- tus Sylvii. Vieussens named it valvula ma- jor cerebri. The medulla oblongata is situated in the mid- dle, lower, and posterior part of the cranium, and may be considered as a production or con- tinuation of the whole medullary substance of the cerebrum and cerebellum, being formed by the union of two considerable medullary processes of the cerebrum, called crura cerebri, with two other smaller ones from the cerebel- lum, which were just now spoken of under the name of crura cerebelli. The crura cerebri arise from the middle and lower part of each hemisphere. They are se- parated from each other at their origin, but are united below, where they terminate in a mid- dle protuberance, the pons Varolii, so called, because Varolius compared it to a bridge. This name, however, can convey no idea of its real appearance. It is, in fact, nothing more than a medullary protuberance, nearly of a se- mi-spherical shape, which unites the crura ce- rebri to those of the cerebellum. Of the Brain and Nerves. 377 Between the crura cerebri, and near the an- terior edge of the pons Varolii, are two tuber- cles, composed externally of medullary, and internally of cineritious, substance, to which Eustachius first gave the name of eminentia mamillares. Along the middle of the posterior surface of the medulla oblongata, where it forms the anterior part of the fourth ventricle, we observe a kind of furrow which runs downwards and terminates in a point. About an inch above the lower extremity of this fissure, several me- dullary filaments are to be seen running to- wards it on each side in an oblique direction, so as to give it the appearance of a writing- pen ; hence it is called calamus scriptorius. From the posterior part of the pons Varolii, the medulla oblongata descends obliquely back- wards ; at its fore-part, immediately behind the pons Varolii, we observe two pair of emi- nences, which were described by Eustachi- us, but received no particular appellation till the time of Vieussens, who gave them the names of corpora olivaria and corpora pyramir dalia. The former are the outermost, being placed one on each side. They are nearly of an oval shape, and are composed of medulla, with streaks of cortical substance. Between these are the corpora pyramidalia, each of which terminates in a point. In the human subject these four eminences are sometimes not easily distinguished. The medulla spinalis or spinal marrow, which is the name given to the medullary chord that is extended down the vertebral ca- 3 B 378 Of the Brain and Nerves, nal, from the great foramen of the occipital bone to the bottom of the last lumbar vertebra, is a continuation of the medulla oblongata. Like the other parts of the brain, it is invested by the dura and pia mater. The first of these, in its passage out of the cranium, adheres to the foramen of the os occipitis. Its connection with the ligamentary substance that lines the cavity of the spine, is only by means of cellular mem- brane ; but between the several vertebrae, where the nerves pass out of the spine, it sends off prolongations, which adhere strongly to the vertebral ligaments. Here, as in the cranium, the dura mater has its sinuses or large veins. These are two in number, and are seen run- ning on each side of the medullary column, from the foramen magnum of the os occipitis to the lower part of the os sacrum. They communicate together by ramifying branches at each vertebra, and terminate in the verte- bral, intercostal, and sacral veins. The pia mater is connected with the dura mater by means of a thin transparent sub- stance which from its indentations between the spinal nerves has obtained the name of ligamentum denticulatum. It is somewhat firm- er than the tunica arachnoidea, but in other re- spects resembles that membrane. Its use is to support the spinal marrow, that it may not affect the medulla oblongata by its weight. The spinal marrow itself is externally of a white colour; but upon cutting into it we find its middle-part composed of a darker coloured mass, resembling the cortex of the brain. When the marrow has reached the first lumbar v^er- Of the Brain and Nerves. 379 tebra, it becomes extremely narrow, and at length terminates in an oblong protuberance ; from the extremity of which the pia mater sends off a prolongation or ligament, resem- bling a nerve, that perforates the dura mater, and is fixed to the os coccygis. The medulla spinalis gives rise to 30 or 31 pair of nerves, but they are not all of the same size, nor do they all run in the same direc- tion. The upper ones are thinner than the rest, and are placed almost transversely: as we descend we find them running more and more obliquely downwards, till at length their course is almost perpendicular, so that the low- ermost nerves exhibit an appearance that is called cauda equina, from its resemblance to a horse's tail. The arteries that ramify through the differ- ent parts of the brain, are derived from the internal carotid and from the vertebral arteries. The medulla spinalis is supplied by the ante- rior and posterior spinal arteries, and likewise receives branches, from the cervical, the infe- rior and superior intercostal, the lumbar, and the sacral arteries. Sect. II. Of the Nerves. The nerves are medullary chords, differing from each other in size, colour, and consist- ence, and deriving their origin from the me- dulla oblongata and medulla spinalis. There are 39, and sometimes 40, pair of these nerves; 380 Of the Brain and Nerves. nine* of which originate from the medulla oblongata, and 30 or 31 from the medulla spi- nalis. They appear to be perfectly inelastic, and likewise to possess no irritability. If we irritate muscular fibres, they immediately con- tract ; but nothing of this sort happens if we irritate a nerve. They carry with them a co- vering from the pia mater; but derive no tu- nic from the dura mater, as hath been gene- rally, though erroneously, supposed, ever since the time of Galen,f the outer covering of the nerves being in fact nothing more than the cel- lular membrane. This covering is very thick where the nerve is exposed to the action of muscles ; but where it runs through a bony ca- nal, or is secure from pressure, the cellular tunic is extremely thin, or altogether wanting. We have instances of this in the portio mollis of the auditory nerve, and in the nerves of the heart. By elevating, carefully and gently, the brain from the basis of the cranium, we find the first nine pair arising in the following order: l. The nervi olfactorii, distributed through the pituitary membrane, which constitutes the or- gan of smell. 2. The optici, which go to the * It has been usual to describe the ten pair of nerves as aris- ing from the medulla oblongata ; but as the tenth pair arise in the fame manner as the other spinal nerves, Santorini, Heister, Haller, and others, seem very properly to have classed them among the nerves of the spine. -j- Baron Haller and Professor Zinn seem to have been the first who demonstrated, that the dura mater is reflected upon and adheres to the periosteum at the edges of the foramina that afford a passage to the nerves out of the cranium, and vertebral eanal, or is soon lost in the cellular substance. Of the Brain and Nerves. 381 eyes, where they receive the impressions of visible objects. 3. The oculorum motores, so called because they are distributed to the mus- cles of the eye. 4. The pathetici, distributed to the superior oblique muscles of the eye, the motion of which is expressive of certain passions of the soul. 5. The nerves of this pair soon divide into three principal branches, and each of these has a different name. Its upper di- vision is the ophthalmicus, which is distribut- ed to various parts of the eyes, eye-lids, fore- head, nose, and integuments of the face. The second is called the maxillaris superior, and the third maxillaris inferior ; both which names allude to their distribution. 6. The abducto- res ; each of these nerves is distributed to the abductor muscle of the eye, so called, because it helps to draw the globe of the eye from the nose. 7. The auditorii,* which are distribut- ed through the organs of hearing. 8. The par vagum, which derives its name from the great number of parts to which it gives branch- es both in the thorax and abdomen. 9. The linguales, or hypo-glossi, which are distribut- ed to the tongue, and appear to contribute both to the organ of taste and to the motions of the tongue.f * This pair, soon after its entrance into the meatus auditorius internus, separates into two branches. One of these is of a very soft and pulpy consistence, it is called the portio mollis of the se- venth pair, and is spread over the inner part of the ear. The other passes out through the aqueduct of Fallopius in a firm chord, which is distinguished as the portio dura, and is distribut- ed to the external ear and other p.irts of the neck and face. •j Heister has summed up the ufes of these nine pair of nerves in the two following Latin verses: 382 Of the Brain arid Nerves. It has already been observed, that the spi- nal marrow sends off 30 or 31 pair of nerves ; these are chiefly distributed to the exterior parts of the trunk and to the extremities. They are commonly distinguished into the cervical, dorsal, lumbar, and sacral nerves. The cervical, which pass out from between the several vertebrae of the neck are eight* in number; the dorsal, twelve ; the lumbar, five; and the sacral, five or six ; the number of the latter depending on the number of holes in the os sacrum. Each spinal nerve at its origin is composed of two fasciculi of medullary fibres. One of these fasciculi arises from the anterior, and the other from the posterior, surface of the medulla. These fasciculi are separated by the ligamen- tum denticulatum; after which we find them contiguous to one another. They then perfo- rate the dura mater, and unite to form a con- siderable knot or ganglion. Each of these " Ol/aciens, cernens, oculosque movens, patiensque, " Gastans, abducens, audiensque, vagansque, loquensque.v * Besides these, there is another pair called accessorii, which arises from the medulla spinalis at its beginning; and ascending through the great foramen of the os occipitis into the cranium, passes out again close to the eighth pair, with which, however, it does not unite j and it is afterwards distributed chiefly to the muscles of the neck, back, and scapula. In this course it sends off filaments to different parts, and likewise communicates with several other nerves. Physiologists are at a loss how to account for the singular origin and course of these nervi accessorii. The ancients considered them as branches of the eighth pair, distri- buted to muscles of the scapula : Willis likewise considered them as appendages to that pair, and on that account named them ac- cessorii. They are sometimes called the spinal pair ; but as this latter name is applicable to all the nerves of the spine indiscrimi- nately, it seems better to adopt that given by Willis. Of the Brain and Nerves. 383 ganglions sends off two branches ; one ante- rior, and the other posterior. The anterior branches communicate with each other at their coming out of the spine, and likewise send off one, and sometimes more branches, to assist in the formation of the intercostal nerve. The knots or ganglions of the nerves just now spoken of, are not only to be met with at their exit from the spine, but likewise in vari- ous parts of the body. They occur in the nerves of the medulla oblongata, as well as in those of the spine. They are not the effects of disease, but are to be met with in the same parts of the same nerves, both in the foetus and adult. They are commonly of an oblong shape, and of a greyish colour, somewhat in- clined to red, which is perhaps owing to their being extremely vascular. Internally we are able to distinguish something like an intermix- ture of the nervous filaments. Some writers have considered them as so many little brains ; Lancisi fancied he had dis- covered muscular fibres in them, but they are certainly not of an irritable nature. A late writer, Dr. Johnstone,* imagines they are in- tended to deprive us of the power of the will over certain parts, as the heart, for instance: but if this hypothesis were well founded, we should meet with them only in the nerves lead- ing to involuntary muscles ; whereas it is cer- tain, that the voluntary muscles receive their nerves through ganglions. Doctor Monro, from observing the accurate intermixture of the * Essay on the Use of the Ganglio-s of the Nerves. 384 Of the Brain and Nerves. minute nerves which compose them, considers them as new sources of nervous energy.f The nerves, like the blood-vessels, in their course through the body, communicate with each other ; and each of these communications constitutes what is called a plexus, from whence branches are again detached to different parts of the body. Some of these are constant and considerable enough to be distinguished by particular names, as the semilunar plexus; the pulmonary plexus; the hepatic, the cardiac, he. It would be foreign to the purpose of this work, to follow the nerves through all their distributions ; but it may be remembered, that in describing the different viscera, mention was made of the nerves distributed to them. There is one pair, however, called the intercostal, or great sympathetic nerve, which seems to require particular notice, because it has an almost universal connection and correspondence with all the other nerves of the body. Authors are not perfectly agreed about the origin of the intercostal ; but it may perhaps not improperly be described, as beginning from filaments of the fifth and sixth pair; it then passes out of the cranium, through the bony canal of the carotid, from whence it descends laterally close to the bodies of the vertebra?, and receives branches from almost all the vertebral nerves ; forming almost as many ganglions in its course through the thorax and abdomen. It sends off an infinite number of branches to the viscera in f Observations on the Nervous System. Of the Brain and Nerves. 385 those cavities, and forms several plexuses with the branches of the eigiith pair or parvagum. That the nerves are destined to convey the principles of motion and sensibility to the brain from all parts of the system, there can be no doubt ; but how these effects are produced, no one has ever yet been able to determine. The inquiry has been a constant source of hypothesis in ail ages, and has produced some ingenious ideas, and many erroneous positions, but with- out having hitherto afforded much satisfactory information. Some physiologists have considered a trunk of nerves as a solid chord, capable of being divided into an infinite number of filaments, by means of which the impressions of feeling are conveyed to the sensorium commune. Others have supposed it to be a canal, which afterwards separates into more minute channels; or, perhaps, as being an assemblage of many very small and distinct tubes, connected to each other, and thus forming a cylindrical chord. They who contend for their being solid bodies, are of opinion, that feeling is occasioned by vibration: so that, for instance according to this system, by pricking the finger, a vibration would be occasioned in the nerve, distributed through its substance; and the effects of this vibration, when extended to the sensorium, would be an excital of pain. But the inelasticity, the softness, the con- nection, and the situation of the nerves, are so many proofs that vibration has no share in the cause of feeling. 3 C 386 Of the Brain and Nerves. Others have supposed, that in the brain and spinal marrow, a very subtile fluid is secreted, and from thence conveyed through the imper- ceptible tubes, which they consider as existing in the nerves. They have farther supposed, that this very subtile fluid, to which they have given the name of animal spirits, is secreted in the cortical substance of the brain and spi- nal marrow, from whence it passes through the medullary substance. This, like the other system, is founded altogether on hypothesis; but it seems to be an hypothesis derived from much more probable principles, and there are many ingenious arguments to be brought in its support* EXPLANATION of PLATE XXIX. Fig. 1. Represents the Inferior part of the Brain;—the Anterior part of the whole Spine, including the Medulla Spinalis ;—with the origin and large portions of all the Nerves. AA, The anterior lobes of the cerebrum. BB, The lateral lobes of the cerebrum. CC, The two lobes of the cerebellum. D, Tuber annulare. E, The passage from the third ventricle to the infundibulum. F, The me- dulla oblongata, which sends off the medulla spinalis through the spine. G G, That part of the os occipitis which is placed above (H H) the transverse processes of the first cervical vertebra. I I, he. The seven cervical verte- Ci- £ S Of the Brain and Nerves. 387 bras, with their intermediate cartilages. K K, he. The twelve dorsal vertebrae, with their intermediate cartilages. L L, he. The five lumbar vertebrae, with their intermediate car- tilages. M, The os sacrum. N, The os coe- cygis. Nerves.— 1 1, The first pair of nerves, nam- ed olfactory, which go to the nose. 2 2, The second pair, named optic, which goes to form the tunica retina of the eye. 3 3, The third pair, named motor oculi; it supplies most of the muscles of the eye-ball. 4 4, The fourth pair, named pathetic,—which is wholly spent upon the musculus trochlearis of the eye. 5 5, The fifth pair divides into three branches.— The first, named ophthalmic, goes to the orbit, supplies the lachrymal gland, and sends branches out to the forehead and nose.— The second, named superior maxillary, sup- plies the teeth of the upper jaw, and some of the muscles of the lips.—The third named in- ferior maxillary, is spent upon the muscles and teeth of the lower jaw, tongue, and mus- cles of the lips. 6 6, The sixth pair, which, after sending off the beginning of the intercos- tal or great sympathetic, is spent upon the ab- ductor oculi. 7 7, The seventh pair, named auditory, divides into two branches.—The largest, named portio mollis, is spent upon the internal ear. The smallest, portio dura, joins to the fifth pair within the internal ear by a re- flected branch from the second of the fifth.; and within the tympanum, by a branch from the third of the fifth named chorda tympani.*— 388 Of the Brain and Nerves. Vid. fig. 3. near B. 8 8, he. The eighth pair, named par vagum,—which accompanies the intercostal, and is spent upon the tongue, la- rynx, pharynx, lungs, and abdominal viscera. 9 9, The ninth pair, which are spent upon the tongue. 10 10, he. The intercostal, or great sympathetic, which is seen from the sixth pair to the bottom of the pelvis on each side of the spine, and joining with all the nerves of the spine ;—in its progress supplying the heart, and, with the par vagum, the contents of the abdomen and pelvis. 11 11, The accessorius, which is spent upon the sternocleido-mastoi- daeus and trapezius muscles. 12 12, The first cervical nerves;—13 13, The second cervical nerves ;—both spent upon the muscles that lie on the neck, and teguments of the neck and head. 14 14, The third cervical nerves, which, after sending off (15 15, he.) the phrenic nerves to the diaphragm, supply the muscles and teguments that lie on the side of the neck and top of the shoulder. 16 16, The brachial plexus, formed by the fourth, fifth, sixth, se- venth cervicals, and first dorsal nerves ; which supply the muscles and teguments of the su- perior extremity. 17 17, The twelve dorsal, or proper intercostal nerves, which are spent upon the intercostal muscles and some of the large muscles which lie upon the thorax. 18 18, The five lumbar pairs of nerves, which supply the lumbar and abdominal muscles, and some of the teguments and muscles of the inferior extremity. 19 19, The sacro-sciatic, or posterior crural nerve, formed by the two inferior lumbar, and three superior of the os Of the Brain and Nerves. 389 sacrum. This large nerve supplies the great- est part of the muscles and teguments of the inferior extremity. 20, The stomachic plexus, formed by the eighth pair. 21 21, Branches of the solar or caehac plexus, formed by the eighth pair and intercostals, which supply the stomach and chylopoietic viscera. 22 22, Branches of the superior and inferior mesen- teric plexuses, formed by the eighth pair and intercostals, which supply the chylopoietic vis- cera, with part of the organs of urine and generation. 23 23, Nerves which accompany the spermatic cord. 24 24, The hypogastric plexus, which supplies the organs of urine and generation within the pelvis. Fig. 2, 3, 4, 5. Shows different Views of the Inferior part of the Brain, cut perpendicu- larly through the Middle,—with the Origin and large Portions of all the Nerves which pass out through the Bones of the Cranium, —and the three first Cervicals. A, The anterior lobe. B, The lateral lobe of the cerebrum. C, One of the lobes of the cerebellum. D, Tuber annulare. E, Corpus pyramidale, in the middle of the medulla ob- longata. F, The corpus olivare, in the side of the medulla oblongata. G, The medulla ob- longata. H, The medulla spinalis. Nerves.— 1 2 3 4 5 6 7 8 and 9, Pairs of nerves. 10 10, Nervus accessorius, which comes from—11, 12, and IS, The three first cervical nerves. 390 Of the Senses and their Organs. PART VI. OF THE SENSES AND THEIR ORGANS. IN treating of the senses, we mean to con- fine ourselves to the external ones of touch, taste, smelling, hearing, and vision. The word sense, when applied to these five, seems to imply not only the sensation excited in the mind by certain impressions made on the body, but likewise the organ destined to receive and transmit these impressions to the sensorium. Each of these organs being of a peculiar struc- ture, is susceptible only of particular impres- sions, which will be pointed out as we pro- ceed to describe each of them separately. Sect. I. Of Touch. The sense of touch maybe defined to be the faculty of distinguishing certain properties of bodies by the feel. In a general acceptation, this definition might perhaps not improperly be extended to every part of the body possess- ed of sensibility,* but it is commonly con- * In the course of this article, mention has often been made of the sensibility or insensibility of different parts of the body : it will therefore, perhaps, not be amiss to observe in this place, that many parts which were formerly supposed to possess the most exquisite sense, are now known to have but little or no feeling, at least in a sound state ; for in an inflamed state, even the bones, the most insensible parts of any, become susceptible Of the Senses and their Organs. 391 fined to the nervous papillae of the cutis, or true skin, which, with its appendages, and their several uses, have been already de- scribed. rI he exterior properties of bodies, such as their solidity, moisture, inequality, smooth- ness, dryness, or fluidity, and likewise their degree of heat, seem all to be capable of mak- ing different impressions on the papillae, and consequently of exciting different ideas in the sensorium commune. But the organ of touch, like all the other senses, is not equally deli- cate in every part of the body, or in every subject; being in some much more exquisite than it is in others. Sect. II. Of the Taste. The sense of taste is seated chiefly in the tongue ; the situation and figure of which are sufficiently known. On the upper surface of this organ we may ' observe a great number of papillae, which, on account of their difference in size and shape, of the most painful sensations. This curious discovery is due to the late Baron Haller. His experiments prove, that the bones, cartilages, ligaments, tendons, epidermis, and membranes (as the pleura, pericardium, dura and pia mater, periosteum, Sec. j may in a healthy state be considered as insensible. As sensibility depends on the brain and nerves, of course different parts will possess a greater or less degree of feeling, in proportion as they are supplied with a greater or smaller number of nerves. Upon this principle it is, that the skin, muscles, stomach, intestines, urinary bladder, ureters, uterus, vatina peni:, tongue, and re- tina, are extremely sensible, while the lung* and glands haw- only an obscure degree of feeling. 392 Of the Senses and their Organs. are commonly divided into three classes. The largest are situated towards the basis of the tongue. Their number commonly varies from seven to nine, and they seem to be mucous follicles. Those of the second class are some- what smaller, and of a cylindrical shape. They are most numerous about the middle of the tongue. Those of the third class are very minute, and of a conical shape. They are very numerous on the apex and edges of the tongue, and have been supposed to be formed by the extremities of its nerves. We observe a line, the linea lingua mediana, running along the middle of the tongue, and dividing it as it were into two portions. To- wards the basis of the tongue, we meet with a little cavity, named by Morgagni foramen ccecum, which seems to be nothing more than a common termination of some of the excretory ducts of mucous glands situated within the substance of the tongue. We have already observed, that this organ is every where covered by the cuticle, which, by forming a reduplication, called the franum, at its under part, serves to prevent the too great motion of the tongue, and to fix it in its situation. But, besides this attachment, the tongue is connected by means of its muscles and membranous ligaments, to the lower jaw, the os hyoides, and the styloid processes. The principal arteries of the tongue are the linguales, which arise from the external carotid. Its veins empty themselves into the external jugulars. Its nerves arise from the fifth, eighth, and ninth pair. Of the Senses and their Organs. 393 The variety of tastes seems to be occasioned by the different impressions made on the papil- lae by the food. The different state of the pa* pUlae with respect to their moisture, their figure, or their covering, seems to produce a consider- able difference in the taste, not only in differ- ent people, but in the same subject, in sickness and in health. The great use of the taste seems to be to enable us to distinguish wholesome and salutary food from that which is unhealthy; and we observe that many quadrupeds, by having their papillae* very large and long, have the faculty of distinguishing flavours with infinite accuracy. Sect. III. Of Smelling. The sense of smelling, like the sense of taste, seems intended to direct us to a proper choice of aliment, and is chiefly seated in the nose, which is distinguished into its external and in- ternal parts. The situation and figure of the former of these do not seem to require a defini- tion. It is composed of bones and cartilages, covered by muscular fibres and by the common integuments. The bones make up the upper portion, and the cartilages the lower one. The septum narium, like the nose, is likewise in part bony, and in part cartilaginous. These bones and their connections were described in the osteology. 3 i) * Malpighi's description of the papillae, which has been copied by many anatomical writers, seems to have been taken chiefly from the tongues o^ -Heep. 394 Of the Senses and their Organs. The internal part of the nose, besides the ossa spongiosa, has six cavities or sinuses, the maxillary, the frontal, and the sphenoid, which were all described with the bones of the head. They all open into the nostrils; and the nose likewise communicates with the mouth, larynx, and pharynx, posteriorly be- hind the velum palati. All these several parts, which are included in the internal division of the nose, viz. the in- ner surface of the nostrils, the lamellae of the ossa spongiosa, and the sinuses, are lined by a thick and very vascular membrane, which, though not unknown to the ancients, was first well described by Schneider,* and is therefore now commonly named membrana pituitaria Schneideri. This membrane is truly the organ of smelling; but its real structure does not yet seem to be perfectly understood. It appears to be a continuation of the cuticle, which lines the inner surface of the mouth. In some parts of the nose it is smooth and firm, and in others it is loose and spongy. It is constantly moist- ened by a mucous secretion; the finer parts of which are carried off by the air we breathe, and the remainder, by being retained in the sinuses, acquires considerable consistence. The manner in which this mucus is secreted has not yet been satisfactorily ascertained; but it seems to be by means of mucous follicles. Its arteries are branches of the internal max- illary and internal carotid. Its veins empty themselves into the internal jugulars. The first pair of nerves, the olfactory, are spread * De Catarrho, lib. iii. Of the Senses and their Organs. 395 over every part of it, and it likewise receives branches from the fifth pair. After what has been said of the pituitary membrane, it will not be difficult to conceive how the air we draw in at the nostrils, being impregnated with the effluvia of bodies, ex- cites in us that kind of sensation we call smell' ing. As these effluvia, from their being ex- ceedingly light and volatile, cannot be capable in a small quantity of making any great im- pression on the extremities of the olfactory nerves, it was necessary to give considerable extent to the pituitary membrane, that by this means a greater number of odoriferous parti- cles might be admitted at the same time. When we wish to take in much of the effluvia of any thing, we naturally close the mouth, that all the air we inspire may pass through the nos- trils ; and at the same time, by means of the muscles of the nose, the nostrils are dilated, and a greater quantity of air is drawn into them. In many quadrupeds, the sense of smelling is much more extensive and delicate than it is in the human subject; and in the human sub- ject it seems to be more perfect the-less it is vitiated by a variety of smells. It is not al- ways in the same state of perfection, being na- turally affected by every change of the pitui- tary membrane, and of the lymph with whicrj 1 that membrane is moistened. 396 Of tiie Senses and their Organs. Sect., IV. Of Hearing. Before we undertake to explain the man- ner in which we are enabled to receive the im- pressions of sound, it will be necessary to de- scribe the ear, which is the organ of hearing. It is commonly distinguished into external and internal. The former of these divisions in- cludes all that we are able to discover without dissection, and the meatus auditorius, as far as the tympanum -r and the latter, all the other parts of the ear. The external ear is a cartilaginous funnel, covered by the common integuments, and at- tached, by means of its ligaments and muscles, to the temporal bone. Although capable only of a very obscure motion, it is found to have several muscles. Different parts of it are dis- tinguished by several names; all its cartila- ginous part is called ala or wing, to distin- guish it from the soft and pendent part below, called the lobe. Its outer circle or border is called helix, and the semicircle within this, antihelix. The moveable cartilage placed im- mediately before the meatus auditorius, which it may be made to close exactly, is named tragus; and an eminence opposite to this at the extremity of the antihelix, is called anti- tragus. The concha is a considerable cavity formed by the extremities of the helix and an- tihelix. The meatus auditorius, which at its opening is cartilaginous, is lined with a very thin membrane, which is a continuation of the cuticle from the surface of the ear. Of the Senses and their Organs. 397 In this canal we find a yellow wax, which is secreted by a number of minute glands or follicles, each of which has an excretory duct. This secretion, which is at first of an oily con- sistence, defends the membrane of the tym- panum from the injuries of the air ; and by its bitterness, prevents minute insects from enter- ing into the ear. But when from neglect or disease it accumulates in too great a quantity, it sometimes occasions deafness. The inner extremity of the meatus is closed by a very1 thin transparent membrane, the membrana tympani, which is set in a bony circle like the head of a drum. In the last century Rivinus, professor at Liepsic, fancied he had discovered a hole in this membrane, surrounded by a sphincter, and affording a passage to the air, between the external and internal ear. Cow- per, Heister, and some other anatomists, have admitted this supposed foramen, which cer- tainly does not exist. Whenever there is any opening in the membrana tympani, it may be considered as accidental. Under the mem- brana tympani runs a branch of the fifth pair of nerves, called chorda tympani; and beyond this membrane is the cavity of the tympanum, which is about seven or eight lines wide, and half so many in depth ; it is semispherical, and every where lined by a very fine membrane. There are four openings to be observed in this cavity. It communicates with the mouth by means of the Eustachian tube. This canal, which is in part bony and in part cartilaginous, begins by a very narrow opening at the anteri- or and almost superior part of the tympanum. 398 Of the Senses and their Organs. increasing in size as it advances towards the palate of the mouth, where it terminates by an oval opening. This tube is every where lined by the same membrane that covers the inside of the mouth. The real use of this ca- nal does not seem to have been hitherto satis- factorily ascertained; but sound would seem to be conveyed through it to the membrana tympani, deaf persons being often observed to listen attentively with their mouths open. Opposite to this is a minute passage, which leads to the sinuosities of the mastoid process; and the two other openings, which are in the internal process of the os petrosum, are the fenestra ovalis, and fenestra rotunda, both of which are covered by a very fine membrane. There are three distinct bones in the cavity of the tympanum; and these are the malleus, incus, and stapes. Besides these there is a fourth, which is the os orbiculare, considered by some anatomists as a process of the stapes, which is necessarily broken off by the violence we are obliged to use in getting at these bones; but when accurately considered, it seems to be a distinct bone. The malleus is supposed to resemble a ham- mer, being larger at one extremity, which is its head, than it is at the other, which is its handle. The latter is attached to the mem- brana tympani, and the head of the bone is articulated with the incus. The incus, as it is called from its shape, though it seems to have less resemblance to an anvil than to one of the dentes molares with its roots widely separated from each other, is Of the Senses and their Organs. 399 distinguished into its body and its legs. One of its legs is placed at the entry of the canal which leads to the mastoid process; and the other, which is somewhat longer, is articulat- ed with the stapes, or rather with the os orbi- culare, which is placed between them. The third bone is very properly named stapes, being perfectly shaped like a stirrup. Its basis is fixed into the fenestra ovalis, and its upper part is articulated with the os orbicu- lare. What is called the fenestra rotunda, though perhaps improperly, as it is more oval than round, is observed a little above the other, in an eminence formed by the os petro- sum, and is closed by a continuation of the membrane that lines the inner surface of the tympanum. The stapes and malleus are each of them furnished with a little muscle, the sta- pedeus and tensor tympani. The first of these, which is the smallest in the body, arises from a little cavern in the posterior and upper part of the cavity of the tympanum; and its tendon, after passing through a hole in the same ca- vern, is inserted at the back part of the head of the stapes. This muscle, by drawing the stapes obliquely upwards, assists in stretch- ing the membrana tympani. The tensor tympani,* or internus mallei, as it is called by some writers, arises from the cartilaginous extremity of the Eustachian tube, and is inserted into the back part of the handle * Some anatomists describe three muscles of the malleus; but only this one seems to deserve the name of muscle; what are called the externus and obliquus mallei, seeming to be ligaments rather than muscles. 400 Of the Senses and their Organs, of the malleus, which it serves to pull inwards, and of course helps to stretch the membrana tympani. The labyrinth is the only part of the ear which remains to be described. It is situated in the os petrosum, and is separated from the tympanum by a partition which is every where bony, except at the two fenestra?. It is com- posed of three parts; and these are the vesti- bulum, the semicircular canals, and the coch- lea. The vestibulum is an irregular cavity, much smaller than the tympanum, situated nearly in the centre of the os petrosum, between the tympanum, the cochlea, and the semicircular canals. It is open on the side of the tympa- num by means of the fenestra ovalis, and com- municates with the upper portion of the coch- lea by an oblong foramen, which is under the fenestra ovalis, from which it is separated only by a very thin partition. Each of the three semicircular canals forms about half a circle of nearly a line in diameter, and running each in a different direction, they are distinguished into vertical, oblique, and ho- rizontal. These three canals open by both their extremities into the vestibulum; but the vertical and the oblique being united together at one of their extremities, there are only five orifices to be seen in the vestibulum. The cochlea is a canal which takes a spiral course, not unlike the shell of a snail. From its basis to its apex it makes two turns and a half; and is divided into two canals by a very thin lamina or septum, which is in part bony Of the Senses and their Organs. 401 and in part membranous, in such a manner that these two canals only communicate with each other at the point. One of them opens into the vestibulum, and the other is covered by the membrane that closes the fenestra ro- tunda. The bony lamella which separates the two canals is exceedingly thin, and fills about two-thirds of the diameter of the canal. The rest of the septum is composed of a most deli- cate membrane, which lines the whole inner surface of the cochlea, and seems to form this division in the same manner as the two mem- branous bags of the pleura, by being applied to each other, form the mediastinum. Every part of the labyrinth is furnished with a very delicate periosteum, and filled with a watery fluid, secreted as in other cavities. This fluid transmits to the nerves the vibrations it receives from the membrane closing the fenes- tra rotunda, and from the basis of the stapes, where it rests on the fenestrum ovale. When this fluid is collected in too great a quantity, or is compressed by the stapes, it is supposed to escape through two minute canals or aque- ducts, lately described by Dr. Cotunni,* an in- genious physician at Naples. One of these aqueducts opens into the bottom of the ves- tibulum, and the other into the cochlea, near the fenestra rotunda. They both pass through the os petrosum, and communicate with the ca- vity of the cranium where the fluid that passes through them is absorbed; and they are lined 3 E * De aquseductibus Auris Humans Interna, 8vo, 1760, 402 Of the Senses and their Organs. by a membrane which is supposed to be a pro- duction of the dura mater. The artei>s of the external ear come from the temporal and other branches of the exter- nal carotid, and its veins pass into the jugular. The internal ear receives branches of arteries from the basilary and carotids, and its veins empty themselves into the sinuses of the dura mater, and into the internal jugular. The portio mollis of the seventh pair is dis- tributed through the cochlea, the vestibulum, and the semi-cireular canals ; and the portio dura sends off a branch to the tympanum, and other branches to the external ear and parts near k. The sense of hearing, in producing which all the parts we have described assist, is occa- sioned by a certain modulation of the air col- lected by the funnel-like shape of the external ear, and conveyed through the meatus audito- rius to the membrana tympani. That sound is propagated by means of the air, is very ea- sily proved by ringing a bell under the receiv- er of an air-pump; the sound it affords being found to diminish gradually as the air becomes exhausted, till at length it ceases to be heard at all. Sound moves through the air with in- finite velocity; but the degree of its motion seems to depend on the state of the air, as it constantly moves faster in a dense and dry, than it does in a moist and rarefied air. That the air vibrating on the membrana tympani communicates its vibration to the dif- ferent parts of the labyrinth, and by means of Of the Sefises and their Organs. 403 the fluid contained in this cavity affects the au- ditory nerve so as to produce sound, seems to be very probable ; but the situation, the mi- nuteness, and the variety of the parts which compose the ear, do not permit much to be ad- vanced with certainty concerning their mode of action. Some of these parts seem to constitute the immediate organ of hearing, and these are all the parts of the vestibulum;; but there ate others which seem intended for the perfection of this sense, without being absolutely essential to it. It has happened, far instance, that the mem- brana tympani, and the little bones of the ear, have been destroyed by disease, without de- priving .the patient of the sense of hearing..* .Sound is'more or less loud in proportion to the strength of the vibration; and the variety of sounds seems to depend on the difference of this vibration ; for the more quick and fre- quent it is, the more acute will be the sound, and vice versa. Before we conclude this article, it will be right to explain certain phenomena, which will be found to fiave a relation to the organ of hearing. Every body has, in consequence of particu- lar sounds, occasionally felt that disagreeable sensation which is usually called setting the teeth on edge: and the cause of this sensation * This observation has led to a supposition, that a perforation of this membrane may in some cases of deafness be useful; and Mr. Cheselden relates, that, some years ago, a malefactor was pardoned on condition that he should submit to this operation ; but the public clamour raised against it was so great, that it Was thought right not to perform it. 404 Of the Senses and their Organs. may be traced to the communication which the portio dura of the auditory nerve has with the branches of the fifth pair that are distributed to the teeth, being probably occasioned by the violent tremor produced in the membrana tym- pani by these very acute sounds. Upon the same principle we may explain the strong idea of sound which a person has who holds a vi- brating string between his teeth. The humming which is sometimes perceiv- ed in the ear, without any exterior cause, may be occasioned either by an increased action of the arteries in the ears, or by convulsive con- tractions of the muscles of the malleus and stapes, affecting the auditory nerve in such a manner as to produce the idea of sound. An ingenious philosophical writer* has lately dis- covered that there are sounds liable to be ex- cited in the ear by irritation, and without any assistance from the vibrations of the air. Sect. V. Of Vision. The eyes, which constitute the organ of vi- sion, are situated in two bony cavities named orbits, where they are surrounded by several parts, which are either intended to protect them from external injury, or to assist in their motion. The globe of the eye is immediately covered by two eye-lids or palpebral, which are com- * Elliot's Philosophical Observations on the Senses of Vision and Hearing, 8vo. Of the Senses and their Organs. 405 posed of muscular fibres covered by the com- mon integuments, and lined by a very fine and smooth membrane, which is from thence ex- tended over part of the globe of the eye, and is called tunica conjunctiva. Each eye-lid is cartilaginous at its edge ; and this border which is called tarsus, is furnished with a row of hairs named cilia or eye-lashes. The cilia serve to protect the eye from in- sects and minute bodies floating in the air, and likewise to moderate the action of the rays of light in their passage to the retina. At the roots of these hairs there are sebaceous folli- cles, first noticed by Meibomius, which dis- charge a glutinous liniment. Sometimes the fluid they secrete has too much viscidity, and the eye-lids become glued to each other. The upper border of the orbit is covered by the eye-brows or supercilia, which by means of their two muscles are capable of being brought towards each other, or of being carried upwards. They have been considered as serv- ing to protect the eyes, but they are probably intended more for ornament than utility.* The orbits in which the eyes are placed, are furnished with a good deal of fat, which af- fords a soft bed on which the eye performs its several motions. The inner angle of each or- bit, or that part of it which is near the nose, is called canthus major, or the great angle; and the outer angle, which is on the opposite side of the eye, is the canthus minor, or little angle. * It is observable, that the eye-brows are peculiar to the hu- man species. 406 Of the Senses and their Organs. The little reddish body which we observe in the great angle of the eye-lids, and which is called caruncula lachrymalis, is supposed to be of a glandular structure, and,, like the follicles of the eye-lids, to secrete an oily humour. But its structure and use do not seem to have been hitherto accurately determined. The surface of the eye is constantly moistened by a very fine limpid fluid called the tears, which is chief- ly, and perhaps wholly, derived from a large gland of the conglomerate kind, situated in a small depression of the os frontis near the out- er angle of the eye. Its excretory ducts pierce the tunica conjunctiva just above the cartila- ginous borders of the upper eye-lids. When the tears were supposed to be secreted by the caruncule, this gland Wats called glandula inno- minata; but now that its structure and uses are ascertained, it very properly has the name of glandula lachrymalis. The tears poured out by the ducts of this gland are, in a natur- al and healthy stated incessantly spread over the surface of the eye, to keep it clear and trans- parent, by means of the eye-lids, and as con- stantly pass out at the opposite corner of the eye or inner angle, through two minute orifi- ces, the puncta lachrymalia ;* being determined into these little openings by a reduplication of the tunica conjunctiva, shaped like a crescent the two points of which answer to the puncta. * It sometimes happens, that this very pellucid fluid, which moistens the eye, being poured out through the excretory ducts of the lachrymal gland faster than it can be carried off through the puncta, trickles down the cheek, and is then strictly and pro- perly called tears. Of the Senses and their Organs. 407 This reduplication is named membrana, or val- vula semilunaris. Each of these puncta is the beginning of a small excretory tube, through which the tears pass into a little pouch or re- servoir, the sacculus lachrymalis, which lies in an excavation formed partly by the nasal process of the os maxillare superius, and part- ly by the os unguis. The lower part of this sac forms a duct called the ductus ad nares, which is continued through a bony channel, and opens into the nose, through which the tears are occasionally discharged.* The motions of the eye are performed by six muscles ; four of which are straight and two oblique. The straight muscles are dis- tinguished by the names of elevator, depres- sor, adductor, and abductor, from their se- veral uses in elevating and depressing the .eye, drawing it towards the nose, or carrying it from the nose towards the temple. All these four muscles arise from the bottom of the or- bit, and are inserted by flat tendons into the globe of the eye. The oblique muscles are intended for the more compound motions of the eye. The first of these muscles, the obli- quus superior, does not, like the other four mus- cles we have described, arise from the bottom of the orbit, but from the edge of the fora- men that transmits the optic nerve, which se- parates the origin of this muscle from that of ,* "When the ductus ad nares becomes obstructed in conse- quence of disease, the tears are no longer able to pass into the nos- trils ; the sacculus lachrymalis becomes distended ; and inflam- mation, and sometimes ulceration taking place, constitute the disease called fistula lachrymal; r. 408 Of the Senses and their Organs. the others. From this beginning it passes in a straight line towards a very small cartilagin- ous ring, the situation of which is marked in the skeleton by a little hollow in the internal orbitar process of the os frontis. The tendon of the muscle after passing through this ring, is inserted into the upper part of the globe of the eye, which it serves to draw forwards, at the same time turning the pupil downwards. The obliquus inferior arises from the edge of the orbit, under the opening of the ductus lachrymalis ; and is inserted somewhat poste- riorly into the outer side of the globe, serving to draw the eye forwards and turn the pupil upwards. When either of these two muscles acts separately, the eye is moved on its axis; but when they act together, it is compressed both above and below. The eye itself, which is now to be described, with its tunics, hu- mours, and component parts, is nearly of a spherical figure. Of its tunics, the conjuncti- va has been already described as a partial co- vering, reflected from the inner surface of the eye-lids over the anterior portion of the eye. What has been named albuginea cannot pro- perly be considered as a coat of the eye, being in fact nothing more than the tendons of the straight muscles spread over some parts of the sclerotica. The immediate tunics of the eye, which are to be demonstrated when its partial coverings, and all the other parts with which it is sur- rounded, are removed, are the sclerotica, cor- nea, choroides, and retina. Of the Senses and their Organs. 409 The sclerotica, which is the exterior coat, is every where white and opaque* and is join- ed at its anterior edge to another, which has more convexity than any other part of the globe, and being exceedingly transparent is called cornea.* These two parts are perfectly different in their structure; so that some ana- tomists suppose them to be as distinct from each other as the glass of a watch is from the case into which it is fixed. The sclerotica is of a compact fibrous structure ; the cornea, on the other hand, is composed of a great num- ber of laminae united by cellular membrane. By macerating them in boiling water, they do not separate from each other, as some writers have asserted; but the cornea soon softens, and becomes of a glutinous consistence. The ancients supposed the sclerotica to be a continuation of the dura mater. Morgagni, and some other modern writers, are of the same opinion ; but this point is disputed by Winslow, Haller, Zin, and others. The truth seems to be, that the sclerotica, though not a production of the dura mater, adheres intimately to that membrane. The choroides is so called because it is fur- nished with a great number of vessels. It has likewise been named uvea*, on account of its re- semblance to a grape. Many modern anatomi- cal writers have considered it as a production of the pia mater. This was likewise the opinion 3 F * Some writers, who have given the name of cornea to all this outer coat, have named what is here and most commonly called sclerotica, cornea opaca ; and its anterior and transparent portion, eomea lucida. 410 Of the Senses and their Organs. of the ancients; but the strength and thickness of the choroides, when compared with the deli- cate structure of the pia mater, are sufficient proofs of their being two distinct membranes. The choroides has of late generally been de- scribed as consisting of two laminae ; the inner- most of which has been named after Ruysch, who first described it. It is certain, however, that Ruysch's distinction is ill founded, at least with respect to the human eye, in which we are unable to demonstrate any such structure, al- though the tunica choroides of sheep and some other quadrupeds may easily be separated into two layers. The choroides adheres intimately to the scle- rotica round the edge of the cornea ; and at the place of this union, we may observe a little whitish areola, named ligamentum ciliare9 though it is not of a ligamentous nature. They who suppose the choroides to be com- posed of two laminae, describe the external one as terminating in the ligamentum ciliare, and the internal one as extending farther to form the iris, which is the circle we are able to distin- guish through the cornea; but this part is of a very different structure from the choroides; so that some late writers have perhaps not im- properly considered the iris as a distinct mem- brane. It derives its name from the variety of its colours, and is perforated in the middle.— This perforation, which is called the pupil or sight of the eye, is closed in the foetus by a very thin vascular membrane. This membrana pu- pillaris commonly disappears about the seventh month. Of the Senses and their Organs. 411 On the under side of the iris we observe ma- ny minute fibres, called ciliary processes, which pass in radii or parallel lines from the circum- ference to the centre. The contraction and dilatation of the pupil are supposed to depend on the action of these processes. Some have considered them as muscular, but they are not of an irritable nature: others have sup- posed them to be filaments of nerves; but their real structure has never yet been clearly ascertained. Besides these ciliary processes, anatomists usually speak of the circular fibres of the iris, but no such seem to exist. The posterior surface of the iris, the ciliary processes, and part of the tunica choroides, are covered by a black mucus for the purposes of accurate and distinct vision; but the man- ner in which it is secreted has not been deter- mined. Immediately under the tunica choroides we find the third and inner coat, called the retina, which seems to be merely an expansion of the pulpy substance of the optic nerve, extending to the border of the crystalline humour- The greatest part of the globe of the eye, within these several tunics, is filled by a very transparent and gelatinous humour of conside- rable consistence, which, from its supposed resemblance to fused glass, is called the vitre- ous humour. It is invested by a very fine and delicate membrane, called tunica vitrea, and sometimes arachnoides.—It is supposed to be composed of two laminae; one of which dips into its substance, and by dividing the hu- 412 Of the Senses and their Organs\ mour into cells adds to its firmness. The fore-part of the vitreous humour is a little hollowed, to receive a very white and trans* parent substance of a firm texture, and of a lenticular and somewhat convex shape, named the crystalline humour. It is included in a capsula, which seems to be formed by a sepa- ration of the two laminae of the tunica vitrea. The fore-part of the eye is filled by a very thin and transparent fluid, named the aqueous humour, which occupies all the space between the crystalline and the prominent cornea— The part of the choroides which is called the iris, and which comes forward to form the pupil, appears to be suspended as it were in this humour, and has occasioned this portion of the eye to be distinguished into two parts. One of these, which is the little space between the anterior surface of the crystalline and the iris, is called the posterior chamber; and the other, which is the space between the iris and the cornea, is called the anterior chamber of the eye.* Both these spaces are completely filled with the aqueous humour.f * We are aware that some anatomists, particularly Lieutaud, are of opinion, that the iris is every where in close contact widi the crystalline, and that it is of course right to speak only of one chamber of the eye ; but as this does not appear to be the case, the situation cf the iris and the two chambers of the eye are here described in the usual way. f When the crystalline becomes opaque, so as to prevent the passage of the rays of light to the retina, it constitutes what is called a cataract; and the operation of couching consists in re- moving the diseased crystalline from its bed in the vitreous hu- mour. In this operation the cornea is perforated, and the aque- ous humour escapes out of the eye, but it is constantly renewed again in a very short time. The manner, however, in which it is secreted, has not yet been determined; Of the Senses mid their Organs. 4,13 ; The eye receives its arteries from the in- ternal carotid through the foramina optica; and its veins pass through the foramina lacera, and empty themselves into the lateral sinuses. Some of the ramifications of these vessels ap- pear on the inner surface of the iris, where they are seen to make very minute convolu- tions, which are sufficiently remarkable to be distinguished by the name of circuius arteriosus^ though perhaps improperly, as they are chief- ly branches of veins. The optic nerve passes in at the posterior part of the eye, in a considerable trunk, to be expanded for the purposes of vision, of which it is now universally supposed to be the im- mediate seat. But Messrs. Mariotte and Mery contended, that the choroides is the seat of this sense; and the ancients supposed the crystalline to be so. Besides, the optic, the eye receives branches from the third, fourth, fifth, and sixth pair of nerves. The humours of the eye, together with the cornea, are calculated to refract and converge the rays of light in such a manner as to form at the bottom of the eye a distinct image of the object we look at; and the point where these rays meet is called the focus of the eye. On the retina, as in the camera obscura, the ob- ject is painted in an inverted position; and it is only by habit that we are enabled to judge of its true situation, and likewise of its dis- tance and magnitude. To a young gentleman who was born blind, and who was couched by Mr. Cheselden, every object (as he expressed himself) seemed to touch his eyes as what he >414 Of the Senses and their Organs. felt did his skin; and he thought no objects so agreeable as those which were smooth and regular, although for some time he could form no judgment of their shape, or guess what it was in any of them that was pleasing to him. In order to paint objects distinctly on the retina, the cornea is required to have such a degree of convexity, that the rays of light may be collected at a certain point, so as to termi- nate exactly on the retina.—If the cornea is too prominent, the rays, by diverging too soon, will be united before they reach the retina, as is the case with near-sighted people or myopes ; and on the contrary, if it is not suf- ficiently convex, the rays will not be perfectly united when they reach the back part of the eye; and this happens to long-sighted people or presbi, being found constantly to take place as we approach to old age, when the eye gra- dually flattens.* These defects are to be sup- plied by means of glasses. He who has too prominent an eye, will find his vision improv- ed by means of a concave glass; and upon the same principles, a convex glass will be found useful to a person whose eye is natu^ rally too flat. * Upon this principle, they who in their youth are near sight- ed may expect to see better as they advance in life, as their eyes gradually become more flat. \natom> I'l.-.te.YW ^/>,.-*>/.:■/«/ Of the Senses and their Organs. 415 EXPLANATION OF PLATE XXX. Figure 1. Shows the Lachrymal Canals, after the Common Teguments and Bones have been cut away. a, The lachrymal gland, b, The two punc- ta lachrymalia, from which the two lachry- mal canals proceed to c, The lachrymal sac. d, The large lachrymal duct, e, Its opening into the nose, f, The caruncula lachrymalis. g, The eye-ball. Fig. 2. An interior View of the Coats and Humours of the Eye. a a a a, The tunica sclerotica cut in four angles, and turned back, b b b b, The tunica choroides adhering to the inside of the sclero- tica, and the ciliary vessels are seen passing over—c c, The retina which covers the vitre- ous humour. d d, The ciliary processes, which were continued from the choroid coat. e e, The iris, f, The pupil. Fig. 3. Shows the Optic Nerves, and Mus- cles of the Eye. a a, The two optic nerves before they meet. b? The two optic nerves conjoined, c, The 415 Of the Senses and their Organs. right optic nerve, d, Musculus attollens pal- pebral superioris. e, Attollens oculi. f, Ab- ductor, g g, Obliquus superior, or trochle- aris. h, Adductor, i, The eye-ball. Fig. 4. Shows the Eye-ball with its Muscles. a, The optic nerve, b, Musculus trochle- aris. c, Part of the os frontis, to which the trochlea or pully is fixed, through which,—d, The tendons of the trochlearis passes, e, At- tollens oculi. f, Adductor oculi. g, Abduc- tor oculi. h, Obliquus inferior, i, Part of the superior maxillary bone to which it is fixed, k, The eye-ball. Fig. 5. Represents the Nerves and Muscles of the Right Eye, after part of the Bones of the orbit have been cut away. A, The eye-ball. B, The lachrymal gland. C, Musculus abductor oculi. D, Attollens. E, Levator palpebral superioris. F, Depres- sor oculi. G, Adductor. H, Obliquus superi- or, with its pully. I, Its insertion into the sclerotic coat. K, Part of the obliquus inferior. L, The anterior part of the os frontis cut. M, The crista galli of the ethmoid bone. N, The posterior part of the sphenoid bone. O, Trans- verse spinous process of the sphenoid bone. P, The carotid artery, denuded where it pass- es through the bones. Q,, The carotid artery within the cranium. R, The ocular artery. Of the Senses and their Organs. 417 Nerves.—a a, The optic nerve, b, The third pair.—c, Its joining with a branch of the first branch of the fifth pair, to form 1,—The lenticular ganglion, which sends off the ciliary nerves, d. e e, The fourth pair, f, The trunk of the fifth pair, g, The first branch of the fifth pair, named ophthalmic.—h, The frontal branch of it. i, Its ciliary branches, along with which the nasal twig is sent to the nose. k, Its branch to the lachrymal gland. 1, The lenticular ganglion. m, The second branch of the fifth pair, named superior max- illary, n, The third branch of the fifth pair, named inferior maxillary, o, The sixth pair of nerves,—which sends off p, The beginning of the great sympathetic, q, The remainder of the sixth pair, spent on c, The abductor oculi. Fig. 6. Represents the head of a youth, where the upper part of the cranium is sawed off,— to show the upper part of the brain, covered by the pia mater, the vessels of which are minutely filled with wax. A A, The cut edges of the upper part of the cranium. B, The two tables and interme- diate diploe. B B, The two hemispheres of the cerebrum. C C, The incisure made by the falx. D, Part of the tentorium cerebello super expansum. E, Part of the falx, which is fixed to the crista galli. 3 G 418 Of the Senses and their, Organs. Fig. 7. Represents the parts of the External Ear, with the Parotid Gland and its Duct. a a, The helix, b, The antihelix. c, The antitragus. d, The tragus. e, The lobe of the ear. f, The cavitas innominata. g, The scapha. h, The concha. i i, The parotid gland, k, A lymphatic gland, which is often found before the tragus. 1, The duct of the parotid gland, m, Its opening into the mouth. Fig. 8. A view of the posterior part of the external ear, meatus auditorius, tympanum, with the small bones, and Eustachian tube of the right side. a, The back part of the meatus, with the small ceruminous glands, b, The incus, c, Malleus, d, The chorda tympani. e, Mem* brana tympani. f, The Eustachian tube, g, Its mouth from the fauces. Fig. 9. Represents the anterior part of the right external ear, the cavity of the tympa- num—its small bones, cochlea, and semicir- cular canals. a, The malleus, b, Incus wit^ its lan£ leg, resting upon the stapes, c, Membrana tym- pani. d, e, The Eustachian tube, covered by part of—f f, The musculus circumflexus palati. 1, 2, 3, The three semicircular canals. 4, The vestible. 5, The cochlea. 6, The por- tio mollis of the seventh pair of nerves. Of the Senses and their Organs, 419 Fig. 10. Shows the muscles which compose the fleshy substance of the Tongue. a a, The tip of the tongue, with some of the papillae minimae. b, The root of the tongue. c, Part of the membrane of the tongue, which covered the epiglottis. d d, Part of the mus- culus hyo-glossus. e, The lingualis. f, Ge- nio-glossus. g g, Part of the stylo-glossus. THE END. Med,. Hist. WZ. X10 "■:;